227 lines
29 KiB
XML
227 lines
29 KiB
XML
<document id="5E4803BBBD3A89B8758FAC953F8C499E" ID-DOI="10.11646/zootaxa.3841.3.4" ID-GBIF-Dataset="fd4f8193-3005-4d9b-ae7f-304b847215a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="225778" ID-ZooBank="E52D2F6B-631D-45FE-97E0-DD26137757FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462027068" checkinUser="plazi" docAuthor="Bianco, Pier Giorgio & Ketmaier, Valerio" docDate="2014" docId="8E3D87DAFFC4FFA266A3FE30F95CFC12" docLanguage="en" docName="zt03841p402.pdf" docOrigin="Zootaxa 3841 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucos ylikiensis Economidis 1991" docType="treatment" docVersion="8" lastPageNumber="389" masterDocId="7204FFA2FFCCFFA86634FFEBFF9AFFB4" masterDocTitle="A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)" masterLastPageNumber="402" masterPageNumber="379" pageNumber="387" updateTime="1698304242431" updateUser="plazi">
|
||
<mods:mods id="4538FBF2FACDECD8EB6F85EBFC4FCF99" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="ADF75EB9A162CC368B816BF3FCB830BC">
|
||
<mods:title id="97EBFC0839EFF13F2D59D77062386DF5">A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="97300A1CE5BF19608B704EE0D1041A62" type="personal">
|
||
<mods:role id="CD59B6BC59533259E66DD73ED14988DF">
|
||
<mods:roleTerm id="69FDCE13C933EF377121D0833EAE8348">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3A08E4511DEDC78D3B7C2039BD3CA0D4">Bianco, Pier Giorgio</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C5994DE36392AE16FFFAA13AC4C4F8BC" type="personal">
|
||
<mods:role id="DD921A57CC728E236C4AFBB12B0794B2">
|
||
<mods:roleTerm id="C328F63B29F4696D1B187CDE142934EE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="451C509071BAC2437A414245D840ADD7">Ketmaier, Valerio</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="47696098C077D81345E91047BBFC657D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2B91B11C4F1F1EDD162A784751C03D71" type="host">
|
||
<mods:titleInfo id="78CAA677FA6C4401BA5686D8F7AD83B9">
|
||
<mods:title id="F4C50CF04FC30577C20B0EA52E705BE3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="16CA1D498E041A6EA187C90473D06F7A">
|
||
<mods:date id="605B8D81ABD76ED6540CF15BE3569B59">2014</mods:date>
|
||
<mods:detail id="AFAC3425406AFD513C31B4E21E94BDA8" type="volume">
|
||
<mods:number id="11897CC31ECDA5964777CC5CE5CABE7A">3841</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F1896C488F4F77450783D9C259016B5C" type="issue">
|
||
<mods:number id="C8AA3C6655840BA188CAB01310E5C442">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="93A8A54E1CF0B2D32B9E7FBE1DDAE6AF" unit="page">
|
||
<mods:start id="6BACC55279646B87A9DAFE508327D9FD">379</mods:start>
|
||
<mods:end id="9535363A6390D61FC0A7045CF33A1E24">402</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="3A47486FFD6C96BFAB11C8D169490466">journal article</mods:classification>
|
||
<mods:identifier id="CE468C38755DE3F7B76AD52BBBE687BC" type="DOI">10.11646/zootaxa.3841.3.4</mods:identifier>
|
||
<mods:identifier id="CB10C9AE124051CAC6B70A3B6B4890ED" type="GBIF-Dataset">fd4f8193-3005-4d9b-ae7f-304b847215a5</mods:identifier>
|
||
<mods:identifier id="1035482ACDD4434D72939167BD5150AC" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="AE60CE516618CF3AEB5962A6B0A6A01F" type="Zenodo-Dep">225778</mods:identifier>
|
||
<mods:identifier id="525213F679944E0B5D781B9E6F5426B7" type="ZooBank">E52D2F6B-631D-45FE-97E0-DD26137757FE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8E3D87DAFFC4FFA266A3FE30F95CFC12" ID-DOI="http://doi.org/10.5281/zenodo.6140507" ID-GBIF-Taxon="119512644" ID-Zenodo-Dep="6140507" LSID="urn:lsid:plazi:treatment:8E3D87DAFFC4FFA266A3FE30F95CFC12" httpUri="http://treatment.plazi.org/id/8E3D87DAFFC4FFA266A3FE30F95CFC12" lastPageId="10" lastPageNumber="389" pageId="8" pageNumber="387">
|
||
<subSubSection id="4E8E6547FFC4FFA066A3FE30FC9EFD2E" pageId="8" pageNumber="387" type="nomenclature">
|
||
<paragraph id="062B36CCFFC4FFA066A3FE30FDC1FE42" blockId="8.[151,603,475,535]" box="[151,603,475,502]" pageId="8" pageNumber="387">
|
||
<heading id="5D6381A0FFC4FFA066A3FE30FDC1FE42" bold="true" box="[151,603,475,502]" fontSize="11" level="1" pageId="8" pageNumber="387" reason="1">
|
||
<taxonomicName id="C1944D4FFFC4FFA066A3FE30FDC1FE42" ID-CoL="725YN" authority="Economidis, 1991" authorityName="Economidis" authorityYear="1991" box="[151,603,475,502]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FE30FDC1FE42" bold="true" box="[151,603,475,502]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FE30FEFFFE41" bold="true" box="[151,357,475,501]" italics="true" pageId="8" pageNumber="387">Leucos ylikiensis</emphasis>
|
||
(Economidis, 1991)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFC4FFA066A3FE15FF67FDA3" blockId="8.[151,603,475,535]" box="[151,253,510,535]" pageId="8" pageNumber="387">
|
||
(
|
||
<figureCitation id="9EAF2A49FFC4FFA066ABFE15FF78FDA3" box="[159,226,510,535]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="8" pageNumber="387">Fig. 2</figureCitation>
|
||
C)
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFC4FFA066A3FDADFC9EFD2E" blockId="8.[151,1435,582,666]" pageId="8" pageNumber="387">
|
||
<taxonomicName id="C1944D4FFFC4FFA066A3FDADFEFBFDE8" box="[151,353,582,604]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="subSpecies" species="aula" subSpecies="rubella">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FDADFEFBFDE8" box="[151,353,582,604]" italics="true" pageId="8" pageNumber="387">Rutilus aula rubella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="C1944D4FFFC4FFA0675CFDACFD36FDE8" authority="Stephanidis, 1939" authorityName="Stephanidis" authorityYear="1939" box="[360,684,582,605]" pageId="8" pageNumber="387" rank="variety" variety="ylikiensis">
|
||
var.
|
||
<emphasis id="34E0EADEFFC4FFA067A6FDADFE6AFDE8" box="[402,496,582,604]" italics="true" pageId="8" pageNumber="387">ylikiensis</emphasis>
|
||
Stephanidis, 1939
|
||
</taxonomicName>
|
||
(infrasubspecific name not available: locality, Lake Yliki)
|
||
<taxonomicName id="C1944D4FFFC4FFA066A3FD8EFDD7FDCF" authority="Stephanidis, 1991: 269" authorityName="Stephanidis" authorityPageNumber="269" authorityYear="1991" box="[151,589,613,635]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FD8EFED3FDCF" box="[151,329,613,635]" italics="true" pageId="8" pageNumber="387">Rutilus ylikiensis</emphasis>
|
||
Stephanidis, 1991: 269
|
||
</taxonomicName>
|
||
. (
|
||
<typeStatus id="D92F886EFFC4FFA06457FD8EFD0DFDCF" box="[611,663,613,635]" pageId="8" pageNumber="387">Type</typeStatus>
|
||
localities: lakes Yliki and Paralimni and River Kifissos). Available by reference to Stephanidis (1991). See
|
||
<bibRefCitation id="62054B3DFFC4FFA06476FD6FFC9EFD2E" author="Kottelat" box="[578,772,644,666]" pageId="8" pageNumber="387" refString="Kottelat, M. (1997) European freshwater fish. Biologia, 52 (Supplement), 1 - 271." type="journal article" year="1997">Kottelat (1997: 82)</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA066A3FD2DFA01FCB7" pageId="8" pageNumber="387" type="materials_examined">
|
||
<paragraph id="062B36CCFFC4FFA066A3FD2DFA01FCB7" blockId="8.[151,1436,710,1959]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FD2DFE1EFD6B" bold="true" box="[151,388,710,735]" pageId="8" pageNumber="387">Examined material.</emphasis>
|
||
<collectionCode id="6085AE09FFC4FFA067B8FD2CFE20FD6B" box="[396,442,711,735]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="8" pageNumber="387">IZA</collectionCode>
|
||
8733, 8,
|
||
<collectingCountry id="7E83765CFFC4FFA06413FD2CFDE3FD6B" box="[551,633,711,735]" name="Greece" pageId="8" pageNumber="387">Greece</collectingCountry>
|
||
, lakes Yliki and Paralimni, P.G. Bianco,
|
||
<date id="722A100CFFC4FFA06273FD2DFB7FFD6B" box="[1095,1253,710,735]" pageId="8" pageNumber="387" value="1987-04-25">25 April 1987</date>
|
||
.—
|
||
<collectionCode id="6085AE09FFC4FFA06333FD2CFAA2FD6B" box="[1287,1336,711,735]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="8" pageNumber="387">IZA</collectionCode>
|
||
0 49, 20,
|
||
<collectingCountry id="7E83765CFFC4FFA066A3FD00FF73FCB7" box="[151,233,747,771]" name="Greece" pageId="8" pageNumber="387">Greece</collectingCountry>
|
||
, L. Yliki, P.G. Bianco,
|
||
<date id="722A100CFFC4FFA067DEFD01FD5FFCB7" box="[490,709,746,771]" pageId="8" pageNumber="387" value="1998-08-11" valueMax="1998-08-16" valueMin="1998-08-11">11–16 August 1998</date>
|
||
.—
|
||
<collectionCode id="6085AE09FFC4FFA064D3FD00FC82FCB7" box="[743,792,747,771]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="8" pageNumber="387">IZA</collectionCode>
|
||
0 479,
|
||
<collectingCountry id="7E83765CFFC4FFA06551FD00FC2DFCB7" box="[869,951,747,771]" name="Greece" pageId="8" pageNumber="387">Greece</collectingCountry>
|
||
, 4, L. Volvi, P.G. Bianco,
|
||
<date id="722A100CFFC4FFA062E3FD00FA0DFCB7" box="[1239,1431,746,771]" pageId="8" pageNumber="387" value="1998-10-21">21 October 1998</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA066F3FCE5FE33FBDF" pageId="8" pageNumber="387" type="diagnosis">
|
||
<paragraph id="062B36CCFFC4FFA066F3FCE5FE33FBDF" blockId="8.[151,1436,710,1959]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066F3FCE5FEDBFC93" bold="true" box="[199,321,782,807]" pageId="8" pageNumber="387">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="C1944D4FFFC4FFA067E8FCE4FDB6FC92" box="[476,556,783,806]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFC4FFA067E8FCE4FDB6FC92" box="[476,556,783,806]" italics="true" pageId="8" pageNumber="387">Leucos</emphasis>
|
||
</taxonomicName>
|
||
characterized by moderate size, not exceeding
|
||
<quantity id="C16C9B29FFC4FFA06277FCE4FB43FC92" box="[1091,1241,783,807]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="387" unit="mm" value="125.0" valueMax="130.0" valueMin="120.0">120–130 mm</quantity>
|
||
SL; absence of a mid-lateral band; peritoneal membrane blackened by dense melanophores; usually 42–43 pored scales on LL, as against
|
||
<quantity id="C16C9B29FFC4FFA066DAFCBDFEC9FCDB" box="[238,339,854,879]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.652" metricValueMin="9.144" pageId="8" pageNumber="387" unit="in" value="37.0" valueMax="38.0" valueMin="36.0">36–38 in</quantity>
|
||
<taxonomicName id="C1944D4FFFC4FFA0676FFCBCFE6EFCDB" box="[347,500,854,879]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFC4FFA0676FFCBCFE6EFCDB" box="[347,500,854,879]" italics="true" pageId="8" pageNumber="387">Leucos basak</emphasis>
|
||
</taxonomicName>
|
||
. It may be distinguished from
|
||
<taxonomicName id="C1944D4FFFC4FFA06561FCBCFC45FCDB" box="[853,991,854,879]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="aula">
|
||
<emphasis id="34E0EADEFFC4FFA06561FCBCFC45FCDB" box="[853,991,854,879]" italics="true" pageId="8" pageNumber="387">Leucos aula</emphasis>
|
||
</taxonomicName>
|
||
for the absence of lateral band and the color of the peritoneal membrane, black in
|
||
<taxonomicName id="C1944D4FFFC4FFA064B6FC97FC92FC27" box="[642,776,890,915]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFC4FFA064B6FC97FC92FC27" box="[642,776,890,915]" italics="true" pageId="8" pageNumber="387">L. ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
and silvery in
|
||
<taxonomicName id="C1944D4FFFC4FFA06587FC97FB9EFC27" box="[947,1028,890,915]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="aula">
|
||
<emphasis id="34E0EADEFFC4FFA06587FC97FB9EFC27" box="[947,1028,890,915]" italics="true" pageId="8" pageNumber="387">L. aula</emphasis>
|
||
</taxonomicName>
|
||
. It differs from all others species of
|
||
<taxonomicName id="C1944D4FFFC4FFA066A3FC74FF7DFC02" box="[151,231,927,950]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FC74FF7DFC02" box="[151,231,927,950]" italics="true" pageId="8" pageNumber="387">Leucos</emphasis>
|
||
</taxonomicName>
|
||
for the high number of total GR, usually
|
||
<quantity id="C16C9B29FFC4FFA064F8FC74FCABFC03" box="[716,817,926,951]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="8" pageNumber="387" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
|
||
<taxonomicName id="C1944D4FFFC4FFA0650EFC74FC59FC03" box="[826,963,926,951]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFC4FFA0650EFC74FC59FC03" box="[826,963,926,951]" italics="true" pageId="8" pageNumber="387">L. ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C16C9B29FFC4FFA065C9FC75FBCEFC03" box="[1021,1108,926,951]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="4.064" metricValueMin="2.032" pageId="8" pageNumber="387" unit="in" value="12.0" valueMax="16.0" valueMin="8.0">8–16 in</quantity>
|
||
all others. The most closely related species, also for the geographical position and according to molecular analyses (
|
||
<figureCitation id="9EAF2A49FFC4FFA06240FC29FB2EFC6F" box="[1140,1204,962,987]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1610,1632]" captionTargetBox="[182,1420,597,1530]" captionTargetId="figure@2.[151,1436,591,1588]" captionTargetPageId="2" captionText="FIGURE 1. A) Collecting localities (L. = lakes, R. = rivers) of Rutilus complex species. 1, R. Thames; 2, R. Danube (Slovakia); 3, R. Danube (Romania); 4, L. Como; 5, L. Maggiore; 6, R. Arno and R. Serchio; 7, R. Albegna and R. Cecina; 8, R. Ombrone and R. Fiora; 9, R. Tiber and R. Mignone; 10, L. Trasimeno; 11, R. Liri and R. Garigliano; 12, R. Volturno and R. Sele; 13, R. Pescara and R. Tavo; 14, R. Trigno; 15, R. Tronto; 16, R. Livenza; 17, L. Pernica; 18, R. Sava; 19, L. Drusino; 20, R. Krupa (R. Neretva basin); 21, L. Besanka (R. Neretva basin); 22, R. Moraca and R. Zeta; 23, L. Skadar; 24, L. Dojran; 25, L. Horid; 26, L. Prespa and L. Micraprespa; 27, L. Vegoritis; 28, L. Volvi; 29, L. Joannina; 30, L. Yliki and L. Paralimni; 31, L. Ambrakia; 32, L. Trichonis. Bold numbers are the different peri-Mediterranean ichthyogeographic districts described in Bianco (1990). B) A schematic temporal reconstruction of the diversification of the genus in southern Mediterranean Europe based on cyt- b is shown (modified from Ketmaier et al., 2008). For each taxon the corresponding number of the district of origin is shown; triangles indicate that multiple populations were analyzed. Numbers at nodes are divergence times in million years; all labeled nodes were robustly supported in the phylogenetic reconstructions (see Ketmaier et al., 2008 for further details)." httpUri="https://zenodo.org/record/225779/files/figure.png" pageId="8" pageNumber="387">Fig. 1</figureCitation>
|
||
B) is
|
||
<taxonomicName id="C1944D4FFFC4FFA062C1FC2FFAC4FC6F" box="[1269,1374,964,987]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="panosi">
|
||
<emphasis id="34E0EADEFFC4FFA062C1FC2FFAC4FC6F" box="[1269,1374,964,987]" italics="true" pageId="8" pageNumber="387">L. panosi</emphasis>
|
||
</taxonomicName>
|
||
from western
|
||
<collectingCountry id="7E83765CFFC4FFA066C1FC0CFEDEFC4B" box="[245,324,999,1023]" name="Greece" pageId="8" pageNumber="387">Greece</collectingCountry>
|
||
. Four specimens from Lake Volvi with 17–18 total GR and 41–42 scales on LL are identified as
|
||
<taxonomicName id="C1944D4FFFC4FFA063B1FC0CFF65FB97" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFC4FFA063B1FC0CFF65FB97" italics="true" pageId="8" pageNumber="387">L. ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
, introduced in this lake similarly to what happened for another endemic species from Lake Yliki,
|
||
<taxonomicName id="C1944D4FFFC4FFA066A3FBC5FD79FBF3" authority="Bianco & Kottelat, 2005" authorityName="Bianco & Kottelat" authorityYear="2005" box="[151,739,1070,1095]" class="Actinopterygii" family="Cyprinidae" genus="Scardinius" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="387" phylum="Chordata" rank="species" species="acarnanicus">
|
||
<emphasis id="34E0EADEFFC4FFA066A3FBC5FE39FBF2" box="[151,419,1070,1095]" italics="true" pageId="8" pageNumber="387">Scardinius acarnanicus</emphasis>
|
||
(
|
||
<bibRefCitation id="62054B3DFFC4FFA06781FBC5FD41FBF3" author="Bianco" box="[437,731,1070,1095]" pageId="8" pageNumber="387" refString="Bianco, P. G. & Kottelat, M. (2005) Scardinius knezevici, a new species of rudd from Lake Skadar, Montenegro (Teleostei: Cyprinidae) Ichthyological Exploration of Freshwaters, 16, 231 - 238." type="journal article" year="2005">Bianco & Kottelat, 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="62054B3DFFC4FFA064C5FBC5FBDCFBF3" author="Bogutskaya" box="[753,1094,1070,1095]" pageId="8" pageNumber="387" refString="Bogutskaya, N. G. & Iliadou, K. (2006) Rutilus panosi, a new roach from western Greece (Teleostei: Cyprinidae). Zoosystematica Rossica, 14, 293 - 298." type="journal article" year="2006">Bogutskaya & Iliadou (2006)</bibRefCitation>
|
||
did not report the species as occurring in Lake Volvi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA066F3FB9DFF36F9D7" pageId="8" pageNumber="387" type="description">
|
||
<paragraph id="062B36CCFFC4FFA066F3FB9DFF36F9D7" blockId="8.[151,1436,710,1959]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066F3FB9DFECBFB3B" bold="true" box="[199,337,1142,1167]" pageId="8" pageNumber="387">Description</emphasis>
|
||
. Body silvery; lateral band absent; color of eye in living fish, yellowish; fins yellowish or pale grayish in preserved specimens; snout pointed; pre-orbital distance near equal to the horizontal diameter of the eye; lips smooth; mouth opening oblique, the corner of maxillae at the same level or a little below; the anteriormost margin of the eye; head length about 3.8–4.2 times in standard length; body depth about
|
||
<quantity id="C16C9B29FFC4FFA06246FB08FB78FB4F" box="[1138,1250,1250,1275]" metricMagnitude="-2" metricUnit="m" metricValue="9.143999999999998" metricValueMax="9.652" metricValueMin="8.636" pageId="8" pageNumber="387" unit="in" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 in</quantity>
|
||
standard length; mouth terminal or slightly inferior; profile of dorsum convex, slightly humped in several cases; paired and unpaired fins yellowish with few scattered melanophores; free margin of dorsal and anal fin concave; caudal fin forked; peritoneal membrane blackened by several melanophores dense and fused; origin of the D at same level of the insertion of the P2; LL complete, and extending from the margin of opercular membrane, to the end of caudal peduncle; 39–45 pored scales, usually 42–43; usually 7.5 row of scales above and 3.5 below the LL; constantly 3 un-branched rays followed by 9 branched rays in the D; constantly 3 un-branched and usually 8 branched rays in the anal fin; P1s with 1 un-branched and 14–15 branched rays; P2 with 1 un-branched ray and constantly 8 branched rays; 16–21 total GR; in adult reproductive males collected in April, there are granular tubercles on the free margin of scales of the body and on the head. For additional description and shape, see
|
||
<figureCitation id="9EAF2A49FFC4FFA0629FF9CDFB77F98B" box="[1195,1261,1574,1599]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="8" pageNumber="387">Fig. 2</figureCitation>
|
||
C and Tables 1, 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA066F3F985FCB5F91F" pageId="8" pageNumber="387" type="distribution">
|
||
<paragraph id="062B36CCFFC4FFA066F3F985FCB5F91F" blockId="8.[151,1436,710,1959]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066F3F985FEFAF933" bold="true" box="[199,352,1646,1671]" pageId="8" pageNumber="387">Distribution.</emphasis>
|
||
Endemic to the two small and adjacent connected lakes Yliki and Paralimni, and River Ksifissos in western
|
||
<collectingCountry id="7E83765CFFC4FFA06726F978FEFFF91F" box="[274,357,1683,1707]" name="Greece" pageId="8" pageNumber="387">Greece</collectingCountry>
|
||
. Introduced in Lake Volvi (present data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA266F3F95DF95CFC12" lastPageId="10" lastPageNumber="389" pageId="8" pageNumber="387" type="discussion">
|
||
<paragraph id="062B36CCFFC4FFA066F3F95DFC91F813" blockId="8.[151,1436,710,1959]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066F3F95DFC90F97B" bold="true" box="[199,778,1718,1743]" pageId="8" pageNumber="387">Remarks on ecology, biology and conservation</emphasis>
|
||
. A preferentially still water species. There are no data available on the species life history. According to local fisherman may reach up to
|
||
<quantity id="C16C9B29FFC4FFA0626EF930FB20F946" box="[1114,1210,1755,1779]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="387" unit="mm" value="200.0">200 mm</quantity>
|
||
TL. The maximum length observed in our materials, collected on April, was for a female of about
|
||
<quantity id="C16C9B29FFC4FFA06226F915FBEAF8A2" box="[1042,1136,1790,1815]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="387" unit="mm" value="150.0">150 mm</quantity>
|
||
TL, aged 3+, with eggs of about 1.0–
|
||
<quantity id="C16C9B29FFC4FFA06724F8C8FEFDF88E" box="[272,359,1827,1851]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="387" unit="mm" value="1.2">1.2 mm</quantity>
|
||
of diameter, all of equal size (a sign of a probable single spawning reproductive habit). Adult males, of
|
||
<quantity id="C16C9B29FFC4FFA06732F8ACFEFEF8EA" box="[262,356,1863,1887]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="387" unit="mm" value="120.0">120 mm</quantity>
|
||
TL, aged, 2+, have fan-shaped minute tubercles on the border of scales, of the body and on the upper part of paired fins. The probable spawning season extends from March to May. Among the IUCN categories, the species is classified as “Endangered” (IUCN, 2013).
|
||
</paragraph>
|
||
<caption id="52EB6644FFC6FFA26643FD80F98AFD34" box="[119,1552,618,640]" pageId="10" pageNumber="389">
|
||
<paragraph id="062B36CCFFC6FFA26643FD80F98AFD34" blockId="10.[119,1552,618,640]" box="[119,1552,618,640]" pageId="10" pageNumber="389">
|
||
<emphasis id="34E0EADEFFC6FFA26643FD80FF40FD34" bold="true" box="[119,218,619,640]" pageId="10" pageNumber="389">TABLE 2</emphasis>
|
||
. Meristic counts and several general features in five species of
|
||
<taxonomicName id="C1944D4FFFC6FFA26576FD81FC1CFD34" box="[834,902,618,640]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFC6FFA26576FD81FC1CFD34" box="[834,902,618,640]" italics="true" pageId="10" pageNumber="389">Leucos</emphasis>
|
||
</taxonomicName>
|
||
. CV: most common values; N= sample size; L. = Lake; R. = River.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="062B36CCFFC6FFA267EEFD02FCE7FCE6" blockId="10.[222,1832,745,934]" pageId="10" pageNumber="389">
|
||
<emphasis id="34E0EADEFFC6FFA267EEFD02F929FD4B" box="[474,1715,745,767]" italics="true" pageId="10" pageNumber="389">
|
||
<taxonomicName id="C1944D4FFFC6FFA267EEFD02FDC6FD4B" box="[474,604,745,767]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="species" species="basak">Leucos basak</taxonomicName>
|
||
<taxonomicName id="C1944D4FFFC6FFA26536FD02FCEDFD4B" box="[770,887,745,767]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="species" species="aula">Leucos aula</taxonomicName>
|
||
<taxonomicName id="C1944D4FFFC6FFA26226FD02FB2FFD4B" box="[1042,1205,745,767]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="species" species="ylikiensis">Leucos ylikiensis</taxonomicName>
|
||
<taxonomicName id="C1944D4FFFC6FFA2630FFD02FA5FFD4B" box="[1339,1477,745,767]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="species" species="panosi">Leucos panosi</taxonomicName>
|
||
<taxonomicName id="C1944D4FFFC6FFA26001FD02F929FD4B" authority="L. Yliki, L. Paralimni, L. Ambrakia, L. Trichonis, R. Krupa, L. Besanka, Several" authorityName="L. Yliki, L. Paralimni, L. Ambrakia, L. Trichonis, R. Krupa, L. Besanka, Several" box="[1589,1715,745,767]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="389" phylum="Chordata" rank="species" species="albus">Leucos albus</taxonomicName>
|
||
</emphasis>
|
||
L. Yliki, L. Paralimni, L. Ambrakia, L. Trichonis, R. Krupa, L. Besanka, Several basins L.Volvi (introduced) eastern L. Joannina (introduced) R. Moraca & Zeta, L. Skadar L. Prespa Micraprespa, Northen
|
||
<collectingCountry id="7E83765CFFC6FFA26565FCD6FCE7FCE6" box="[849,893,829,850]" name="Italy" pageId="10" pageNumber="389">Italy</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFC6FFA266EAFCA0FDE5FCD9" blockId="10.[222,1832,745,934]" pageId="10" pageNumber="389">
|
||
Characters
|
||
<collectingCountry id="7E83765CFFC6FFA26276FCA1FB1CFCEB" box="[1090,1158,842,863]" name="Greece" pageId="10" pageNumber="389">Greece</collectingCountry>
|
||
Western
|
||
<collectingCountry id="7E83765CFFC6FFA263BDFCA1FA57FCEB" box="[1417,1485,842,863]" name="Greece" pageId="10" pageNumber="389">Greece</collectingCountry>
|
||
L.Skadar, L.Ohrid
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFC6FFA2643EFC9FF95CFC12" blockId="10.[222,1832,745,934]" pageId="10" pageNumber="389">N=130 N=152 N=32 N=90 N=11 Range CV Range CV Range CV Range CV Range CV</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |