188 lines
21 KiB
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188 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="9742492B37265D68AFFB450AED24E5A4" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Rhaphaulus chrysalis" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199839046" LSID="urn:lsid:plazi:treatment:9742492B37265D68AFFB450AED24E5A4" httpUri="http://treatment.plazi.org/id/9742492B37265D68AFFB450AED24E5A4" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="9742492B-3726-5D68-AFFB-450AED24E5A4" authority="(Pfeiffer, 1853)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Rhaphaulus chrysalis (Pfeiffer, 1853)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">Fig. 14B-D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="L. Pfeiffer" authorityYear="1854" class="Gastropoda" family="Pupinidae" genus="Cyclostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclostoma chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Cyclostoma chrysalis</taxonomicName>
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Pfeiffer, 1853: 239, pl. 31, figs 23, 24. Type locality: Arva [Mandalay Region, Myanmar].
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="156 - 160" refId="B122" refString="Pfeiffer, L, 1854. Descriptions of eighteen new species of land-shells, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London 20 (1852): 156 - 160" title="Descriptions of eighteen new species of land-shells, from the collection of H. Cuming, Esq." volume="20" year="1854">Pfeiffer 1854</bibRefCitation>
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: 158.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Rhaphaulus chrysalis</taxonomicName>
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-
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="245 - 254" refId="B167" refString="Theobald, W, 1858. Notes on the distribution of some of the land and freshwater shells of India. Journal of the Asiatic Society of Bengal 26 (1857): 245 - 254" title="Notes on the distribution of some of the land and freshwater shells of India." volume="26" year="1858">Theobald 1858</bibRefCitation>
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[1857]: 247, Maulmein [Mawlamyine, Mawlamyine Township, Mawlamyine District, Mon State, Myanmar]. Sowerby I 1866:
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<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
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, pl. 2 (pl. 264),
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<taxonomicName authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhaphaulus</taxonomicName>
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, figs 6, 7, Siam.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14456" author="Hanley, SCT" journalOrPublisher="Annals of the Lyceum of Natural History of New York" pageId="0" pageNumber="1" refId="B65" refString="Hanley, SCT, Theobald, W (1870, 1875, [1870-1876]) Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London, i-xviii, + 1-65. [pp. 1-18, pls, 1-40, 1870. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. Londoni-xviii+ 1-65. [pp. 1-18pls1-40 (1870) ; pp. 49-56, pls 121-140 (1875)] [Published in parts, dates follow Prashad (1927)] https://doi.org/10.5962/bhl.title.14456" title="Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. Londoni-xviii + 1 - 65. [pp. 1 - 18 pls 1 - 40 (1870); pp. 49 - 56, pls 121 - 140 (1875)] [Published in parts, dates follow Prashad (1927)]" url="https://doi.org/10.5962/bhl.title.14456" year="1870">Hanley and Theobald 1875</bibRefCitation>
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: 53, pl. 133, fig. 7.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
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: 301.
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<bibRefCitation author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve & Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve & Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
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:
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<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
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, pl. 10, sp. 95.
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<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B50" refString="Godwin-Austen, HH, 1886. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 1, Part 5: 165-205." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 1, Part 5: 165 - 205." year="1886">Godwin-Austen 1886</bibRefCitation>
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: 200, 201, pl. 47, fig. 1, 1a.
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<bibRefCitation author="Tapparone-Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="295 - 359" refId="B161" refString="Tapparone-Canefri, C, 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine, XVIII molluschi terrestri e d'acqua dolce. Annali del Museo Civico di Storia Naturale di Genova 27: 295 - 359" title="Viaggio di Leonardo Fea in Birmania e regioni vicine, XVIII molluschi terrestri e d'acqua dolce." volume="27" year="1889">Tapparone-Canefri 1889</bibRefCitation>
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: 310.
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<bibRefCitation author="Smith, EA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="17 - 19" refId="B145" refString="Smith, EA, 1898. Description of Rhaphaulus perakensis, n.sp., with a list of the known species of the genus. Proceedings of the Malacological Society of London 3: 17 - 19" title="Description of Rhaphaulus perakensis, n. sp., with a list of the known species of the genus." volume="3" year="1898">Smith 1898</bibRefCitation>
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: 19.
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 275, 276.
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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: 165, 166, fig. 24.
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2014
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</bibRefCitation>
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: 572, fig. 11, north-eastern India and Myanmar.
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<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
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: 95.
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<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
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: 157.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="L.Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Raphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphaulus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Raphaulus</taxonomicName>
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[sic]
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<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1854" class="Gastropoda" family="Pupinidae" genus="Raphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Raphaulus chrysalis</taxonomicName>
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-
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<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="143 - 259" refId="B150" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 259" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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: 151, farm caves, near Moulmein, Myanmar.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Possible syntype</emphasis>
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NHMUK 2013.04.16 (1 shell; Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14B</figureCitation>
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) from Siam.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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NHMUK 1871.9.23.52 (1 shell; Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14C</figureCitation>
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) from
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<collectingCountry name="Myanmar">Burma</collectingCountry>
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. NHMUK 1903.7.1.3073 (2 shells; Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14D</figureCitation>
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) from Molmein.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate; body whorls slightly bulging. Tube cylindrical, pointing upward and backward.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhaphaulus chrysalis</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName lsidName="R. lorraini" pageId="0" pageNumber="1" rank="species" species="lorraini">
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<emphasis italics="true" pageId="0" pageNumber="1">R. lorraini</emphasis>
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</taxonomicName>
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in shell shape, but differs in having a cylindrical tube pointing upward and backward, instead of forward as in
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<taxonomicName lsidName="R. lorraini" pageId="0" pageNumber="1" rank="species" species="lorraini">
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<emphasis italics="true" pageId="0" pageNumber="1">R. lorraini</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Northeastern India, Myanmar, and an uncertain record from Thailand (
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2014
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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No material of this species was found during this survey, and the record in Thailand needs further confirmation. The type locality on the label of the possible type specimen is
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<normalizedToken originalValue="“Siam”">"Siam"</normalizedToken>
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, which is different from that reported in the original description as
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<normalizedToken originalValue="“Arva”">"Arva"</normalizedToken>
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. A lack of a tube in a possible syntype NHMUK 2013.04.16 (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14B</figureCitation>
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) is possibly due to damage.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |