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<document id="9F45130B48A052C939B1F8A05DAE142B" ID-CLB-Dataset="169982" ID-DOI="10.11646/zootaxa.5195.6.1" ID-GBIF-Dataset="2f972211-63f6-447e-9bdd-2e8965659ee3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7223832" ID-ZooBank="1A42B1B9-9D3F-44D8-B9C7-0D96CB2312ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666164657878" checkinUser="plazi" docAuthor="Kawase, Naoki" docDate="2022" docId="E84587EDFFA40871E7BCFA6C62C0FDFA" docLanguage="en" docName="zootaxa.5195.6.1.pdf" docOrigin="Zootaxa 5195 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psilotreta spatulata Kawase 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="510" masterDocId="147CFF95FFA20878E72BFFA66054FFE3" masterDocTitle="The genus Psilotreta Banks (Trichoptera, Odontoceridae) in Japan" masterLastPageNumber="521" masterPageNumber="501" pageNumber="507" updateTime="1699408918723" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BC87CA2B704610410BA2553C3E7C5BC1">
<mods:title id="D3CF80216506C78B70C28642BB3BE4AD">The genus Psilotreta Banks (Trichoptera, Odontoceridae) in Japan</mods:title>
</mods:titleInfo>
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<mods:namePart id="123988ECB41CD61A620410EE7FC825B2">Kawase, Naoki</mods:namePart>
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<mods:title id="B14D64615EDE6D6EA82FE648DD4CD4F8">Zootaxa</mods:title>
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<mods:part id="5B6E8A6AD043A443F88CF2CF6E171711">
<mods:date id="01CA0B7E26FE086FD96C2AACFDE47D35">2022</mods:date>
<mods:detail id="9A4BC135969C2C81A131BF1BD1F50AF6" type="pubDate">
<mods:number id="2B075C73687D48FFDF6AB2BC48A42A82">2022-10-17</mods:number>
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<mods:number id="B008CA1EE43451A079DB1ED630D34C51">5195</mods:number>
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<mods:classification id="9C79923EA48830CBA08CC3E58315592A">journal article</mods:classification>
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<mods:identifier id="5704ADCEF109D827F95E0705E15C2DAF" type="DOI">10.11646/zootaxa.5195.6.1</mods:identifier>
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<treatment id="E84587EDFFA40871E7BCFA6C62C0FDFA" ID-DOI="http://doi.org/10.5281/zenodo.7231569" ID-GBIF-Taxon="203119556" ID-Zenodo-Dep="7231569" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA" httpUri="http://treatment.plazi.org/id/E84587EDFFA40871E7BCFA6C62C0FDFA" lastPageId="9" lastPageNumber="510" pageId="6" pageNumber="507">
<subSubSection id="28F66570FFA4087EE7BCFA6C61B2FA06" box="[151,486,1482,1509]" pageId="6" pageNumber="507" type="nomenclature">
<paragraph id="605336FBFFA4087EE7BCFA6C61B2FA06" blockId="6.[151,486,1482,1545]" box="[151,486,1482,1509]" pageId="6" pageNumber="507">
<heading id="3B1B8197FFA4087EE7BCFA6C61B2FA06" bold="true" box="[151,486,1482,1509]" fontSize="11" level="1" pageId="6" pageNumber="507" reason="1">
<emphasis id="5298EAE9FFA4087EE7BCFA6C61B2FA06" bold="true" box="[151,486,1482,1509]" pageId="6" pageNumber="507">
<taxonomicName id="A7EC4D78FFA4087EE7BCFA6C61D5FA06" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[151,385,1482,1509]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFA4087EE7BCFA6C61D5FA06" bold="true" box="[151,385,1482,1509]" italics="true" pageId="6" pageNumber="507">Psilotreta spatulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49AB5792FFA4087EE6A3FA6D61B2FA06" box="[392,486,1483,1509]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFA4087EE7BCFA496197F9EA" box="[151,451,1519,1545]" pageId="6" pageNumber="507" type="description">
<paragraph id="605336FBFFA4087EE7BCFA496197F9EA" blockId="6.[151,486,1482,1545]" box="[151,451,1519,1545]" pageId="6" pageNumber="507">
(
<figureCitation id="F8D72A7EFFA4087EE7B4FA496165F9EA" box="[159,305,1519,1545]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Figs 3A–3M</figureCitation>
,
<figureCitation id="F8D72A7EFFA4087EE616FA49612BF9EA" box="[317,383,1519,1545]" captionStart="FIGURE 9" captionStartId="16.[151,250,1855,1879]" captionTargetBox="[219,1369,181,1810]" captionTargetId="figure-25@16.[218,1370,181,1831]" captionTargetPageId="16" captionText="FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9)." figureDoi="http://doi.org/10.5281/zenodo.7223858" httpUri="https://zenodo.org/record/7223858/files/figure.png" pageId="6" pageNumber="507">9F, 9I</figureCitation>
,
<figureCitation id="F8D72A7EFFA4087EE6A7FA4961EEF9EA" box="[396,442,1519,1545]" captionStart="FIGURE 11" captionStartId="18.[151,250,1991,2015]" captionTargetId="figure-25@18.[282,1306,181,1967]" captionTargetPageId="18" captionText="FIGURE 11. Distributions of Japanese Psilotreta species." figureDoi="http://doi.org/10.5281/zenodo.7231471" httpUri="https://zenodo.org/record/7231471/files/figure.png" pageId="6" pageNumber="507">11C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFA4087EE7BCF98B6499F9A5" box="[151,1229,1581,1606]" pageId="6" pageNumber="507" type="reference_group">
<paragraph id="605336FBFFA4087EE7BCF98B6499F9A5" blockId="6.[151,1229,1581,1606]" box="[151,1229,1581,1606]" pageId="6" pageNumber="507">
<taxonomicName id="A7EC4D78FFA4087EE7BCF98B61C9F9A5" box="[151,413,1581,1606]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5298EAE9FFA4087EE7BCF98B614DF9A5" bold="true" box="[151,281,1581,1606]" pageId="6" pageNumber="507">
<emphasis id="5298EAE9FFA4087EE7BCF98B60AFF9A6" bold="true" box="[151,251,1581,1605]" italics="true" pageId="6" pageNumber="507">Psilotreta</emphasis>
sp
</emphasis>
.
<bibRefCitation id="047D4B0AFFA4087EE60DF98861C9F9A5" author="Tsuda, M." box="[294,413,1582,1606]" pageId="6" pageNumber="507" pagination="100 - 104" refId="ref13218" refString="Tsuda, M. (1938) Zur Kenntnis der Trichopteren von Liukiu auf Grund des materials der 1935 Liukiu-Expedition. Transactions of the Biogeographical Society of Japan, 3, 100 - 104." type="journal article" year="1938">Tsuda 1938</bibRefCitation>
</taxonomicName>
, 104, larva, case,
<collectingRegion id="A228F819FFA4087EE579F98862F9F9A5" box="[594,685,1582,1606]" country="Japan" name="Okinawa" pageId="6" pageNumber="507">Okinawa</collectingRegion>
Island;
<bibRefCitation id="047D4B0AFFA4087EE5D5F988632AF9A5" author="Tanida, K." box="[766,894,1582,1606]" pageId="6" pageNumber="507" pagination="370 - 392" refId="ref13046" refString="Tanida, K. (2003) Trichoptera. In: Nishida, M., Shikatani, N. &amp; Shokita, S. (Eds.), The Flora and Fauna of Inland Waters in the Ryukyu Islands. Tokai University Press, Hadano, pp. 370 - 392. [in Japanese]" type="book chapter" year="2003">Tanida 2003</bibRefCitation>
, 374, 380, case,
<collectingRegion id="A228F819FFA4087EE30DF98864D5F9A5" box="[1062,1153,1582,1606]" country="Japan" name="Okinawa" pageId="6" pageNumber="507">Okinawa</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFA4087FE7BCF9C860A4FBBA" lastPageId="7" lastPageNumber="508" pageId="6" pageNumber="507" type="diagnosis">
<paragraph id="605336FBFFA4087EE7BCF9C865C5F8FB" blockId="6.[151,1437,1646,2033]" pageId="6" pageNumber="507">
<emphasis id="5298EAE9FFA4087EE7BCF9C86146F96B" bold="true" box="[151,274,1646,1672]" pageId="6" pageNumber="507">Diagnosis.</emphasis>
The male genitalia of
<taxonomicName id="A7EC4D78FFA4087EE525F9C962C5F96A" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[526,657,1647,1673]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFA4087EE525F9C962C5F96A" box="[526,657,1647,1673]" italics="true" pageId="6" pageNumber="507">P. spatulata</emphasis>
</taxonomicName>
<emphasis id="5298EAE9FFA4087EE5B2F9C962A6F96A" bold="true" box="[665,754,1647,1673]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="49AB5792FFA4087EE5B2F9C962A6F96A" box="[665,754,1647,1673]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to those of
<taxonomicName id="A7EC4D78FFA4087EE319F9C964F4F96B" authorityName="Kawase" authorityYear="2021" box="[1074,1184,1647,1672]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="moritai">
<emphasis id="5298EAE9FFA4087EE319F9C964F4F96B" box="[1074,1184,1647,1672]" italics="true" pageId="6" pageNumber="507">P. moritai</emphasis>
</taxonomicName>
in the shapes of a pair of flattened spatula-shaped lateral processes but can be distinguished from the latter by the following structures: (1) the shape of each lateral process of tergum X is sub-triangular in lateral view in
<taxonomicName id="A7EC4D78FFA4087EE327F91164DAF932" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[1036,1166,1719,1745]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFA4087EE327F91164DAF932" box="[1036,1166,1719,1745]" italics="true" pageId="6" pageNumber="507">P. spatulata</emphasis>
</taxonomicName>
<emphasis id="5298EAE9FFA4087EE3BEF91164B9F932" bold="true" box="[1173,1261,1719,1745]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="49AB5792FFA4087EE3BEF91164B9F932" box="[1173,1261,1719,1745]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F8D72A7EFFA4087EE3D0F911652AF932" box="[1275,1406,1719,1745]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 1" captionStart-2="FIGURE 2" captionStartId-0="8.[151,250,1747,1771]" captionStartId-1="2.[151,250,1685,1709]" captionStartId-2="5.[151,250,1785,1809]" captionTargetBox-0="[225,1361,194,1709]" captionTargetBox-1="[206,1381,196,1647]" captionTargetBox-2="[210,1376,194,1747]" captionTargetId-0="figure-25@8.[225,1362,181,1723]" captionTargetId-1="figure-25@2.[200,1387,181,1660]" captionTargetId-2="figure-16@5.[210,1377,181,1761]" captionTargetPageId-0="8" captionTargetPageId-1="2" captionTargetPageId-2="5" captionText-0="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." captionText-1="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." captionText-2="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi-0="http://doi.org/10.5281/zenodo.7223842" figureDoi-1="http://doi.org/10.5281/zenodo.7223836" figureDoi-2="http://doi.org/10.5281/zenodo.7223838" httpUri-0="https://zenodo.org/record/7223842/files/figure.png" httpUri-1="https://zenodo.org/record/7223836/files/figure.png" httpUri-2="https://zenodo.org/record/7223838/files/figure.png" pageId="6" pageNumber="507">Figs 3I1–3I</figureCitation>
<figureCitation id="F8D72A7EFFA4087EE255F91165DAF932" box="[1406,1422,1719,1745]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="6" pageNumber="507">4</figureCitation>
), but narrow rectangular in
<taxonomicName id="A7EC4D78FFA4087EE693F97D6271F917" authorityName="Kawase" authorityYear="2021" box="[440,549,1755,1780]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="moritai">
<emphasis id="5298EAE9FFA4087EE693F97D6271F917" box="[440,549,1755,1780]" italics="true" pageId="6" pageNumber="507">P. moritai</emphasis>
</taxonomicName>
(
<figureCitation id="F8D72A7EFFA4087EE518F97D62C6F916" box="[563,658,1755,1781]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="6" pageNumber="507">Figs 4C</figureCitation>
<figureCitation id="F8D72A7EFFA4087EE5B9F97D62C8F916" box="[658,668,1755,1781]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[206,1381,196,1647]" captionTargetId="figure-25@2.[200,1387,181,1660]" captionTargetPageId="2" captionText="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223836" httpUri="https://zenodo.org/record/7223836/files/figure.png" pageId="6" pageNumber="507">1</figureCitation>
,
<figureCitation id="F8D72A7EFFA4087EE58CF97D629FF916" box="[679,715,1755,1781]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="6" pageNumber="507">4C</figureCitation>
<figureCitation id="F8D72A7EFFA4087EE5E0F97D628EF916" box="[715,730,1755,1781]" captionStart="FIGURE 2" captionStartId="5.[151,250,1785,1809]" captionTargetBox="[210,1376,194,1747]" captionTargetId="figure-16@5.[210,1377,181,1761]" captionTargetPageId="5" captionText="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223838" httpUri="https://zenodo.org/record/7223838/files/figure.png" pageId="6" pageNumber="507">2</figureCitation>
); and (2) a pair of parameres in the phallus are long and located laterally in
<taxonomicName id="A7EC4D78FFA4087EE63CF95961CEF8FA" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[279,410,1791,1817]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFA4087EE63CF95961CEF8FA" box="[279,410,1791,1817]" italics="true" pageId="6" pageNumber="507">P. spatulata</emphasis>
</taxonomicName>
<emphasis id="5298EAE9FFA4087EE68AF95961AFF8FA" bold="true" box="[417,507,1791,1817]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="49AB5792FFA4087EE68AF95961AFF8FA" box="[417,507,1791,1817]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F8D72A7EFFA4087EE521F95962DBF8FA" box="[522,655,1791,1817]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Figs 3J, 3K</figureCitation>
), but they are short and located ventrally in
<taxonomicName id="A7EC4D78FFA4087EE357F95964BDF8FB" authorityName="Kawase" authorityYear="2021" box="[1148,1257,1791,1816]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="moritai">
<emphasis id="5298EAE9FFA4087EE357F95964BDF8FB" box="[1148,1257,1791,1816]" italics="true" pageId="6" pageNumber="507">P. moritai</emphasis>
</taxonomicName>
(
<figureCitation id="F8D72A7EFFA4087EE3D3F95965D6F8FA" box="[1272,1410,1791,1817]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="6" pageNumber="507">Figs 4D, 4E</figureCitation>
).
</paragraph>
<paragraph id="605336FBFFA4087EE7ECF8846294F812" blockId="6.[151,1437,1646,2033]" pageId="6" pageNumber="507">
<emphasis id="5298EAE9FFA4087EE7ECF884615FF8DF" bold="true" box="[199,267,1826,1852]" pageId="6" pageNumber="507">Adult</emphasis>
(
<figureCitation id="F8D72A7EFFA4087EE630F88561FDF8DE" box="[283,425,1827,1853]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Figs 3A–3C</figureCitation>
). Body and antennae brown, wings light brown in alcohol. Length of each forewing: male
<quantity id="A7149B1EFFA4087EE7BCF8E1614BF882" box="[151,287,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.3" metricValueMin="8.2" pageId="6" pageNumber="507" unit="mm" value="8.75" valueMax="9.3" valueMin="8.2">8.2–9.3 mm</quantity>
(mean = 8.7, n = 10), female
<quantity id="A7149B1EFFA4087EE55EF8E162A9F882" box="[629,765,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.9" metricValueMin="8.2" pageId="6" pageNumber="507" unit="mm" value="9.05" valueMax="9.9" valueMin="8.2">8.2–9.9 mm</quantity>
(mean = 9.3, n = 7). On head dorsally, eyes hemispherical, large especially in male (
<figureCitation id="F8D72A7EFFA4087EE687F8CD6252F866" box="[428,518,1899,1925]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Fig. 3A</figureCitation>
); locations and shapes of setal warts most similar to those of
<taxonomicName id="A7EC4D78FFA4087EE380F8CD6573F867" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1195,1319,1899,1924]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFA4087EE380F8CD6573F867" box="[1195,1319,1899,1924]" italics="true" pageId="6" pageNumber="507">P. japonica</emphasis>
</taxonomicName>
(
<figureCitation id="F8D72A7EFFA4087EE21DF8CD65DAF866" box="[1334,1422,1899,1925]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[206,1381,196,1647]" captionTargetId="figure-25@2.[200,1387,181,1660]" captionTargetPageId="2" captionText="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223836" httpUri="https://zenodo.org/record/7223836/files/figure.png" pageId="6" pageNumber="507">Fig. 1A</figureCitation>
), but pair of anterior warts round oval (narrow transverse in
<taxonomicName id="A7EC4D78FFA4087EE408F82963F4F84B" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[803,928,1935,1960]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFA4087EE408F82963F4F84B" box="[803,928,1935,1960]" italics="true" pageId="6" pageNumber="507">P. japonica</emphasis>
</taxonomicName>
). Venation similar to that of
<taxonomicName id="A7EC4D78FFA4087EE3F4F8296508F84B" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1247,1372,1935,1960]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFA4087EE3F4F8296508F84B" box="[1247,1372,1935,1960]" italics="true" pageId="6" pageNumber="507">P. japonica</emphasis>
</taxonomicName>
(
<figureCitation id="F8D72A7EFFA4087EE241F82960BDF82E" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[206,1381,196,1647]" captionTargetId="figure-25@2.[200,1387,181,1660]" captionTargetPageId="2" captionText="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223836" httpUri="https://zenodo.org/record/7223836/files/figure.png" pageId="6" pageNumber="507">Figs 1B, 1C</figureCitation>
), but crossvein
<emphasis id="5298EAE9FFA4087EE6B6F8156181F82F" box="[413,469,1971,1996]" italics="true" pageId="6" pageNumber="507">m-cu</emphasis>
absent in male hind wings (
<figureCitation id="F8D72A7EFFA4087EE43DF815633AF82E" box="[790,878,1971,1997]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Fig. 3B</figureCitation>
), and base of M3+4 and Cu1a directly connected without cross vein in female forewings (
<figureCitation id="F8D72A7EFFA4087EE570F87162E7F812" box="[603,691,2007,2033]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="6" pageNumber="507">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFF3161B4FDDE" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<emphasis id="5298EAE9FFA5087FE7ECFF316127FF53" bold="true" box="[199,371,150,177]" pageId="7" pageNumber="508">Male genitalia</emphasis>
(
<figureCitation id="F8D72A7EFFA5087FE6A9FF316246FF52" box="[386,530,151,177]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3D–3K</figureCitation>
). Tergum IX elongate, subtriangular in dorsal view, with steep sides above basal setal warts (
<figureCitation id="F8D72A7EFFA5087FE632FF1D619AFF36" box="[281,462,187,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3D, 3E, 3H</figureCitation>
). Basal segment of each inferior appendage as long as preanal appendages, thick, slightly dilated at middle in lateral view, covered with setae; apical segment about 1/2 as long as basal segment, cylindrical with many small brown teeth on apical half (
<figureCitation id="F8D72A7EFFA5087FE5AEFEA56316FEFE" box="[645,834,259,285]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3D, 3G, 3H</figureCitation>
). Preanal appendages elongate, compressed (
<figureCitation id="F8D72A7EFFA5087FE21DFEA565DBFEFE" box="[1334,1423,259,285]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Fig. 3D</figureCitation>
). Median dorsal process of tergum X extending posteriorly, apical half clavate with truncate or round apex in dorsal view (
<figureCitation id="F8D72A7EFFA5087FE7F5FEED613DFE86" box="[222,361,331,357]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3D–3F</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE642FEED612CFE86" box="[361,376,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1785,1809]" captionTargetBox="[210,1376,194,1747]" captionTargetId="figure-16@5.[210,1377,181,1761]" captionTargetPageId="5" captionText="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223838" httpUri="https://zenodo.org/record/7223838/files/figure.png" pageId="7" pageNumber="508">2</figureCitation>
). Pair of lateral processes of tergum X sclerotized, as long as median dorsal process, forming subtriangular plate in lateral view (
<figureCitation id="F8D72A7EFFA5087FE50BFEC9622FFE6A" box="[544,635,367,393]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Fig. 3H</figureCitation>
); each with ventral projection extending antero-ventrad (
<figureCitation id="F8D72A7EFFA5087FE3D2FEC9652AFE6A" box="[1273,1406,367,393]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 1" captionStart-2="FIGURE 2" captionStartId-0="8.[151,250,1747,1771]" captionStartId-1="2.[151,250,1685,1709]" captionStartId-2="5.[151,250,1785,1809]" captionTargetBox-0="[225,1361,194,1709]" captionTargetBox-1="[206,1381,196,1647]" captionTargetBox-2="[210,1376,194,1747]" captionTargetId-0="figure-25@8.[225,1362,181,1723]" captionTargetId-1="figure-25@2.[200,1387,181,1660]" captionTargetId-2="figure-16@5.[210,1377,181,1761]" captionTargetPageId-0="8" captionTargetPageId-1="2" captionTargetPageId-2="5" captionText-0="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." captionText-1="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." captionText-2="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi-0="http://doi.org/10.5281/zenodo.7223842" figureDoi-1="http://doi.org/10.5281/zenodo.7223836" figureDoi-2="http://doi.org/10.5281/zenodo.7223838" httpUri-0="https://zenodo.org/record/7223842/files/figure.png" httpUri-1="https://zenodo.org/record/7223836/files/figure.png" httpUri-2="https://zenodo.org/record/7223838/files/figure.png" pageId="7" pageNumber="508">Figs 3I1–3I</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE255FEC965DAFE6A" box="[1406,1422,367,393]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="7" pageNumber="508">4</figureCitation>
); posterior portion of each lateral process occasionally bearing few denticles ventrally (marked with arrows in
<figureCitation id="F8D72A7EFFA5087FE247FE3560E7FE32" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3I</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE798FE1160EAFE32" box="[179,190,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[206,1381,196,1647]" captionTargetId="figure-25@2.[200,1387,181,1660]" captionTargetPageId="2" captionText="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223836" httpUri="https://zenodo.org/record/7223836/files/figure.png" pageId="7" pageNumber="508">1</figureCitation>
,
<figureCitation id="F8D72A7EFFA5087FE7E5FE1160B1FE32" box="[206,229,439,465]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">3I</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE7CDFE1160A2FE32" box="[230,246,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1785,1809]" captionTargetBox="[210,1376,194,1747]" captionTargetId="figure-16@5.[210,1377,181,1761]" captionTargetPageId="5" captionText="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi="http://doi.org/10.5281/zenodo.7223838" httpUri="https://zenodo.org/record/7223838/files/figure.png" pageId="7" pageNumber="508">2</figureCitation>
), dorsal margins of lateral processes sometimes partially concave beside their intermediate appendages (marked with arrows in
<figureCitation id="F8D72A7EFFA5087FE68BFE7D624FFE16" box="[416,539,475,501]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3E, 3I</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE531FE7D627EFE16" box="[538,554,475,501]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="7" pageNumber="508">4</figureCitation>
). Pair of intermediate appendages dark brown, heavily sclerotized, and strongly curved, C-shaped in lateral view, with acute apex directed posterad (
<figureCitation id="F8D72A7EFFA5087FE486FE5964FDFDFA" box="[941,1193,511,537]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 1" captionStart-2="FIGURE 2" captionStartId-0="8.[151,250,1747,1771]" captionStartId-1="2.[151,250,1685,1709]" captionStartId-2="5.[151,250,1785,1809]" captionTargetBox-0="[225,1361,194,1709]" captionTargetBox-1="[206,1381,196,1647]" captionTargetBox-2="[210,1376,194,1747]" captionTargetId-0="figure-25@8.[225,1362,181,1723]" captionTargetId-1="figure-25@2.[200,1387,181,1660]" captionTargetId-2="figure-16@5.[210,1377,181,1761]" captionTargetPageId-0="8" captionTargetPageId-1="2" captionTargetPageId-2="5" captionText-0="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." captionText-1="FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X." captionText-2="FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. &amp; pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X." figureDoi-0="http://doi.org/10.5281/zenodo.7223842" figureDoi-1="http://doi.org/10.5281/zenodo.7223836" figureDoi-2="http://doi.org/10.5281/zenodo.7223838" httpUri-0="https://zenodo.org/record/7223842/files/figure.png" httpUri-1="https://zenodo.org/record/7223836/files/figure.png" httpUri-2="https://zenodo.org/record/7223838/files/figure.png" pageId="7" pageNumber="508">Figs 3D, 3H, 3I1–3I3</figureCitation>
), sometimes directed posterolaterad (
<figureCitation id="F8D72A7EFFA5087FE66EFD856197FDDE" box="[325,451,547,573]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3E, 3I</figureCitation>
<figureCitation id="F8D72A7EFFA5087FE6E8FD856187FDDE" box="[451,467,547,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1558,1582]" captionTargetBox="[233,1352,361,1521]" captionTargetId="figure-99@10.[219,1368,345,1535]" captionTargetPageId="10" captionText="FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired)." figureDoi="http://doi.org/10.5281/zenodo.7223844" httpUri="https://zenodo.org/record/7223844/files/figure.png" pageId="7" pageNumber="508">4</figureCitation>
).
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFDE162B9FD4A" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
Phallus with phallotheca long, cylindrical; endotheca with pair of long, acute parameres laterally, each apex directed posterolaterad; aedeagus membranous with weakly sclerotized ventral plate, phallotremal sclerite V-shaped in ventral view, strongly curved dorsad (
<figureCitation id="F8D72A7EFFA5087FE570FD2962B4FD4A" box="[603,736,655,681]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3J, 3K</figureCitation>
).
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFD156575FCDA" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<emphasis id="5298EAE9FFA5087FE7ECFD1561C6FD2F" bold="true" box="[199,402,690,717]" pageId="7" pageNumber="508">Female genitalia</emphasis>
(
<figureCitation id="F8D72A7EFFA5087FE68DFD156268FD2E" box="[422,572,691,717]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Figs 3L–3M</figureCitation>
). Sternum IX semicircular to sub-triangular in ventral view with median longitudinal sulcus at posterior 2/3 of sternum IX between pair of indefinite broad pigmentations (
<figureCitation id="F8D72A7EFFA5087FE3E3FD716571FD12" box="[1224,1317,727,753]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Fig. 3M</figureCitation>
). Posterior margin of segment X transverse, almost straight in ventral view; segment X with pair of flattened (depressed) dorsolateral lobes, sparsely setose (
<figureCitation id="F8D72A7EFFA5087FE509FCB9622DFCDA" box="[546,633,799,825]" captionStart="FIGURE 3" captionStartId="8.[151,250,1747,1771]" captionTargetBox="[225,1361,194,1709]" captionTargetId="figure-25@8.[225,1362,181,1723]" captionTargetPageId="8" captionText="FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="7" pageNumber="508">Fig. 3L</figureCitation>
). Length of vaginal apparatus 2 times as long as sternum IX.
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFCE460A4FBBA" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<emphasis id="5298EAE9FFA5087FE7ECFCE461C3FCBE" bold="true" box="[199,407,834,861]" pageId="7" pageNumber="508">Final instar larva</emphasis>
(
<figureCitation id="F8D72A7EFFA5087FE68DFCE56276FCBE" box="[422,546,835,861]" captionStart="FIGURE 9" captionStartId="16.[151,250,1855,1879]" captionTargetBox="[219,1369,181,1810]" captionTargetId="figure-25@16.[218,1370,181,1831]" captionTargetPageId="16" captionText="FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9)." figureDoi="http://doi.org/10.5281/zenodo.7223858" httpUri="https://zenodo.org/record/7223858/files/figure.png" pageId="7" pageNumber="508">Figs 9F, 9I</figureCitation>
). Length up to
<quantity id="A7149B1EFFA5087FE5E7FCE56344FCBE" box="[716,784,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="508" unit="mm" value="9.0">9 mm</quantity>
. Generally similar to those of
<taxonomicName id="A7EC4D78FFA5087FE34AFCE5648AFCBF" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1121,1246,835,860]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFA5087FE34AFCE5648AFCBF" box="[1121,1246,835,860]" italics="true" pageId="7" pageNumber="508">P. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="A7EC4D78FFA5087FE23EFCE565C1FCBE" authorityName="Iwata" authorityYear="1928" box="[1301,1429,835,861]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="kisoensis">
<emphasis id="5298EAE9FFA5087FE23EFCE565C1FCBE" box="[1301,1429,835,861]" italics="true" pageId="7" pageNumber="508">P. kisoensis</emphasis>
</taxonomicName>
, but head and thoracic nota uniformly brown to dark brown without markings. Among primary setae on head, setae 9, 12, 14, and 15 thick, pale brown, other setae fine, white to near transparent; setae 14 longest and 1.5 to 2 times as long as 9; setae 9 second longest and about 1.5 to 2 times as long as 15. Branched gills as in
<figureCitation id="F8D72A7EFFA5087FE3EBFC09655FFC2A" box="[1216,1291,943,969]" captionStart="FIGURE 9" captionStartId="16.[151,250,1855,1879]" captionTargetBox="[219,1369,181,1810]" captionTargetId="figure-25@16.[218,1370,181,1831]" captionTargetPageId="16" captionText="FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9)." figureDoi="http://doi.org/10.5281/zenodo.7223858" httpUri="https://zenodo.org/record/7223858/files/figure.png" pageId="7" pageNumber="508">Fig. 9I</figureCitation>
on each side of abdomen (numbers of branches in parenthesis, n = 3): anterior dorsal gills on segments II (8–10), III (8–14), IV (10), V (7–8), VI (4–7), VII (3–5) and VIII (0-4); anterior ventral gills on segments II (8), III (10–14), IV (8–12), V (8–11), VI (6–7), VII (4–7) and VIII (2–4); anterior lateral gills lacking. Case similar to that of
<taxonomicName id="A7EC4D78FFA5087FE3C0FBBD653EFBD7" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1259,1386,1051,1076]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFA5087FE3C0FBBD653EFBD7" box="[1259,1386,1051,1076]" italics="true" pageId="7" pageNumber="508">P. japonica</emphasis>
</taxonomicName>
, but smaller.
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFA5087FE7ECFBC564D6F876" pageId="7" pageNumber="508" type="materials_examined">
<paragraph id="605336FBFFA5087FE7ECFBC563C1FB42" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<materialsCitation id="D0843CA6FFA5087FE7ECFBC563C1FB42" ID-GBIF-Occurrence="3938587367" collectingDate="1999-03-21" collectionCode="T" collectorName="Ito &amp; A. Ohkawa" county="Male" latitude="26.734167" location="Kunigami-son" longLatPrecision="20" longitude="128.22665" municipality="Iji" pageId="7" pageNumber="508" specimenCode="LBM1410012539" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="5298EAE9FFA5087FE7ECFBC5616FFB9E" bold="true" box="[199,315,1123,1149]" pageId="7" pageNumber="508">
<typeStatus id="BF578859FFA5087FE7ECFBC56162FB9E" box="[199,310,1123,1149]" pageId="7" pageNumber="508" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="89324E77FFA5087FE66FFBC5612AFB9E" box="[324,382,1123,1149]" pageId="7" pageNumber="508">Male</collectingCounty>
(in alcohol),
<emphasis id="5298EAE9FFA5087FE532FBC4628EFB9E" bold="true" box="[537,730,1122,1149]" pageId="7" pageNumber="508">
<collectingRegion id="A228F819FFA5087FE532FBC462D2FB9F" box="[537,646,1122,1148]" country="Japan" name="Okinawa" pageId="7" pageNumber="508">Okinawa</collectingRegion>
Island
</emphasis>
. Okuni-rindo,
<collectingMunicipality id="8037AC81FFA5087FE4AFFBC563F4FB9E" box="[900,928,1123,1149]" pageId="7" pageNumber="508">Iji</collectingMunicipality>
,
<location id="65336020FFA5087FE487FBC5641BFB9E" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE487FBC5641BFB9E" box="[940,1103,1123,1149]" county="Male" latitude="26.734167" longLatPrecision="20" longitude="128.22665" municipality="Iji" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<collectingRegion id="A228F819FFA5087FE377FBC56557FB9E" box="[1116,1283,1123,1149]" country="Japan" name="Okinawa" pageId="7" pageNumber="508">Okinawa Pref.</collectingRegion>
,
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<geoCoordinate id="05D8503CFFA5087FE7BCFB21617CFB42" box="[151,296,1159,1185]" degrees="128" direction="east" minutes="13" orientation="longitude" pageId="7" pageNumber="508" precision="15" seconds="36" value="128.22665">128°13'36&quot;E</geoCoordinate>
,
<date id="1452103BFFA5087FE61FFB2161FBFB42" box="[308,431,1159,1185]" pageId="7" pageNumber="508" value="1999-03-21">
<collectingDate id="0416E9D3FFA5087FE61FFB2161FBFB42" box="[308,431,1159,1185]" pageId="7" pageNumber="508" value="1999-03-21">21.iii.1999</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE690FB216199FB42" box="[443,461,1159,1185]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
.
<collectorName id="CD19532DFFA5087FE6F3FB2161A3FB42" box="[472,503,1159,1185]" pageId="7" pageNumber="508">Ito</collectorName>
and
<collectorName id="CD19532DFFA5087FE507FB2162FFFB42" box="[556,683,1159,1185]" pageId="7" pageNumber="508">A. Ohkawa</collectorName>
(
<specimenCode id="304A9E80FFA5087FE591FB2063DEFB42" box="[698,906,1158,1185]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012539</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFB0D622AFA7E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<materialsCitation id="D0843CA6FFA5087FE7ECFB0D6253FB26" ID-GBIF-Occurrence="3938587369" box="[199,519,1194,1221]" collectorName="Island" location="Okinawa" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="5298EAE9FFA5087FE7ECFB0D6253FB26" bold="true" box="[199,519,1194,1221]" pageId="7" pageNumber="508">
<typeStatus id="BF578859FFA5087FE7ECFB0D6115FB26" box="[199,321,1195,1221]" pageId="7" pageNumber="508" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="A228F819FFA5087FE661FB0C61E3FB27" box="[330,439,1194,1220]" country="Japan" name="Okinawa" pageId="7" pageNumber="508">Okinawa</collectingRegion>
<collectorName id="CD19532DFFA5087FE697FB0D6253FB26" box="[444,519,1195,1221]" pageId="7" pageNumber="508">Island</collectorName>
</emphasis>
</materialsCitation>
:
<materialsCitation id="D0843CA6FFA5087FE538FB0D65D2FB26" ID-GBIF-Occurrence="3938587313" box="[531,1414,1194,1221]" collectingDate="1999-03-21" collectionCode="LBM" collectorName="Ito &amp; A. Ohkawa" county="Male" latitude="26.734167" location="Kunigami-son" longLatPrecision="20" longitude="128.22665" municipality="Iji" pageId="7" pageNumber="508" specimenCode="LBM1410012540, LBM1410012549" specimenCount="10" specimenCount-female="1" specimenCount-male="9" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE538FB0D623DFB26" box="[531,617,1195,1221]" pageId="7" pageNumber="508" type="male">9 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE559FB0D6280FB26" box="[626,724,1195,1221]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
, same data as holotype (
<specimenCode id="304A9E80FFA5087FE4FCFB0C64F0FB26" box="[983,1188,1194,1221]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012540</specimenCode>
–
<specimenCode id="304A9E80FFA5087FE399FB0C652AFB26" box="[1202,1406,1194,1221]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012549</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE2A5FB0D65CCFB0A" ID-GBIF-Occurrence="3938587305" collectingDate="1999-03-22" collectionCode="T" collectorName="Ito &amp; A. Ohkawa" county="Sukuna-gawa" latitude="26.75" location="Kunigami-son" longLatPrecision="1240" longitude="128.21666" municipality="Yona" pageId="7" pageNumber="508" specimenCode="LBM1410012550" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE2A5FB0D6084FB0A" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE7F2FB696120FB0A" box="[217,372,1231,1257]" pageId="7" pageNumber="508">Sukuna-gawa</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE656FB6961EEFB0A" box="[381,442,1231,1257]" pageId="7" pageNumber="508">Yona</collectingMunicipality>
,
<location id="65336020FFA5087FE6E9FB696231FB0A" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE6E9FB696231FB0A" box="[450,613,1231,1257]" county="Sukuna-gawa" latitude="26.75" longLatPrecision="1240" longitude="128.21666" municipality="Yona" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<geoCoordinate id="05D8503CFFA5087FE545FB68629BFB0A" box="[622,719,1230,1257]" degrees="26" direction="north" minutes="45" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.75">26°45'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE5FEFB69636AFB0A" box="[725,830,1231,1257]" degrees="128" direction="east" minutes="13" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.21666">128°13'E</geoCoordinate>
,
<date id="1452103BFFA5087FE463FB696397FB0A" box="[840,963,1231,1257]" pageId="7" pageNumber="508" value="1999-03-22">
<collectingDate id="0416E9D3FFA5087FE463FB696397FB0A" box="[840,963,1231,1257]" pageId="7" pageNumber="508" value="1999-03-22">22.iii.1999</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE4E7FB69638AFB0A" box="[972,990,1231,1257]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
.
<collectorName id="CD19532DFFA5087FE4CCFB696452FB0A" box="[999,1030,1231,1257]" pageId="7" pageNumber="508">Ito</collectorName>
and
<collectorName id="CD19532DFFA5087FE31CFB6964E0FB0A" box="[1079,1204,1231,1257]" pageId="7" pageNumber="508">A. Ohkawa</collectorName>
(
<specimenCode id="304A9E80FFA5087FE3EAFB6865C4FB0A" box="[1217,1424,1230,1257]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012550</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE7BCFB556410FAEE" ID-GBIF-Occurrence="3938587344" box="[151,1092,1266,1293]" collectingDate="2011-04-10" collectionCode="T" collectorName="Ito" county="Sukuna-gawa" latitude="26.75" location="Kunigami-son" longLatPrecision="1240" longitude="128.21666" municipality="Yona" pageId="7" pageNumber="508" specimenCode="LBM1410012551, LBM1410012554" specimenCount="4" specimenCount-male="4" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE7BCFB5560A2FAEE" box="[151,246,1267,1293]" pageId="7" pageNumber="508" type="male">4 males</specimenCount>
, same locality,
<date id="1452103BFFA5087FE69EFB556279FAEE" box="[437,557,1267,1293]" pageId="7" pageNumber="508" value="2011-04-10">
<collectingDate id="0416E9D3FFA5087FE69EFB556279FAEE" box="[437,557,1267,1293]" pageId="7" pageNumber="508" value="2011-04-10">10.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE515FB556204FAEE" box="[574,592,1267,1293]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
.
<collectorName id="CD19532DFFA5087FE54AFB5562D4FAEE" box="[609,640,1267,1293]" pageId="7" pageNumber="508">Ito</collectorName>
(
<specimenCode id="304A9E80FFA5087FE5BEFB546334FAEE" box="[661,864,1266,1293]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012551</specimenCode>
–
<specimenCode id="304A9E80FFA5087FE444FB54646FFAEE" box="[879,1083,1266,1293]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012554</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE37EFB556409FAD2" ID-GBIF-Occurrence="3938587348" collectingDate="2011-04-08" collectionCode="T" collectorName="Ito" latitude="26.6" location="Nago-shi" longLatPrecision="1241" longitude="128.06667" municipality="Genka" pageId="7" pageNumber="508" specimenCode="LBM1410012555, LBM1410012567" specimenCount="13" specimenCount-female="4" specimenCount-male="9" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE37EFB5564E0FAEE" box="[1109,1204,1267,1293]" pageId="7" pageNumber="508" type="male">9 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE3EDFB55656FFAEE" box="[1222,1339,1267,1293]" pageId="7" pageNumber="508" type="female">4 females</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE267FB5565CCFAEE" box="[1356,1432,1267,1293]" pageId="7" pageNumber="508">Genka</collectingMunicipality>
,
<location id="65336020FFA5087FE7BCFAB16155FAD2" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE7BCFAB16155FAD2" box="[151,257,1303,1329]" latitude="26.6" longLatPrecision="1241" longitude="128.06667" municipality="Genka" name="Nago-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Nago-shi</location>
,
<geoCoordinate id="05D8503CFFA5087FE625FAB1613EFAD2" box="[270,362,1303,1329]" degrees="26" direction="north" minutes="36" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.6">26°36'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE658FAB16189FAD2" box="[371,477,1303,1329]" degrees="128" direction="east" minutes="04" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.06667">128°04'E</geoCoordinate>
,
<date id="1452103BFFA5087FE6C1FAB16207FAD2" box="[490,595,1303,1329]" pageId="7" pageNumber="508" value="2011-04-08">
<collectingDate id="0416E9D3FFA5087FE6C1FAB16207FAD2" box="[490,595,1303,1329]" pageId="7" pageNumber="508" value="2011-04-08">8.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE54BFAB16226FAD2" box="[608,626,1303,1329]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
.
<collectorName id="CD19532DFFA5087FE554FAB162CAFAD2" box="[639,670,1303,1329]" pageId="7" pageNumber="508">Ito</collectorName>
(
<specimenCode id="304A9E80FFA5087FE584FAB0632EFAD2" box="[687,890,1302,1329]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012555</specimenCode>
–
<specimenCode id="304A9E80FFA5087FE4A2FAB06401FAD2" box="[905,1109,1302,1329]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="7" pageNumber="508">LBM1410012567</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE341FAB16330FAB6" ID-GBIF-Occurrence="3938587334" collectingDate="1996-04-09" collectionCode="OMNH" collectorName="M. Aoyagi" latitude="26.133333" location="Kunigami-son" longLatPrecision="1243" longitude="128.05" municipality="Benoki" pageId="7" pageNumber="508" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE341FAB164EDFAD2" box="[1130,1209,1303,1329]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE3ECFAB06563FAD2" box="[1223,1335,1302,1329]" pageId="7" pageNumber="508" type="female">5 females</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE26FFAB165CCFAD2" box="[1348,1432,1303,1329]" pageId="7" pageNumber="508">Benoki</collectingMunicipality>
,
<location id="65336020FFA5087FE7BCFA9D616EFAB6" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE7BCFA9D616EFAB6" box="[151,314,1339,1365]" latitude="26.133333" longLatPrecision="1243" longitude="128.05" municipality="Benoki" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<geoCoordinate id="05D8503CFFA5087FE66DFA9D61C0FAB6" box="[326,404,1339,1365]" degrees="26" direction="north" minutes="8" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.133333">26°8'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE6B0FA9D61A3FAB6" box="[411,503,1339,1365]" degrees="128" direction="east" minutes="3" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.05">128°3'E</geoCoordinate>
,
<date id="1452103BFFA5087FE528FA9D6239FAB6" box="[515,621,1339,1365]" pageId="7" pageNumber="508" value="1996-04-09">
<collectingDate id="0416E9D3FFA5087FE528FA9D6239FAB6" box="[515,621,1339,1365]" pageId="7" pageNumber="508" value="1996-04-09">9.iv.1996</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE552FA9D62A4FAB6" box="[633,752,1339,1365]" pageId="7" pageNumber="508">M. Aoyagi</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE42BFA9D6308FAB6" box="[768,860,1339,1365]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE444FA9D627DFA9A" ID-GBIF-Occurrence="3938587360" collectingDate="1996-04-12" collectionCode="OMNH" collectorName="M. Aoyagi" latitude="26.116667" location="Kunigami-son" longLatPrecision="1243" longitude="128.03334" municipality="Hiji river" pageId="7" pageNumber="508" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE444FA9D639CFAB6" box="[879,968,1339,1365]" pageId="7" pageNumber="508" type="male">2 males</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE4FFFA9D646AFAB6" box="[980,1086,1339,1365]" pageId="7" pageNumber="508">Hiji river</collectingMunicipality>
,
<location id="65336020FFA5087FE362FA9D64B8FAB6" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE362FA9D64B8FAB6" box="[1097,1260,1339,1365]" latitude="26.116667" longLatPrecision="1243" longitude="128.03334" municipality="Hiji river" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<geoCoordinate id="05D8503CFFA5087FE3DCFA9D6519FAB6" box="[1271,1357,1339,1365]" degrees="26" direction="north" minutes="7" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.116667">26°7' N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE27FFA9D60FEFA9A" degrees="128" direction="east" minutes="2" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.03334">128°2' E</geoCoordinate>
,
<date id="1452103BFFA5087FE79CFAF96164FA9A" box="[183,304,1375,1401]" pageId="7" pageNumber="508" value="1996-04-12">
<collectingDate id="0416E9D3FFA5087FE79CFAF96164FA9A" box="[183,304,1375,1401]" pageId="7" pageNumber="508" value="1996-04-12">12.iv.1996</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE617FAF961E1FA9A" box="[316,437,1375,1401]" pageId="7" pageNumber="508">M. Aoyagi</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE6EEFAF96275FA9A" box="[453,545,1375,1401]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE51EFAF9622EFA7E" ID-GBIF-Occurrence="3938587332" collectingDate="1996-04-11" collectionCode="OMNH" collectorName="M. Aoyagi" county="Ryukyu University Station" location="Kunigami-son" longitude="128.35" municipality="Yona" pageId="7" pageNumber="508" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="76EAFD72FFA5087FE51EFAF962DBFA9A" box="[565,655,1375,1401]" pageId="7" pageNumber="508" type="male">2 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE5B0FAF96354FA9A" box="[667,768,1375,1401]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE426FAF96468FA9A" box="[781,1084,1375,1401]" pageId="7" pageNumber="508">Ryukyu University Station</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE363FAF964D1FA9A" box="[1096,1157,1375,1401]" pageId="7" pageNumber="508">Yona</collectingMunicipality>
,
<location id="65336020FFA5087FE3BAFAF96560FA9A" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE3BAFAF96560FA9A" box="[1169,1332,1375,1401]" county="Ryukyu University Station" longitude="128.35" municipality="Yona" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
, 26°76'N 128°21'E,
<date id="1452103BFFA5087FE626FA2561D0FA7E" box="[269,388,1411,1437]" pageId="7" pageNumber="508" value="1996-04-11">
<collectingDate id="0416E9D3FFA5087FE626FA2561D0FA7E" box="[269,388,1411,1437]" pageId="7" pageNumber="508" value="1996-04-11">11.iv.1996</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE6A4FA256253FA7E" box="[399,519,1411,1437]" pageId="7" pageNumber="508">M. Aoyagi</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE53DFA256226FA7E" box="[534,626,1411,1437]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="605336FBFFA5087FE7ECFA0064D6F876" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<emphasis id="5298EAE9FFA5087FE7ECFA0062B9FA22" bold="true" box="[199,749,1446,1473]" pageId="7" pageNumber="508">
Other specimens examined.
<collectingRegion id="A228F819FFA5087FE503FA0062C1FA23" box="[552,661,1446,1472]" country="Japan" name="Okinawa" pageId="7" pageNumber="508">Okinawa</collectingRegion>
Island
</emphasis>
:
<materialsCitation id="D0843CA6FFA5087FE429FA016175FA06" ID-GBIF-Occurrence="3938587363" collectingDate="2011-04-08" collectionCode="T" latitude="26.6" location="Nago-shi" longLatPrecision="1241" longitude="128.06667" municipality="Genka" pageId="7" pageNumber="508" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE429FA016302FA22" box="[770,854,1447,1473]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE44CFA016388FA22" box="[871,988,1447,1473]" pageId="7" pageNumber="508" type="female">2 females</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE4C6FA01646DFA22" box="[1005,1081,1447,1473]" pageId="7" pageNumber="508">Genka</collectingMunicipality>
,
<location id="65336020FFA5087FE361FA0164E0FA22" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE361FA0164E0FA22" box="[1098,1204,1447,1473]" latitude="26.6" longLatPrecision="1241" longitude="128.06667" municipality="Genka" name="Nago-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Nago-shi</location>
,
<geoCoordinate id="05D8503CFFA5087FE3EEFA016575FA22" box="[1221,1313,1447,1473]" degrees="26" direction="north" minutes="36" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.6">26°36'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE205FA0165CCFA22" box="[1326,1432,1447,1473]" degrees="128" direction="east" minutes="04" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.06667">128°04'E</geoCoordinate>
,
<date id="1452103BFFA5087FE7BCFA6D6154FA06" box="[151,256,1483,1509]" pageId="7" pageNumber="508" value="2011-04-08">
<collectingDate id="0416E9D3FFA5087FE7BCFA6D6154FA06" box="[151,256,1483,1509]" pageId="7" pageNumber="508" value="2011-04-08">8.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE624FA6D6175FA06" box="[271,289,1483,1509]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE605FA6D6104FA06" ID-GBIF-Occurrence="3938587370" box="[302,336,1483,1509]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE605FA6D6104FA06" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE605FA6D6104FA06" box="[302,336,1483,1509]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE674FA6D6358FA06" ID-GBIF-Occurrence="3938587361" box="[351,780,1483,1509]" collectingDate="2012-03-18" collectionCode="T" location="Ito" pageId="7" pageNumber="508" specimenCount="3" specimenCount-male="3" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE674FA6D61EFFA06" box="[351,443,1483,1509]" pageId="7" pageNumber="508" type="male">3 males</specimenCount>
, same locality,
<date id="1452103BFFA5087FE55BFA6D62BFFA06" box="[624,747,1483,1509]" pageId="7" pageNumber="508" value="2012-03-18">
<collectingDate id="0416E9D3FFA5087FE55BFA6D62BFFA06" box="[624,747,1483,1509]" pageId="7" pageNumber="508" value="2012-03-18">18.iii.2012</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE5D1FA6D6358FA06" box="[762,780,1483,1509]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE431FA6D6368FA06" ID-GBIF-Occurrence="3938587307" box="[794,828,1483,1509]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE431FA6D6368FA06" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE431FA6D6368FA06" box="[794,828,1483,1509]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE461FA6D6175F9EA" ID-GBIF-Occurrence="3938587338" collectingDate="2011-04-08" collectionCode="T" county="Hogen-hashi" location="Nago-shi" municipality="Genka" pageId="7" pageNumber="508" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE461FA6D63F2FA06" box="[842,934,1483,1509]" pageId="7" pageNumber="508" type="male">2 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE49EFA6D6473FA06" box="[949,1063,1483,1509]" pageId="7" pageNumber="508" type="female">2 females</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE31FFA6D6492FA06" box="[1076,1222,1483,1509]" pageId="7" pageNumber="508">Hogen-hashi</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE3FFFA6D6574FA06" box="[1236,1312,1483,1509]" pageId="7" pageNumber="508">Genka</collectingMunicipality>
,
<location id="65336020FFA5087FE205FA6D65CCFA06" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE205FA6D65CCFA06" box="[1326,1432,1483,1509]" county="Hogen-hashi" municipality="Genka" name="Nago-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Nago-shi</location>
,
<date id="1452103BFFA5087FE7BCFA496154F9EA" box="[151,256,1519,1545]" pageId="7" pageNumber="508" value="2011-04-08">
<collectingDate id="0416E9D3FFA5087FE7BCFA496154F9EA" box="[151,256,1519,1545]" pageId="7" pageNumber="508" value="2011-04-08">8.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE624FA496175F9EA" box="[271,289,1519,1545]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE61BFA496106F9EA" ID-GBIF-Occurrence="3938587362" box="[304,338,1519,1545]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE61BFA496106F9EA" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE61BFA496106F9EA" box="[304,338,1519,1545]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE64AFA496368F9EA" ID-GBIF-Occurrence="3938587331" box="[353,828,1519,1545]" collectingDate="2010-11-22" collectingDateMax="2010-11-24" collectingDateMin="2010-11-22" collectionCode="T" location="Ito" pageId="7" pageNumber="508" specimenCount="3" specimenCount-larva="3" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE64AFA496194F9EA" box="[353,448,1519,1545]" pageId="7" pageNumber="508" type="larva">3 larvae</specimenCount>
, same locality,
<date id="1452103BFFA5087FE55CFA496348F9EA" box="[631,796,1519,1545]" pageId="7" pageNumber="508" value="2010-11-22" valueMax="2010-11-24" valueMin="2010-11-22">
<collectingDate id="0416E9D3FFA5087FE55CFA496348F9EA" box="[631,796,1519,1545]" pageId="7" pageNumber="508" value="2010-11-22" valueMax="2010-11-24" valueMin="2010-11-22">22–24.xi.2010</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE401FA496368F9EA" box="[810,828,1519,1545]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE461FA496338F9EA" ID-GBIF-Occurrence="3938587356" box="[842,876,1519,1545]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE461FA496338F9EA" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE461FA496338F9EA" box="[842,876,1519,1545]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE457FA49651AF9EA" ID-GBIF-Occurrence="3938587304" box="[892,1358,1519,1545]" collectingDate="2014-10-17" collectingDateMax="2014-10-19" collectingDateMin="2014-10-17" collectionCode="T" location="Ito" pageId="7" pageNumber="508" specimenCount="6" specimenCount-larva="6" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE457FA49638FF9EA" box="[892,987,1519,1545]" pageId="7" pageNumber="508" type="larva">6 larvae</specimenCount>
, same locality,
<date id="1452103BFFA5087FE3BAFA49657AF9EA" box="[1169,1326,1519,1545]" pageId="7" pageNumber="508" value="2014-10-17" valueMax="2014-10-19" valueMin="2014-10-17">
<collectingDate id="0416E9D3FFA5087FE3BAFA49657AF9EA" box="[1169,1326,1519,1545]" pageId="7" pageNumber="508" value="2014-10-17" valueMax="2014-10-19" valueMin="2014-10-17">17–19.x.2014</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE217FA49651AF9EA" box="[1340,1358,1519,1545]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE276FA49652BF9EA" ID-GBIF-Occurrence="3938587314" box="[1373,1407,1519,1545]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE276FA49652BF9EA" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE276FA49652BF9EA" box="[1373,1407,1519,1545]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE2A5FA4963E7F9CE" ID-GBIF-Occurrence="3938587321" collectingDate="2012-03-18" collectionCode="T" county="Heigi-hashi" latitude="26.75" location="Nago-shi" longLatPrecision="1240" longitude="128.21666" municipality="Genka" pageId="7" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE2A5FA4960B2F9CE" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE7DBF9B56122F9CE" box="[240,374,1555,1581]" pageId="7" pageNumber="508">Heigi-hashi</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE655F9B5619EF9CE" box="[382,458,1555,1581]" pageId="7" pageNumber="508">Genka</collectingMunicipality>
,
<location id="65336020FFA5087FE6FFF9B5626AF9CE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE6FFF9B5626AF9CE" box="[468,574,1555,1581]" county="Heigi-hashi" latitude="26.75" longLatPrecision="1240" longitude="128.21666" municipality="Genka" name="Nago-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Nago-shi</location>
,
<geoCoordinate id="05D8503CFFA5087FE56DF9B462F7F9CE" box="[582,675,1554,1581]" degrees="26" direction="north" minutes="45" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.75">26°45'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE583F9B56346F9CE" box="[680,786,1555,1581]" degrees="128" direction="east" minutes="13" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.21666">128°13'E</geoCoordinate>
,
<date id="1452103BFFA5087FE437F9B563C3F9CE" box="[796,919,1555,1581]" pageId="7" pageNumber="508" value="2012-03-18">
<collectingDate id="0416E9D3FFA5087FE437F9B563C3F9CE" box="[796,919,1555,1581]" pageId="7" pageNumber="508" value="2012-03-18">18.iii.2012</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE48AF9B563E7F9CE" box="[929,947,1555,1581]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE497F9B5638AF9CE" ID-GBIF-Occurrence="3938587343" box="[956,990,1555,1581]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE497F9B5638AF9CE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE497F9B5638AF9CE" box="[956,990,1555,1581]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE4C3F9B561C1F9B2" ID-GBIF-Occurrence="3938587322" collectingDate="1989-01-10" collectionCode="T" latitude="26.933332" location="Nago-shi" longLatPrecision="1239" longitude="128.15" municipality="Teina-gawa" pageId="7" pageNumber="508" specimenCount="1" specimenCount-larva="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE4C3F9B56462F9CE" box="[1000,1078,1555,1581]" pageId="7" pageNumber="508" type="larva">1 larva</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE314F9B56490F9CE" box="[1087,1220,1555,1581]" pageId="7" pageNumber="508">Teina-gawa</collectingMunicipality>
,
<location id="65336020FFA5087FE3E6F9B56563F9CE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE3E6F9B56563F9CE" box="[1229,1335,1555,1581]" latitude="26.933332" longLatPrecision="1239" longitude="128.15" municipality="Teina-gawa" name="Nago-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Nago-shi</location>
,
<geoCoordinate id="05D8503CFFA5087FE26BF9B465C8F9CE" box="[1344,1436,1554,1581]" degrees="26" direction="north" minutes="56" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.933332">26°56'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE7BCF9916155F9B2" box="[151,257,1591,1617]" degrees="128" direction="east" minutes="09" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.15">128°09'E</geoCoordinate>
,
<date id="1452103BFFA5087FE627F991612CF9B2" box="[268,376,1591,1617]" pageId="7" pageNumber="508" value="1989-01-10">
<collectingDate id="0416E9D3FFA5087FE627F991612CF9B2" box="[268,376,1591,1617]" pageId="7" pageNumber="508" value="1989-01-10">10.i.1989</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE6A8F99161C1F9B2" box="[387,405,1591,1617]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE6B4F99162D3F9B2" ID-GBIF-Occurrence="3938587359" box="[415,647,1591,1617]" collectionCode="OMNH" location="Kishimoto" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE6B4F9916241F9B2" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE6B4F9916241F9B2" box="[415,533,1591,1617]" name="Kishimoto" pageId="7" pageNumber="508" stateProvince="Okinawa">Kishimoto</location>
(
<collectionCode id="06FDAE3EFFA5087FE508F991622BF9B2" box="[547,639,1591,1617]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE5B9F991615BF996" ID-GBIF-Occurrence="3938587333" collectingDate="1997-04-12" collectorName="M. Yoshio" latitude="26.133333" location="Kunigami-son" longLatPrecision="1243" longitude="128.05" municipality="Benoki" pageId="7" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE5B9F991628BF9B2" box="[658,735,1591,1617]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE5C1F9916319F9B2" box="[746,845,1591,1617]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE473F99163F8F9B2" box="[856,940,1591,1617]" pageId="7" pageNumber="508">Benoki</collectingMunicipality>
,
<location id="65336020FFA5087FE49DF991640DF9B2" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE49DF991640DF9B2" box="[950,1113,1591,1617]" latitude="26.133333" longLatPrecision="1243" longitude="128.05" municipality="Benoki" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<geoCoordinate id="05D8503CFFA5087FE34FF99164E6F9B2" box="[1124,1202,1591,1617]" degrees="26" direction="north" minutes="8" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.133333">26°8'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE392F9916540F9B2" box="[1209,1300,1591,1617]" degrees="128" direction="east" minutes="3" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.05">128°3'E</geoCoordinate>
,
<date id="1452103BFFA5087FE234F99165CCF9B2" box="[1311,1432,1591,1617]" pageId="7" pageNumber="508" value="1997-04-12">
<collectingDate id="0416E9D3FFA5087FE234F99165CCF9B2" box="[1311,1432,1591,1617]" pageId="7" pageNumber="508" value="1997-04-12">12.iv.1997</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE7BCF9FD615BF996" box="[151,271,1627,1653]" pageId="7" pageNumber="508">M. Yoshio</collectorName>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE631F9FD62E1F996" ID-GBIF-Occurrence="3938587316" box="[282,693,1627,1653]" collectingDate="1997-04-12" collectionCode="T" collectorName="M. Yoshio" latitude="26.133333" location="Kunigami-son" longLatPrecision="1243" longitude="128.05" municipality="Benoki" pageId="7" pageNumber="508" specimenCount="9" specimenCount-male="9" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE631F9FD6127F996" box="[282,371,1627,1653]" pageId="7" pageNumber="508" type="male">9 males</specimenCount>
, same locality,
<date id="1452103BFFA5087FE534F9FD62CCF996" box="[543,664,1627,1653]" pageId="7" pageNumber="508" value="1997-04-12">
<collectingDate id="0416E9D3FFA5087FE534F9FD62CCF996" box="[543,664,1627,1653]" pageId="7" pageNumber="508" value="1997-04-12">12.iv.1997</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE588F9FD62E1F996" box="[675,693,1627,1653]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE5EBF9FD62ABF996" ID-GBIF-Occurrence="3938587320" box="[704,767,1627,1653]" location="Ueda" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE5EBF9FD62ABF996" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE5EBF9FD62ABF996" box="[704,767,1627,1653]" name="Ueda" pageId="7" pageNumber="508" stateProvince="Okinawa">Ueda</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE421F9FD60BEF97A" ID-GBIF-Occurrence="3938587350" collectingDate="2001-04-11" collectingDateMax="2001-04-15" collectingDateMin="2001-04-11" collectorName="K. Uesugi" county="Ie-rindo" location="Kunigami-son" municipality="Sosu" pageId="7" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE421F9FD633AF996" box="[778,878,1627,1653]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE452F9FD6382F996" box="[889,982,1627,1653]" pageId="7" pageNumber="508">Ie-rindo</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE4CAF9FD644EF996" box="[993,1050,1627,1653]" pageId="7" pageNumber="508">Sosu</collectingMunicipality>
,
<location id="65336020FFA5087FE30DF9FD649DF996" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE30DF9FD649DF996" box="[1062,1225,1627,1653]" county="Ie-rindo" municipality="Sosu" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<date id="1452103BFFA5087FE3FFF9FC6521F996" box="[1236,1397,1626,1653]" pageId="7" pageNumber="508" value="2001-04-11" valueMax="2001-04-15" valueMin="2001-04-11">
<collectingDate id="0416E9D3FFA5087FE3FFF9FC6521F996" box="[1236,1397,1626,1653]" pageId="7" pageNumber="508" value="2001-04-11" valueMax="2001-04-15" valueMin="2001-04-11">11–15.iv.2001</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE2AAF9FD60BEF97A" pageId="7" pageNumber="508">K. Uesugi</collectorName>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE7DEF9D96448F97A" ID-GBIF-Occurrence="3938587302" box="[245,1052,1663,1689]" collectingDate="2006-03-27" collectorName="K. Konishi" latitude="26.7" location="Kunigami-son" longLatPrecision="1240" longitude="128.18333" municipality="Hiji" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE7DEF9D96116F97A" box="[245,322,1663,1689]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE666F9D9612FF97A" box="[333,379,1663,1689]" pageId="7" pageNumber="508">Hiji</collectingMunicipality>
,
<location id="65336020FFA5087FE6ADF9D9627DF97A" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE6ADF9D9627DF97A" box="[390,553,1663,1689]" latitude="26.7" longLatPrecision="1240" longitude="128.18333" municipality="Hiji" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
,
<geoCoordinate id="05D8503CFFA5087FE51FF9D962C3F97A" box="[564,663,1663,1689]" degrees="26" direction="north" minutes="42" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.7">26°42' N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE5B6F9D96358F97A" box="[669,780,1663,1689]" degrees="128" direction="east" minutes="11" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.18333">128°11' E</geoCoordinate>
,
<date id="1452103BFFA5087FE43CF9D963C7F97A" box="[791,915,1663,1689]" pageId="7" pageNumber="508" value="2006-03-27">
<collectingDate id="0416E9D3FFA5087FE43CF9D963C7F97A" box="[791,915,1663,1689]" pageId="7" pageNumber="508" value="2006-03-27">27.iii.2006</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE4B5F9D96448F97A" box="[926,1052,1663,1689]" pageId="7" pageNumber="508">K. Konishi</collectorName>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE30CF9D9614BF95E" ID-GBIF-Occurrence="3938587310" collectingDate="2011-04-08" collectionCode="T" collectorName="K. Konishi" latitude="26.7" location="Kunigami-son" longLatPrecision="1240" longitude="128.18333" municipality="Hiji" pageId="7" pageNumber="508" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE30CF9D9642BF97A" box="[1063,1151,1663,1689]" pageId="7" pageNumber="508" type="male">8 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE3A1F9D964ADF97A" box="[1162,1273,1663,1689]" pageId="7" pageNumber="508" type="female">2 females</specimenCount>
, same locality,
<date id="1452103BFFA5087FE7BCF9056154F95E" box="[151,256,1699,1725]" pageId="7" pageNumber="508" value="2011-04-08">
<collectingDate id="0416E9D3FFA5087FE7BCF9056154F95E" box="[151,256,1699,1725]" pageId="7" pageNumber="508" value="2011-04-08">8.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE626F905614BF95E" box="[269,287,1699,1725]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE607F905611AF95E" ID-GBIF-Occurrence="3938587309" box="[300,334,1699,1725]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE607F905611AF95E" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE607F905611AF95E" box="[300,334,1699,1725]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE677F9056332F95E" ID-GBIF-Occurrence="3938587366" box="[348,870,1699,1725]" collectingDate="2011-04-10" collectionCode="T" location="Ito" pageId="7" pageNumber="508" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE677F90561FFF95E" box="[348,427,1699,1725]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE692F905624AF95E" box="[441,542,1699,1725]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
, same locality,
<date id="1452103BFFA5087FE5E4F9056313F95E" box="[719,839,1699,1725]" pageId="7" pageNumber="508" value="2011-04-10">
<collectingDate id="0416E9D3FFA5087FE5E4F9056313F95E" box="[719,839,1699,1725]" pageId="7" pageNumber="508" value="2011-04-10">10.iv.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE47FF9056332F95E" box="[852,870,1699,1725]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE459F90563C0F95E" ID-GBIF-Occurrence="3938587371" box="[882,916,1699,1725]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE459F90563C0F95E" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE459F90563C0F95E" box="[882,916,1699,1725]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE489F905656BF95E" ID-GBIF-Occurrence="3938587358" box="[930,1343,1699,1725]" collectingDate="1999-03-21" collectionCode="T" location="Ito" pageId="7" pageNumber="508" specimenCount="1" specimenCount-larva="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE489F90563A0F95E" box="[930,1012,1699,1725]" pageId="7" pageNumber="508" type="larva">1 larva</specimenCount>
, same locality,
<date id="1452103BFFA5087FE38EF9056574F95E" box="[1189,1312,1699,1725]" pageId="7" pageNumber="508" value="1999-03-21">
<collectingDate id="0416E9D3FFA5087FE38EF9056574F95E" box="[1189,1312,1699,1725]" pageId="7" pageNumber="508" value="1999-03-21">21.iii.1999</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE206F905656BF95E" box="[1325,1343,1699,1725]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE267F905614FF902" ID-GBIF-Occurrence="3938587351" collectorName="A. Ohkawa" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE267F905653FF95E" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE267F905653FF95E" box="[1356,1387,1699,1725]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
and
<collectorName id="CD19532DFFA5087FE7BCF961614FF902" box="[151,283,1735,1761]" pageId="7" pageNumber="508">A. Ohkawa</collectorName>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE60DF9616157F8E6" ID-GBIF-Occurrence="3938587336" collectingDate="1986-04-02" collectionCode="OMNH" collectorName="N. Kobayashi" county="Ryukyu University Station" location="Kunigami-son" longitude="128.35" municipality="Yona" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE60DF9616121F902" box="[294,373,1735,1761]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<collectingCounty id="89324E77FFA5087FE6AAF96162E4F902" box="[385,688,1735,1761]" pageId="7" pageNumber="508">Ryukyu University Station</collectingCounty>
,
<collectingMunicipality id="8037AC81FFA5087FE591F96162A3F902" box="[698,759,1735,1761]" pageId="7" pageNumber="508">Yona</collectingMunicipality>
,
<location id="65336020FFA5087FE428F96163F2F902" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE428F96163F2F902" box="[771,934,1735,1761]" county="Ryukyu University Station" longitude="128.35" municipality="Yona" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
, 26°76'N 128°21'E,
<date id="1452103BFFA5087FE3A0F96164A2F902" box="[1163,1270,1735,1761]" pageId="7" pageNumber="508" value="1986-04-02">
<collectingDate id="0416E9D3FFA5087FE3A0F96164A2F902" box="[1163,1270,1735,1761]" pageId="7" pageNumber="508" value="1986-04-02">2.iv.1986</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE229F96165C8F902" box="[1282,1436,1735,1761]" pageId="7" pageNumber="508">N. Kobayashi</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE7B4F94D60AFF8E6" box="[159,251,1771,1797]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE625F94D62A1F8E6" ID-GBIF-Occurrence="3938587325" box="[270,757,1771,1797]" collectingDate="1996-04-08" collectionCode="OMNH" collectorName="M. Aoyagi" county="Ryukyu University Station" location="Kunigami-son" longitude="128.35" municipality="Yona" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE625F94D610FF8E6" box="[270,347,1771,1797]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
, same locality,
<date id="1452103BFFA5087FE52EF94D6224F8E6" box="[517,624,1771,1797]" pageId="7" pageNumber="508" value="1996-04-08">
<collectingDate id="0416E9D3FFA5087FE52EF94D6224F8E6" box="[517,624,1771,1797]" pageId="7" pageNumber="508" value="1996-04-08">8.iv.1996</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE551F94D62A1F8E6" box="[634,757,1771,1797]" pageId="7" pageNumber="508">M. Aoyagi</collectorName>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE42BF94D61D6F8CA" ID-GBIF-Occurrence="3938587323" collectingDate="1988-10-16" collectionCode="OMNH" collectorName="S. Shokita" location="Kunigami-son" longitude="128.46666" municipality="Aha" pageId="7" pageNumber="508" specimenCount="2" specimenCount-larva="2" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE42BF94D630FF8E6" box="[768,859,1771,1797]" pageId="7" pageNumber="508" type="larva">2 larvae</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE44FF94D63C1F8E6" box="[868,917,1771,1797]" pageId="7" pageNumber="508">Aha</collectingMunicipality>
,
<location id="65336020FFA5087FE48BF94D6417F8E6" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE48BF94D6417F8E6" box="[928,1091,1771,1797]" longitude="128.46666" municipality="Aha" name="Kunigami-son" pageId="7" pageNumber="508" stateProvince="Okinawa">Kunigami-son</location>
, 26°70'N 128°28'E,
<date id="1452103BFFA5087FE20EF94D65C3F8E6" box="[1317,1431,1771,1797]" pageId="7" pageNumber="508" value="1988-10-16">
<collectingDate id="0416E9D3FFA5087FE20EF94D65C3F8E6" box="[1317,1431,1771,1797]" pageId="7" pageNumber="508" value="1988-10-16">16.x.1988</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE7BCF8A96159F8CA" box="[151,269,1807,1833]" pageId="7" pageNumber="508">S. Shokita</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE635F8A9612EF8CA" box="[286,378,1807,1833]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE6A4F8A96439F8CA" ID-GBIF-Occurrence="3938587328" box="[399,1133,1806,1833]" collectingDate="1994-09-05" collectionCode="OMNH" collectorName="M. Aoyagi" location="Kunigami-son" longitude="128.46666" municipality="Aha" pageId="7" pageNumber="508" specimenCount="2" specimenCount-larva="2" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE6A4F8A961BAF8CA" box="[399,494,1807,1833]" pageId="7" pageNumber="508" type="larva">2 larvae</specimenCount>
, 4 cases, same locality,
<date id="1452103BFFA5087FE428F8A8633BF8CA" box="[771,879,1806,1833]" pageId="7" pageNumber="508" value="1994-09-05">
<collectingDate id="0416E9D3FFA5087FE428F8A8633BF8CA" box="[771,879,1806,1833]" pageId="7" pageNumber="508" value="1994-09-05">5.ix.1994</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE455F8A963A3F8CA" box="[894,1015,1807,1833]" pageId="7" pageNumber="508">M. Aoyagi</collectorName>
(
<collectionCode id="06FDAE3EFFA5087FE322F8A96431F8CA" box="[1033,1125,1807,1833]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="508" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE351F8A96200F8AE" ID-GBIF-Occurrence="3938587355" collectingDate="2012-03-17" collectionCode="T" latitude="26.683332" location="Ogimison" longLatPrecision="1240" longitude="128.18333" municipality="Takasato" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE351F8A9649FF8CA" box="[1146,1227,1807,1833]" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE3F3F8A9656AF8CA" box="[1240,1342,1807,1833]" pageId="7" pageNumber="508">Takasato</collectingMunicipality>
,
<location id="65336020FFA5087FE260F8A96094F8AE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE260F8A96094F8AE" latitude="26.683332" longLatPrecision="1240" longitude="128.18333" municipality="Takasato" name="Ogimison" pageId="7" pageNumber="508" stateProvince="Okinawa">Ogimison</location>
,
<geoCoordinate id="05D8503CFFA5087FE7FBF895617BF8AE" box="[208,303,1843,1869]" degrees="26" direction="north" minutes="41" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.683332">26°41'N</geoCoordinate>
,
<geoCoordinate id="05D8503CFFA5087FE614F89561FCF8AE" box="[319,424,1843,1869]" degrees="128" direction="east" minutes="11" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="128.18333">128°11'E</geoCoordinate>
,
<date id="1452103BFFA5087FE69CF8956267F8AE" box="[439,563,1843,1869]" pageId="7" pageNumber="508" value="2012-03-17">
<collectingDate id="0416E9D3FFA5087FE69CF8956267F8AE" box="[439,563,1843,1869]" pageId="7" pageNumber="508" value="2012-03-17">17.iii.2012</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE569F8956200F8AE" box="[578,596,1843,1869]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE548F89562D1F8AE" ID-GBIF-Occurrence="3938587311" box="[611,645,1843,1869]" location="Ito" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE548F89562D1F8AE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE548F89562D1F8AE" box="[611,645,1843,1869]" name="Ito" pageId="7" pageNumber="508" stateProvince="Okinawa">Ito</location>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE5BEF8956147F892" ID-GBIF-Occurrence="3938587353" collectingDate="2004-06-22" latitude="26.472223" location="Kin-cho" longLatPrecision="20" longitude="127.926384" municipality="Okukubi" pageId="7" pageNumber="508" specimenCount="3" specimenCount-larva="1" specimenCount-pupa="2" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE5BEF89562B3F8AE" box="[661,743,1843,1869]" pageId="7" pageNumber="508" type="larva">1 larva</specimenCount>
and
<specimenCount id="76EAFD72FFA5087FE40EF89563D0F8AE" box="[805,900,1843,1869]" pageId="7" pageNumber="508" type="pupa">2 pupae</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE4B8F89563ACF8AE" box="[915,1016,1843,1869]" pageId="7" pageNumber="508">Okukubi</collectingMunicipality>
,
<location id="65336020FFA5087FE323F8956433F8AE" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE323F8956433F8AE" box="[1032,1127,1843,1869]" latitude="26.472223" longLatPrecision="20" longitude="127.926384" municipality="Okukubi" name="Kin-cho" pageId="7" pageNumber="508" stateProvince="Okinawa">Kin-cho</location>
,
<geoCoordinate id="05D8503CFFA5087FE35EF89564AEF8AE" box="[1141,1274,1843,1869]" degrees="26" direction="north" minutes="28" orientation="latitude" pageId="7" pageNumber="508" precision="15" seconds="20" value="26.472223">26°28'20”N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE22EF89465C3F8AE" box="[1285,1431,1842,1869]" degrees="127" direction="east" minutes="55" orientation="longitude" pageId="7" pageNumber="508" precision="15" seconds="35" value="127.926384">127°55'35”E</geoCoordinate>
,
<date id="1452103BFFA5087FE7BCF8F16147F892" box="[151,275,1879,1905]" pageId="7" pageNumber="508" value="2004-06-22">
<collectingDate id="0416E9D3FFA5087FE7BCF8F16147F892" box="[151,275,1879,1905]" pageId="7" pageNumber="508" value="2004-06-22">22.vi.2004</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D0843CA6FFA5087FE634F8F16414F892" ID-GBIF-Occurrence="3938587342" box="[287,1088,1878,1905]" collectingDate="2011-03-01" collectionCode="R" latitude="26.483334" location="Kin-cho" longLatPrecision="1241" longitude="127.916664" municipality="Okukubi" pageId="7" pageNumber="508" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE634F8F1612EF892" box="[287,378,1879,1905]" pageId="7" pageNumber="508" type="male">3 males</specimenCount>
,
<specimenCount id="76EAFD72FFA5087FE6ADF8F161B8F892" box="[390,492,1879,1905]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE6D3F8F16209F892" box="[504,605,1879,1905]" pageId="7" pageNumber="508">Okukubi</collectingMunicipality>
,
<location id="65336020FFA5087FE541F8F1629DF892" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE541F8F1629DF892" box="[618,713,1879,1905]" latitude="26.483334" longLatPrecision="1241" longitude="127.916664" municipality="Okukubi" name="Kin-cho" pageId="7" pageNumber="508" stateProvince="Okinawa">Kin-cho</location>
,
<geoCoordinate id="05D8503CFFA5087FE5FEF8F16366F892" box="[725,818,1879,1905]" degrees="26" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="26.483334">26°29'N</geoCoordinate>
<geoCoordinate id="05D8503CFFA5087FE411F8F063F7F892" box="[826,931,1878,1905]" degrees="127" direction="east" minutes="55" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="127.916664">127°55'E</geoCoordinate>
,
<date id="1452103BFFA5087FE49AF8F16449F892" box="[945,1053,1879,1905]" pageId="7" pageNumber="508" value="2011-03-01">
<collectingDate id="0416E9D3FFA5087FE49AF8F16449F892" box="[945,1053,1879,1905]" pageId="7" pageNumber="508" value="2011-03-01">1.iii.2011</collectingDate>
</date>
,
<collectionCode id="06FDAE3EFFA5087FE301F8F16414F892" box="[1066,1088,1879,1905]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="508">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE367F8F1652AF892" ID-GBIF-Occurrence="3938587368" box="[1100,1406,1878,1905]" location="Kuranishi. Ishigaki Island" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE367F8F1652AF892" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE367F8F1652AF892" box="[1100,1406,1878,1905]" name="Kuranishi. Ishigaki Island" pageId="7" pageNumber="508" stateProvince="Okinawa">
Kuranishi.
<emphasis id="5298EAE9FFA5087FE3E1F8F0652AF892" bold="true" box="[1226,1406,1878,1905]" pageId="7" pageNumber="508">Ishigaki Island</emphasis>
</location>
</materialsCitation>
:
<materialsCitation id="D0843CA6FFA5087FE2A5F8F16442F876" ID-GBIF-Occurrence="3938587346" collectingDate="2002-03-18" collectingDateMax="2002-03-19" collectingDateMin="2002-03-18" collectionCode="T" collectorName="I. Oshima &amp; K. Sugishima" location="Ishigaki-shi" municipality="Omoto-dake" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="76EAFD72FFA5087FE2A5F8F16084F876" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
,
<collectingMunicipality id="8037AC81FFA5087FE7F7F8DD613FF876" box="[220,363,1915,1941]" pageId="7" pageNumber="508">Omoto-dake</collectingMunicipality>
,
<location id="65336020FFA5087FE65DF8DD61A9F876" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE65DF8DD61A9F876" box="[374,509,1915,1941]" municipality="Omoto-dake" name="Ishigaki-shi" pageId="7" pageNumber="508" stateProvince="Okinawa">Ishigaki-shi</location>
,
<date id="1452103BFFA5087FE523F8DD62FAF876" box="[520,686,1915,1941]" pageId="7" pageNumber="508" value="2002-03-18" valueMax="2002-03-19" valueMin="2002-03-18">
<collectingDate id="0416E9D3FFA5087FE523F8DD62FAF876" box="[520,686,1915,1941]" pageId="7" pageNumber="508" value="2002-03-18" valueMax="2002-03-19" valueMin="2002-03-18">18–19.iii.2002</collectingDate>
</date>
,
<collectorName id="CD19532DFFA5087FE592F8DD637EF876" box="[697,810,1915,1941]" pageId="7" pageNumber="508">I. Oshima</collectorName>
,
<collectorName id="CD19532DFFA5087FE41DF8DD639AF876" box="[822,974,1915,1941]" pageId="7" pageNumber="508">K. Sugishima</collectorName>
and
<collectionCode id="06FDAE3EFFA5087FE32FF8DD6442F876" box="[1028,1046,1915,1941]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="508">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="D0843CA6FFA5087FE30BF8DD642AF876" ID-GBIF-Occurrence="3938587308" box="[1056,1150,1915,1941]" location="Yoshida" pageId="7" pageNumber="508" specimenCount="1" stateProvince="Okinawa">
<location id="65336020FFA5087FE30BF8DD642AF876" LSID="urn:lsid:plazi:treatment:E84587EDFFA40871E7BCFA6C62C0FDFA:65336020FFA5087FE30BF8DD642AF876" box="[1056,1150,1915,1941]" name="Yoshida" pageId="7" pageNumber="508" stateProvince="Okinawa">Yoshida</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFA5087FE7ECF83960ABF83E" pageId="7" pageNumber="508" type="etymology">
<paragraph id="605336FBFFA5087FE7ECF83960ABF83E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="508">
<emphasis id="5298EAE9FFA5087FE7ECF839611CF85A" bold="true" box="[199,328,1951,1977]" pageId="7" pageNumber="508">Etymology</emphasis>
. The Latin species epithet
<taxonomicName id="A7EC4D78FFA5087FE55AF839628DF85A" authorityName="Kawase" authorityYear="2022" box="[625,729,1951,1977]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="5298EAE9FFA5087FE55AF839628DF85A" box="[625,729,1951,1977]" italics="true" pageId="7" pageNumber="508">spatulata</emphasis>
</taxonomicName>
is derived from the spatula-shaped lateral processes in the male genitalia.
</paragraph>
</subSubSection>
<caption id="34936673FFAA0870E7BCF97561AFF804" ID-DOI="http://doi.org/10.5281/zenodo.7223842" ID-Zenodo-Dep="7223842" httpUri="https://zenodo.org/record/7223842/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,1747,1771]" targetBox="[225,1361,194,1709]" targetPageId="8">
<paragraph id="605336FBFFAA0870E7BCF97561AFF804" blockId="8.[151,1437,1747,2023]" pageId="8" pageNumber="509">
<emphasis id="5298EAE9FFAA0870E7BCF9756269F908" bold="true" box="[151,573,1747,1771]" pageId="8" pageNumber="509">
FIGURE 3.
<taxonomicName id="A7EC4D78FFAA0870E630F97561B1F908" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[283,485,1747,1771]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFAA0870E630F97561B1F908" bold="true" box="[283,485,1747,1771]" italics="true" pageId="8" pageNumber="509">Psilotreta spatulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49AB5792FFAA0870E6C7F9756269F908" box="[492,573,1747,1771]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4;
<emphasis id="5298EAE9FFAA0870E47BF82E63D7F87C" box="[848,899,1928,1951]" italics="true" pageId="8" pageNumber="509">m-cu</emphasis>
= crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus. Arrows see text.
</paragraph>
</caption>
<subSubSection id="28F66570FFAB0871E7ECFF3060A0FF1A" pageId="9" pageNumber="510" type="distribution">
<paragraph id="605336FBFFAB0871E7ECFF3060A0FF1A" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="510">
<emphasis id="5298EAE9FFAB0871E7ECFF3061AEFF53" bold="true" box="[199,506,150,177]" pageId="9" pageNumber="510">Distribution and habitat.</emphasis>
<taxonomicName id="A7EC4D78FFAB0871E52EFF3162B1FF52" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[517,741,151,177]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFAB0871E52EFF3162B1FF52" box="[517,741,151,177]" italics="true" pageId="9" pageNumber="510">Psilotreta spatulata</emphasis>
</taxonomicName>
<emphasis id="5298EAE9FFAB0871E5DAFF31631AFF52" bold="true" box="[753,846,151,177]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="49AB5792FFAB0871E5DAFF31631AFF52" box="[753,846,151,177]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is an Oriental species distributed in
<collectingRegion id="A228F819FFAB0871E22FFF31653DFF52" box="[1284,1385,151,177]" country="Japan" name="Okinawa" pageId="9" pageNumber="510">Okinawa</collectingRegion>
and Ishigaki Islands in the Ryukyu Archipelago, southwestern
<collectingCountry id="18FB766BFFAB0871E41AFF1D6325FF36" box="[817,881,187,213]" name="Japan" pageId="9" pageNumber="510">Japan</collectingCountry>
(
<figureCitation id="F8D72A7EFFAB0871E4A9FF1D63B2FF36" box="[898,998,187,213]" captionStart="FIGURE 11" captionStartId="18.[151,250,1991,2015]" captionTargetId="figure-25@18.[282,1306,181,1967]" captionTargetPageId="18" captionText="FIGURE 11. Distributions of Japanese Psilotreta species." figureDoi="http://doi.org/10.5281/zenodo.7231471" httpUri="https://zenodo.org/record/7231471/files/figure.png" pageId="9" pageNumber="510">Fig. 11C</figureCitation>
) and was found in small mountainous streams.
</paragraph>
</subSubSection>
<subSubSection id="28F66570FFAB0871E7ECFEA562C0FDFA" pageId="9" pageNumber="510" type="discussion">
<paragraph id="605336FBFFAB0871E7ECFEA56344FE16" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="510">
<emphasis id="5298EAE9FFAB0871E7ECFEA5616FFEFE" bold="true" box="[199,315,259,285]" pageId="9" pageNumber="510">Remarks.</emphasis>
<bibRefCitation id="047D4B0AFFAB0871E66EFEA56189FEFE" author="Tsuda, M." box="[325,477,259,285]" pageId="9" pageNumber="510" pagination="100 - 104" refId="ref13218" refString="Tsuda, M. (1938) Zur Kenntnis der Trichopteren von Liukiu auf Grund des materials der 1935 Liukiu-Expedition. Transactions of the Biogeographical Society of Japan, 3, 100 - 104." type="journal article" year="1938">Tsuda (1938)</bibRefCitation>
recorded larvae and pupae of unidentified
<taxonomicName id="A7EC4D78FFAB0871E4FEFEA564CAFEFE" box="[981,1182,259,285]" pageId="9" pageNumber="510">
<emphasis id="5298EAE9FFAB0871E4FEFEA56416FEFE" box="[981,1090,259,285]" italics="true" pageId="9" pageNumber="510">Psilotreta</emphasis>
species
</taxonomicName>
from
<collectingRegion id="A228F819FFAB0871E3C3FEA56519FEFE" box="[1256,1357,259,285]" country="Japan" name="Okinawa" pageId="9" pageNumber="510">Okinawa</collectingRegion>
Island and
<bibRefCitation id="047D4B0AFFAB0871E7E2FE81613DFEA2" author="Tanida, K." box="[201,361,295,321]" pageId="9" pageNumber="510" pagination="445 - 451" refId="ref12981" refString="Tanida, K. (1997) Trichoptera fauna of the Ryukyu Islands: Taxonomic and ecological prospects. In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 445 - 451." type="book chapter" year="1997">Tanida (1997)</bibRefCitation>
also recorded unidentified larvae and males from
<collectingRegion id="A228F819FFAB0871E49FFE81644DFEA2" box="[948,1049,295,321]" country="Japan" name="Okinawa" pageId="9" pageNumber="510">Okinawa</collectingRegion>
Island. These larvae do not have distinct markings on their heads and thoracic nota as found in previously known Japanese species,
<taxonomicName id="A7EC4D78FFAB0871E3C4FEED6538FE87" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1263,1388,331,356]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="5298EAE9FFAB0871E3C4FEED6538FE87" box="[1263,1388,331,356]" italics="true" pageId="9" pageNumber="510">P. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="A7EC4D78FFAB0871E7BCFEC9614EFE6A" authorityName="Iwata" authorityYear="1928" box="[151,282,367,393]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="kisoensis">
<emphasis id="5298EAE9FFAB0871E7BCFEC9614EFE6A" box="[151,282,367,393]" italics="true" pageId="9" pageNumber="510">P. kisoensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="047D4B0AFFAB0871E601FEC961E7FE6A" author="Tsuda, M." box="[298,435,367,393]" pageId="9" pageNumber="510" pagination="100 - 104" refId="ref13218" refString="Tsuda, M. (1938) Zur Kenntnis der Trichopteren von Liukiu auf Grund des materials der 1935 Liukiu-Expedition. Transactions of the Biogeographical Society of Japan, 3, 100 - 104." type="journal article" year="1938">Tsuda 1938</bibRefCitation>
,
<bibRefCitation id="047D4B0AFFAB0871E694FEC96205FE6A" author="Tanida, K." box="[447,593,367,393]" pageId="9" pageNumber="510" pagination="370 - 392" refId="ref13046" refString="Tanida, K. (2003) Trichoptera. In: Nishida, M., Shikatani, N. &amp; Shokita, S. (Eds.), The Flora and Fauna of Inland Waters in the Ryukyu Islands. Tokai University Press, Hadano, pp. 370 - 392. [in Japanese]" type="book chapter" year="2003">Tanida 2003</bibRefCitation>
). The author examined the male and larval specimens deposited in
<collectingRegion id="A228F819FFAB0871E27DFEC965C8FE6A" box="[1366,1436,367,393]" country="Japan" name="Osaka" pageId="9" pageNumber="510">Osaka</collectingRegion>
Museum of Natural History that were used by
<bibRefCitation id="047D4B0AFFAB0871E580FE35631EFE4E" author="Tanida, K." box="[683,842,403,429]" pageId="9" pageNumber="510" pagination="445 - 451" refId="ref12981" refString="Tanida, K. (1997) Trichoptera fauna of the Ryukyu Islands: Taxonomic and ecological prospects. In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 445 - 451." type="book chapter" year="1997">Tanida (1997)</bibRefCitation>
and confirmed that both the adults and larvae agree with those of
<taxonomicName id="A7EC4D78FFAB0871E612FE1161EBFE32" authority="Kawase, 2022" authorityName="Kawase" authorityYear="2022" box="[313,447,439,465]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="5298EAE9FFAB0871E612FE1161EBFE32" box="[313,447,439,465]" italics="true" pageId="9" pageNumber="510">P. spatulata</emphasis>
</taxonomicName>
<emphasis id="5298EAE9FFAB0871E6E2FE116271FE32" bold="true" box="[457,549,439,465]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="49AB5792FFAB0871E6E2FE116271FE32" box="[457,549,439,465]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
described above. Although the author could not examine specimens used by
<bibRefCitation id="047D4B0AFFAB0871E7BCFE7D6179FE16" author="Tsuda, M." box="[151,301,475,501]" pageId="9" pageNumber="510" pagination="100 - 104" refId="ref13218" refString="Tsuda, M. (1938) Zur Kenntnis der Trichopteren von Liukiu auf Grund des materials der 1935 Liukiu-Expedition. Transactions of the Biogeographical Society of Japan, 3, 100 - 104." type="journal article" year="1938">Tsuda (1938)</bibRefCitation>
, they most probably belong to this species.
</paragraph>
<paragraph id="605336FBFFAB0871E7ECFE5962C0FDFA" blockId="9.[151,1437,150,537]" box="[199,660,511,537]" pageId="9" pageNumber="510">
<emphasis id="5298EAE9FFAB0871E7ECFE5961D7FDFA" bold="true" box="[199,387,511,537]" pageId="9" pageNumber="510">Japanese name.</emphasis>
Okinawa-kiso-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>
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