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<document id="8AA04EC3B4F7FA6EAC213D5C36A3443C" ID-CLB-Dataset="24789" ID-DOI="10.11646/zootaxa.4701.2.2" ID-GBIF-Dataset="2254e402-c6b6-4b90-95d8-587e979af76f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557927" ID-ZooBank="69045F2D-1E16-427B-939E-9F0A6D047D05" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1575043726066" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="C17B87FF3C39123EC0E6FC33FC82F69F" docLanguage="en" docName="zootaxa.4701.2.2.pdf" docOrigin="Zootaxa 4701 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus metallipes Knisch" docType="treatment" docVersion="8" lastPageNumber="157" masterDocId="3D42FF873C3E1236C071FA4AFF92F730" masterDocTitle="Type designations for Neotropical water beetles in the genus Hydrochus Leach 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="167" masterPageNumber="149" pageNumber="156" updateTime="1698769658320" updateUser="ExternalLinkService">
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<mods:title id="D69E0E2C81F46FAFB4BD037EC8541C59">Type designations for Neotropical water beetles in the genus Hydrochus Leach 1817 (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="C17B87FF3C39123EC0E6FC33FC82F69F" ID-DOI="http://doi.org/10.5281/zenodo.5944922" ID-GBIF-Taxon="160605448" ID-Zenodo-Dep="5944922" LSID="urn:lsid:plazi:treatment:C17B87FF3C39123EC0E6FC33FC82F69F" httpUri="http://treatment.plazi.org/id/C17B87FF3C39123EC0E6FC33FC82F69F" lastPageId="8" lastPageNumber="157" pageId="7" pageNumber="156">
<subSubSection id="01C865623C391231C0E6FC33FE6CF1A3" box="[151,510,1657,1684]" pageId="7" pageNumber="156" type="nomenclature">
<paragraph id="496D36E93C391231C0E6FC33FE6CF1A3" blockId="7.[151,510,1657,1720]" box="[151,510,1657,1684]" pageId="7" pageNumber="156">
<heading id="122581853C391231C0E6FC33FE6CF1A3" bold="true" box="[151,510,1657,1684]" fontSize="11" level="1" pageId="7" pageNumber="156" reason="1">
<taxonomicName id="8ED24D6A3C391231C0E6FC33FE6CF1A3" ID-CoL="6MJVD" authority="Knisch" authorityName="Knisch" box="[151,510,1657,1684]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C391231C0E6FC33FE6CF1A3" bold="true" box="[151,510,1657,1684]" pageId="7" pageNumber="156">
<emphasis id="7BA6EAFB3C391231C0E6FC33FE0CF1A4" bold="true" box="[151,414,1657,1684]" italics="true" pageId="7" pageNumber="156">Hydrochus metallipes</emphasis>
Knisch
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01C865623C391231C0E6FCD7FEB0F188" box="[151,290,1693,1720]" pageId="7" pageNumber="156" type="description">
<paragraph id="496D36E93C391231C0E6FCD7FEB0F188" blockId="7.[151,510,1657,1720]" box="[151,290,1693,1720]" pageId="7" pageNumber="156">
<figureCitation id="D1E92A6C3C391231C0E6FCD7FF77F187" box="[151,229,1693,1719]" captionStart="FIGURE 6" captionStartId="8.[151,250,1779,1804]" captionTargetBox="[165,1435,482,1710]" captionTargetId="figure@8.[151,1435,453,1753]" captionTargetPageId="8" captionText="FIGURE 6. Hydrochus metallipes Knisch, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3557939" httpUri="https://zenodo.org/record/3557939/files/figure.png" pageId="7" pageNumber="156">Figs. 6</figureCitation>
,
<figureCitation id="D1E92A6C3C391231C080FCD4FEB0F188" box="[241,290,1694,1720]" captionStart="FIGURES 13" captionStartId="16.[151,264,1786,1811]" captionTargetBox="[168,1423,197,1749]" captionTargetId="figure@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURES 13AE. Labels of lectotypes and neotype. (A) Hydrochus metallipes Knisch; (B) Hydrochus pumilio Knisch; (C) Hydrochus purpureus Knisch; (D) Hydrochus tarsalis Chevrolat; (E) Hydrochus secretus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557953" httpUri="https://zenodo.org/record/3557953/files/figure.png" pageId="7" pageNumber="156">13A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="01C865623C39123EC0E6FCAFFB96F62F" lastPageId="8" lastPageNumber="157" pageId="7" pageNumber="156" type="reference_group">
<paragraph id="496D36E93C391231C0E6FCAFFDF2F030" blockId="7.[151,608,1765,1792]" box="[151,608,1765,1792]" pageId="7" pageNumber="156">
<treatmentCitationGroup id="69C211C73C391231C0E6FCAFFDF2F030" box="[151,608,1765,1792]" pageId="7" pageNumber="156">
<taxonomicName id="8ED24D6A3C391231C0E6FCAFFDCEF030" authority="Knisch 1920: 56" authorityName="Knisch" authorityPageNumber="56" authorityYear="1920" box="[151,604,1765,1792]" class="Aves" family="Apodidae" genus="Hydrochous" kingdom="Animalia" order="Apodiformes" pageId="7" pageNumber="156" phylum="Chordata" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C391231C0E6FCAFFE0BF1CF" box="[151,409,1765,1791]" italics="true" pageId="7" pageNumber="156">Hydrochous metallipes</emphasis>
<treatmentCitation id="C87310F83C391231C1D1FCAFFDCEF030" author="Knisch, A." box="[416,604,1765,1792]" page="56" pageId="7" pageNumber="156" year="1920">
<bibRefCitation id="2D434B183C391231C1D1FCAFFDCEF030" author="Knisch, A." box="[416,604,1765,1792]" pageId="7" pageNumber="156" pagination="55 - 89" refId="ref5572" refString="Knisch, A. (1920) Die exotischen Hydrophiliden des Deutschen Entomologischen Museum (Col.). Archiv fur Naturegeschichte, 85 A, 55 - 89." type="journal article" year="1920">Knisch 1920: 56</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="496D36E93C39123EC0E6FD67FB96F62F" blockId="7.[151,1437,1837,2044]" lastBlockId="8.[151,1437,152,431]" lastPageId="8" lastPageNumber="157" pageId="7" pageNumber="156">
The RBINS kindly provided eighteen
<typeStatus id="9669884B3C391231C235FD64FD34F078" box="[580,678,1838,1864]" pageId="7" pageNumber="156" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="8ED24D6A3C391231C2BDFD67FB8BF077" ID-CoL="6MJVD" authority="Knisch" authorityName="Knisch" box="[716,1049,1837,1863]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C391231C2BDFD67FC52F077" box="[716,960,1837,1863]" italics="true" pageId="7" pageNumber="156">Hydrochus metallipes</emphasis>
Knisch
</taxonomicName>
, which are part of the Orchymont/ Knisch types collection. The
<typeStatus id="9669884B3C391231C1AEFD18FDD3F05C" box="[479,577,1874,1900]" pageId="7" pageNumber="156" type="syntype">syntypes</typeStatus>
consist of both males and females. I have selected one of the males, a specimen in good condition, to serve as
<typeStatus id="9669884B3C391231C251FD3FFD18F0BF" box="[544,650,1909,1935]" pageId="7" pageNumber="156" type="lectotype">lectotype</typeStatus>
, have labeled it such, and have imaged the habitus, male genitalia and labels of this specimen (
<figureCitation id="D1E92A6C3C391231C1D8FDD3FE6BF083" box="[425,505,1945,1971]" captionStart="FIGURE 6" captionStartId="8.[151,250,1779,1804]" captionTargetBox="[165,1435,482,1710]" captionTargetId="figure@8.[151,1435,453,1753]" captionTargetPageId="8" captionText="FIGURE 6. Hydrochus metallipes Knisch, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3557939" httpUri="https://zenodo.org/record/3557939/files/figure.png" pageId="7" pageNumber="156">Figs. 6</figureCitation>
,
<figureCitation id="D1E92A6C3C391231C277FDD0FDA8F084" box="[518,570,1946,1972]" captionStart="FIGURES 13" captionStartId="16.[151,264,1786,1811]" captionTargetBox="[168,1423,197,1749]" captionTargetId="figure@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURES 13AE. Labels of lectotypes and neotype. (A) Hydrochus metallipes Knisch; (B) Hydrochus pumilio Knisch; (C) Hydrochus purpureus Knisch; (D) Hydrochus tarsalis Chevrolat; (E) Hydrochus secretus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557953" httpUri="https://zenodo.org/record/3557953/files/figure.png" pageId="7" pageNumber="156">13A</figureCitation>
). The seventeen non-lectotype
<typeStatus id="9669884B3C391231C3E8FDD0FC69F084" box="[921,1019,1946,1972]" pageId="7" pageNumber="156" type="syntype">syntypes</typeStatus>
have been labeled as
<typeStatus id="9669884B3C391231C485FDD3FA0BF083" box="[1268,1433,1945,1971]" pageId="7" pageNumber="156" type="paralectotype">paralectotypes</typeStatus>
.
<typeStatus id="9669884B3C391231C0E6FDF4FE95F0E8" box="[151,263,1982,2008]" pageId="7" pageNumber="156" type="lectotype">Lectotype</typeStatus>
labels (
<figureCitation id="D1E92A6C3C391231C12EFDF7FE5AF0E8" box="[351,456,1981,2008]" captionStart="FIGURES 13" captionStartId="16.[151,264,1786,1811]" captionTargetBox="[168,1423,197,1749]" captionTargetId="figure@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURES 13AE. Labels of lectotypes and neotype. (A) Hydrochus metallipes Knisch; (B) Hydrochus pumilio Knisch; (C) Hydrochus purpureus Knisch; (D) Hydrochus tarsalis Chevrolat; (E) Hydrochus secretus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557953" httpUri="https://zenodo.org/record/3557953/files/figure.png" pageId="7" pageNumber="156">Fig. 13A</figureCitation>
): “Bras.Corumba [Matto Grosso] // det. Kniž
<emphasis id="7BA6EAFB3C391231C3BBFDF7FBAFF0E7" box="[970,1085,1981,2007]" italics="true" pageId="7" pageNumber="156">metallipes</emphasis>
m. // Coll. A.Knisch TYPUS // coll. A. dOrchymont:
<taxonomicName id="8ED24D6A3C391231C1E4FDABFD9DF0CB" ID-CoL="8VRRD" authorityName="Leach" authorityYear="1817" box="[405,527,2017,2043]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="7BA6EAFB3C391231C1E4FDABFD9DF0CB" box="[405,527,2017,2043]" italics="true" pageId="7" pageNumber="156">Hydrochus</emphasis>
</taxonomicName>
R.M.H.N.B. 15.962 // TYPE // HABITUS/GENIT. DIGITAL IMAGES captured 2019 P.D. Perkins // [red]
<typeStatus id="9669884B3C36123EC1B4FAD3FDF7F783" box="[453,613,153,179]" pageId="8" pageNumber="157" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="8ED24D6A3C36123EC21CFAD3FC29F783" ID-CoL="6MJVD" authority="Knisch" authorityName="Knisch" box="[621,955,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C36123EC21CFAD3FCF1F783" box="[621,867,153,179]" italics="true" pageId="8" pageNumber="157">Hydrochus metallipes</emphasis>
Knisch
</taxonomicName>
desig. P. D. Perkins //
<taxonomicName id="8ED24D6A3C36123EC4B4FAD3FEDCF7E7" ID-CoL="3N6CM" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="7BA6EAFB3C36123EC4B4FAD3FA0EF783" box="[1221,1436,153,179]" italics="true" pageId="8" pageNumber="157">Hydrochus tarsalis</emphasis>
<bibRefCitation id="2D434B183C36123EC0E6FAF7FEDCF7E7" author="Chevrolat, A." box="[151,334,189,215]" pageId="8" pageNumber="157" pagination="183 - 210" refId="ref5281" refString="Chevrolat, A. (1863) Coleopteres del l'ile de Cuba (suite). Notes, synonymies et descriptions d'especes Nouvelles. Familles des Cicindeletes, Carabiques, Dytiscides, Gyrinides et Palpicornes. Annales de la Societe entomologique de France, 4 eme Serie, 3, 1863, 183 - 210. https: // doi. org / 10.1080 / 00379271.1863.11755444" type="journal article" year="1863">Chevrolat, 1863</bibRefCitation>
</taxonomicName>
, det. P. D. Perkins, 2019.” The male genitalia of the
<typeStatus id="9669884B3C36123EC3FDFAF7FC61F7E7" box="[908,1011,189,215]" pageId="8" pageNumber="157" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="8ED24D6A3C36123EC467FAF7FA0EF7E7" ID-CoL="6MJVD" authority="Knisch 1920" authorityName="Knisch" authorityYear="1920" box="[1046,1436,189,215]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C36123EC467FAF7FA9BF7E7" box="[1046,1289,189,215]" italics="true" pageId="8" pageNumber="157">Hydrochus metallipes</emphasis>
<bibRefCitation id="2D434B183C36123EC57EFAF7FA0EF7E7" author="Knisch, A." box="[1295,1436,189,215]" pageId="8" pageNumber="157" pagination="55 - 89" refId="ref5572" refString="Knisch, A. (1920) Die exotischen Hydrophiliden des Deutschen Entomologischen Museum (Col.). Archiv fur Naturegeschichte, 85 A, 55 - 89." type="journal article" year="1920">Knisch 1920</bibRefCitation>
</taxonomicName>
does not differ significantly from the male genitalia of the
<typeStatus id="9669884B3C36123EC350FAABFCE8F7CB" box="[801,890,225,251]" pageId="8" pageNumber="157" type="neotype">neotype</typeStatus>
of
<taxonomicName id="8ED24D6A3C36123EC3EFFAABFABBF7CB" ID-CoL="3N6CM" authority="Chevrolat 1863" authorityName="Chevrolat" authorityYear="1863" box="[926,1321,225,251]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="7BA6EAFB3C36123EC3EFFAABFBE0F7CB" box="[926,1138,225,251]" italics="true" pageId="8" pageNumber="157">Hydrochus tarsalis</emphasis>
<bibRefCitation id="2D434B183C36123EC408FAABFABBF7CB" author="Chevrolat, A." box="[1145,1321,225,251]" pageId="8" pageNumber="157" pagination="183 - 210" refId="ref5281" refString="Chevrolat, A. (1863) Coleopteres del l'ile de Cuba (suite). Notes, synonymies et descriptions d'especes Nouvelles. Familles des Cicindeletes, Carabiques, Dytiscides, Gyrinides et Palpicornes. Annales de la Societe entomologique de France, 4 eme Serie, 3, 1863, 183 - 210. https: // doi. org / 10.1080 / 00379271.1863.11755444" type="journal article" year="1863">Chevrolat 1863</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="8ED24D6A3C36123EC542FAA8FF5EF62F" ID-CoL="6MJVD" authority="Knisch 1920" authorityName="Knisch" authorityYear="1920" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C36123EC542FAA8FF5EF62F" italics="true" pageId="8" pageNumber="157">H. metallipes</emphasis>
</taxonomicName>
is therefore reduced to status of junior synonym of
<taxonomicName id="8ED24D6A3C36123EC361FB4CFB96F62F" ID-CoL="3N6CM" authority="Chevrolat." authorityName="Chevrolat." box="[784,1028,261,287]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="7BA6EAFB3C36123EC361FB4CFC14F62F" box="[784,902,261,287]" italics="true" pageId="8" pageNumber="157">H. tarsalis</emphasis>
Chevrolat.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="01C865623C36123EC0B6FB63FC82F69F" pageId="8" pageNumber="157" type="discussion">
<paragraph id="496D36E93C36123EC0B6FB63FC82F69F" blockId="8.[151,1437,152,431]" pageId="8" pageNumber="157">
<emphasis id="7BA6EAFB3C36123EC0B6FB63FEA9F673" bold="true" box="[199,315,297,323]" pageId="8" pageNumber="157">Remarks.</emphasis>
<bibRefCitation id="2D434B183C36123EC130FB63FE42F673" author="Oliva, A." box="[321,464,297,323]" pageId="8" pageNumber="157" pagination="87 - 104" refId="ref6088" refString="Oliva, A. (1992) The species of Hydrochus (Coleoptera; Hydrophiloidea; Hydrochidae) described from South America. Bulletin et annales de la Societe royale belge d'entomologie, 128, 87 - 104." type="journal article" year="1992">Oliva (1992)</bibRefCitation>
published a drawing of the male genitalia of, supposedly,
<taxonomicName id="8ED24D6A3C36123EC427FB60FB79F673" box="[1110,1259,297,323]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="metallipes">
<emphasis id="7BA6EAFB3C36123EC427FB60FB79F673" box="[1110,1259,297,323]" italics="true" pageId="8" pageNumber="157">H. metallipes</emphasis>
</taxonomicName>
. The parameres are depicted as having a more pronounced enlargement of the medial margin than is present in the
<typeStatus id="9669884B3C36123EC483FB07FACEF657" box="[1266,1372,333,359]" pageId="8" pageNumber="157" type="lectotype">lectotype</typeStatus>
, or in all specimens of
<taxonomicName id="8ED24D6A3C36123EC122FB38FE5AF6BB" box="[339,456,369,395]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="7BA6EAFB3C36123EC122FB38FE5AF6BB" box="[339,456,369,395]" italics="true" pageId="8" pageNumber="157">H. tarsalis</emphasis>
</taxonomicName>
for which the male genitalia have been examined.
<bibRefCitation id="2D434B183C36123EC38FFB3BFB1CF6BB" author="Oliva, A." box="[1022,1166,369,395]" pageId="8" pageNumber="157" pagination="87 - 104" refId="ref6088" refString="Oliva, A. (1992) The species of Hydrochus (Coleoptera; Hydrophiloidea; Hydrochidae) described from South America. Bulletin et annales de la Societe royale belge d'entomologie, 128, 87 - 104." type="journal article" year="1992">Oliva (1992)</bibRefCitation>
did not give the locality data of the specimen that was used to make the drawing.
</paragraph>
</subSubSection>
</treatment>
</document>