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<document id="4C0133BBE2FC0045A5FDEA02351EB009" ID-CLB-Dataset="5523" ID-DOI="10.11646/zootaxa.133.1.1" ID-GBIF-Dataset="8e961a32-fadb-471a-bf45-51eeaa44d4df" ID-ISSN="1175­5334" ID-Zenodo-Dep="5014171" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624381892612" checkinUser="felipe" docAuthor="Boyer, Sarah L. &amp; Giribet, Gonzalo" docDate="2003" docId="97070A43FF975A2A5708F9FAB9DCADEB" docLanguage="en" docName="zt00133.pdf" docOrigin="Zootaxa 133 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.133.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Rakaia Hirst 1925" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="6B3E723BFF965A265600FFC3BB32AC4E" masterDocTitle="A new Rakaia species (Opiliones, Cyphophthalmi, Pettalidae) from Otago, New Zealand" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1699020046264" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="81AF8375D137E73C0FF8012C05F811B7">
<mods:title id="81EDB9E57C393856893B45185A12A890">A new Rakaia species (Opiliones, Cyphophthalmi, Pettalidae) from Otago, New Zealand</mods:title>
</mods:titleInfo>
<mods:name id="5CE4E83BD99C9D5837E3BFA59A28F51D" type="personal">
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<mods:namePart id="0E7D71FDD939CEB217688A867FFD5882">Boyer, Sarah L.</mods:namePart>
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<mods:role id="A405BC8074FA45CEC1D80EE4F2DABEFC">
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<mods:namePart id="5B52D475D4434C98CEC18C3E783145C2">Giribet, Gonzalo</mods:namePart>
</mods:name>
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<mods:title id="45E0667488032C71A88DA285FEC1D63C">Zootaxa</mods:title>
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<mods:date id="5784D2454E776B5E5FCA0B032A92A728">2003</mods:date>
<mods:detail id="733C16FD8C577436C6AB70728E297CFF" type="pubDate">
<mods:number id="54581F3A30942DC4403E25E36EADAAD0">2003-01-17</mods:number>
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<mods:number id="05E328AD2B2E0DCD1B48C379C21DB34F">133</mods:number>
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<treatment id="97070A43FF975A2A5708F9FAB9DCADEB" ID-DOI="http://doi.org/10.5281/zenodo.5093221" ID-GBIF-Taxon="183501690" ID-Zenodo-Dep="5093221" LSID="urn:lsid:plazi:treatment:97070A43FF975A2A5708F9FAB9DCADEB" httpUri="http://treatment.plazi.org/id/97070A43FF975A2A5708F9FAB9DCADEB" lastPageId="12" lastPageNumber="12" pageId="1" pageNumber="2">
<subSubSection id="57B4E8DEFF975A275708F9FABADBAA1D" box="[264,489,1593,1619]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="1F11BB55FF975A275708F9FABADBAA1D" blockId="1.[264,489,1593,1619]" box="[264,489,1593,1619]" pageId="1" pageNumber="2">
<heading id="44590C39FF975A275708F9FABADBAA1D" bold="true" box="[264,489,1593,1619]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<taxonomicName id="D8AEC0D6FF975A275708F9FABADBAA1D" ID-CoL="7PXC5" authority="Hirst 1925" authorityName="Hirst" authorityYear="1925" box="[264,489,1593,1619]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="2DDA6747FF975A275708F9FABADBAA1D" bold="true" box="[264,489,1593,1619]" pageId="1" pageNumber="2">
<emphasis id="2DDA6747FF975A275708F9FABA6CAA1D" bold="true" box="[264,350,1593,1619]" italics="true" pageId="1" pageNumber="2">Rakaia</emphasis>
Hirst 1925
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="57B4E8DEFF975A245708F940BACEAF70" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="type_taxon">
<paragraph id="1F11BB55FF975A275708F940B81AAAD0" blockId="1.[264,1323,1667,1814]" box="[264,808,1667,1694]" pageId="1" pageNumber="2">
<emphasis id="2DDA6747FF975A275708F940BA9EAAD3" bold="true" box="[264,428,1667,1693]" pageId="1" pageNumber="2">
<typeStatus id="C01505F7FF975A275708F940BA77AAD3" box="[264,325,1667,1693]" pageId="1" pageNumber="2">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="D8AEC0D6FF975A2757B4F947B816AAD0" ID-CoL="7YQK9" authority="Hirst 1925" authorityName="Hirst" authorityYear="1925" box="[436,804,1668,1694]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="antipodiana">
<emphasis id="2DDA6747FF975A2757B4F947B9AFAAD3" box="[436,669,1668,1693]" italics="true" pageId="1" pageNumber="2">Rakaia antipodiana</emphasis>
Hirst 1925
</taxonomicName>
.
</paragraph>
<paragraph id="1F11BB55FF975A275738F968B8E0AB58" blockId="1.[264,1323,1667,1814]" pageId="1" pageNumber="2">
<emphasis id="2DDA6747FF975A275738F968B9C2AA8B" bold="true" box="[312,752,1707,1733]" pageId="1" pageNumber="2">Species account and distribution:</emphasis>
23 valid species (including the new species described here) are known, with 3 occurring in
<collectingRegion id="DD6A75B7FF975A275531F917B889AAA0" box="[817,955,1748,1774]" country="Australia" name="Queensland" pageId="1" pageNumber="2">Queensland</collectingRegion>
(
<collectingCountry id="67B9FBC5FF975A2755CBF917BF08AAA0" box="[971,1082,1748,1774]" name="Australia" pageId="1" pageNumber="2">Australia</collectingCountry>
) and the rest in
<collectingCountry id="67B9FBC5FF975A2752F4F917BA58AB58" name="New Zealand" pageId="1" pageNumber="2">New Zealand</collectingCountry>
. Six of these species comprise two subspecies each.
</paragraph>
<paragraph id="1F11BB55FF945A245708FED8B924AD78" blockId="2.[264,534,283,350]" box="[264,534,283,310]" pageId="2" pageNumber="3">
<taxonomicName id="D8AEC0D6FF945A245708FED8BA80AD7B" ID-CoL="7YQKY" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[264,434,283,309]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF945A245708FED8BA80AD7B" bold="true" box="[264,434,283,309]" italics="true" pageId="2" pageNumber="3">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF945A2457BAFEDFB924AD78" box="[442,534,284,310]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="1F11BB55FF945A245708FE87BAA0AD10" blockId="2.[264,534,283,350]" box="[264,402,324,350]" pageId="2" pageNumber="3">
(
<figureCitation id="8795A7D0FF945A245711FE87BABBAD10" box="[273,393,324,350]" captionStart-0="FIGURES 1-2" captionStart-1="FIGURE 3" captionStart-2="FIGURES 4­5" captionStart-3="FIGURES 6­7" captionStart-4="FIGURES 8­11" captionStart-5="FIGURES 12­16" captionStart-6="FIGURES 17­22" captionStart-7="FIGURES 23­24" captionStart-8="FIGURES 25­28" captionStart-9="FIGURES 29­30" captionStartId-0="2.[264,382,1831,1855]" captionStartId-1="3.[264,368,1833,1857]" captionStartId-2="4.[264,383,1837,1861]" captionStartId-3="5.[264,383,1773,1797]" captionStartId-4="6.[264,382,995,1019]" captionStartId-5="7.[264,383,1630,1654]" captionStartId-6="8.[264,382,1730,1754]" captionStartId-7="9.[264,382,1303,1327]" captionStartId-8="10.[264,383,1797,1821]" captionStartId-9="11.[264,383,1673,1697]" captionTargetBox-0="[492,1094,969,1804]" captionTargetBox-1="[267,1319,1258,1815]" captionTargetBox-2="[265,1321,689,1810]" captionTargetBox-3="[315,1271,876,1740]" captionTargetBox-4="[265,1321,310,970]" captionTargetBox-5="[270,1317,609,1603]" captionTargetBox-6="[264,1323,449,1704]" captionTargetBox-7="[490,1097,609,1276]" captionTargetBox-8="[268,1323,738,1777]" captionTargetBox-9="[493,1093,777,1642]" captionTargetId-0="figure-213@2.[492,1094,969,1804]" captionTargetId-1="figure-331@3.[267,1319,1258,1815]" captionTargetId-2="figure-177@4.[265,1321,689,1810]" captionTargetId-3="figure-202@5.[311,1276,873,1746]" captionTargetId-4="figure-273@6.[265,1321,310,970]" captionTargetId-5="figure-183@7.[270,1317,609,1603]" captionTargetId-6="figure-115@8.[268,1319,917,1704]" captionTargetId-7="figure-276@9.[490,1097,609,1276]" captionTargetId-8="figure-205@10.[264,1323,729,1786]" captionTargetId-9="figure-215@11.[493,1093,777,1642]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURES 1-2. Rakaia macra sp. nov. 1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm." captionText-1="FIGURE 3. Rakaia macra sp. nov. Dorsal view of male paratype. Scale bar 1 mm." captionText-2="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." captionText-3="FIGURES 6­7. Rakaia macra sp. nov., lateral view. 6. Male holotype; 7. Female paratype. Scale bar 1 mm." captionText-4="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." captionText-5="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." captionText-6="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." captionText-7="FIGURES 23­24. Rakaia macra sp. nov. male paratype anal region. 23. Posterior view of anal region, showing scopulae on margin of the anal plate and on lobes of tergite 8 (scale bar 200 µm); 24. Dorsal angle, showing lobes and scopulae of tergite 8 (scale bar 300 µm)." captionText-8="FIGURES 25­28. Rakaia macra sp. nov. apex of penis. 25. Dorsal view; 26. Dorsal view showing movable fingers (mf); 27. Ventral view; 28. Light microscope image, dorsal view. Scale bar 100 µm for all." captionText-9="FIGURES 29­30. Rakaia macra sp. nov. ovipositor. 29. Entire ovipositor structure (scale bar 400 µm); 30. Ovipositor tip (scale bar 200 µm)." figureDoi-0="http://doi.org/10.5281/zenodo.5014173" figureDoi-1="http://doi.org/10.5281/zenodo.5014175" figureDoi-2="http://doi.org/10.5281/zenodo.5014177" figureDoi-3="http://doi.org/10.5281/zenodo.5014179" figureDoi-4="http://doi.org/10.5281/zenodo.5014183" figureDoi-5="http://doi.org/10.5281/zenodo.5014185" figureDoi-6="http://doi.org/10.5281/zenodo.5014187" figureDoi-7="http://doi.org/10.5281/zenodo.5014189" figureDoi-8="http://doi.org/10.5281/zenodo.5014193" figureDoi-9="http://doi.org/10.5281/zenodo.5014195" httpUri-0="https://zenodo.org/record/5014173/files/figure.png" httpUri-1="https://zenodo.org/record/5014175/files/figure.png" httpUri-2="https://zenodo.org/record/5014177/files/figure.png" httpUri-3="https://zenodo.org/record/5014179/files/figure.png" httpUri-4="https://zenodo.org/record/5014183/files/figure.png" httpUri-5="https://zenodo.org/record/5014185/files/figure.png" httpUri-6="https://zenodo.org/record/5014187/files/figure.png" httpUri-7="https://zenodo.org/record/5014189/files/figure.png" httpUri-8="https://zenodo.org/record/5014193/files/figure.png" httpUri-9="https://zenodo.org/record/5014195/files/figure.png" pageId="2" pageNumber="3">Figs. 1-30</figureCitation>
)
</paragraph>
<paragraph id="1F11BB55FF945A245708FE50BE16AE00" blockId="2.[264,1323,403,910]" pageId="2" pageNumber="3">
<emphasis id="2DDA6747FF945A245708FE50BA68ADE3" bold="true" box="[264,346,403,429]" pageId="2" pageNumber="3">Types.</emphasis>
<materialsCitation id="AFC6B108FF945A245765FE57B9C6AD98" ID-GBIF-Occurrence="3320620304" collectingDate="1977-03-18" collectionCode="OMNZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
<typeStatus id="C01505F7FF945A245765FE57BAE5ADE0" box="[357,471,404,430]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
: male from
<location id="1A71ED8EFF945A24546BFE57B9F9ADE0" LSID="urn:lsid:plazi:treatment:97070A43FF975A2A5708F9FAB9DCADEB:1A71ED8EFF945A24546BFE57B9F9ADE0" box="[619,715,404,430]" country="New Zealand" municipality="South Island" name="Waipori" pageId="2" pageNumber="3" stateProvince="Otago">Waipori</location>
(
<collectingRegion id="DD6A75B7FF945A2454DDFE57B81BADE0" box="[733,809,404,430]" country="New Zealand" name="Otago" pageId="2" pageNumber="3">Otago</collectingRegion>
,
<collectingMunicipality id="FF75212FFF945A245538FE57B8FDADE0" box="[824,975,404,430]" pageId="2" pageNumber="3">South Island</collectingMunicipality>
of
<collectingCountry id="67B9FBC5FF945A2455FBFE57BF92ADE0" box="[1019,1184,404,430]" name="New Zealand" pageId="2" pageNumber="3">New Zealand</collectingCountry>
),
<date id="6B109D95FF945A2452B7FE57BA77AD98" pageId="2" pageNumber="3" value="1977-03-18">
<collectingDate id="7B54647DFF945A2452B7FE57BA77AD98" pageId="2" pageNumber="3" value="1977-03-18">18 March 1977</collectingDate>
</date>
,
<collectorName id="B25BDE83FF945A245751FE7FBAF4AD98" box="[337,454,444,470]" pageId="2" pageNumber="3">R. Forster</collectorName>
leg., deposited in
<collectionCode id="79BF2390FF945A24549AFE7FB9C6AD98" box="[666,756,444,470]" country="New Zealand" httpUri="http://grbio.org/cool/1rhp-izex" name="Otago Museum" pageId="2" pageNumber="3">OMNZ</collectionCode>
</materialsCitation>
.
<materialsCitation id="AFC6B108FF945A2454FFFE7FB9C4ADB0" ID-GBIF-Occurrence="3320620303" collectingDate="1977-03-18" collectionCode="OMNZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Otago" typeStatus="paratype">
<typeStatus id="C01505F7FF945A2454FFFE7FB844AD98" box="[767,886,444,470]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="09A870DCFF945A24557FFE7FB8EBAD98" box="[895,985,444,470]" pageId="2" pageNumber="3" type="male">5 males</specimenCount>
and
<specimenCount id="09A870DCFF945A245211FE7FBFB6AD98" box="[1041,1156,444,470]" pageId="2" pageNumber="3" type="female">6 females</specimenCount>
, same collecting data as holotype, deposited in
<collectionCode id="79BF2390FF945A24549CFE27B9C4ADB0" box="[668,758,484,510]" country="New Zealand" httpUri="http://grbio.org/cool/1rhp-izex" name="Otago Museum" pageId="2" pageNumber="3">OMNZ</collectionCode>
</materialsCitation>
;
<materialsCitation id="AFC6B108FF945A245503FE27B941AE68" ID-GBIF-Occurrence="3320620301" collectingDate="1977-03-18" collectionCode="MONZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Otago" typeStatus="paratype">
<specimenCount id="09A870DCFF945A245503FE27B86FADB0" box="[771,861,484,510]" pageId="2" pageNumber="3" type="male">6 males</specimenCount>
and
<specimenCount id="09A870DCFF945A245598FE27BF3FADB0" box="[920,1037,484,510]" pageId="2" pageNumber="3" type="female">6 females</specimenCount>
, same collecting data as holotype, deposited in
<collectionCode id="79BF2390FF945A245419FDCFB941AE68" box="[537,627,524,550]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="2" pageNumber="3" type="Museum">MONZ</collectionCode>
</materialsCitation>
;
<materialsCitation id="AFC6B108FF945A245481FDCFB94DAE00" ID-GBIF-Occurrence="3320620302" accessionNumber="MCZ 48184" collectingDate="1977-03-18" collectionCode="MCZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCode="MCZ 48184-48195" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Otago" typeStatus="paratype">
<specimenCount id="09A870DCFF945A245481FDCFB9EFAE68" box="[641,733,524,550]" pageId="2" pageNumber="3" type="male">6 males</specimenCount>
and
<specimenCount id="09A870DCFF945A24551AFDCFB8A2AE68" box="[794,912,524,550]" pageId="2" pageNumber="3" type="female">6 females</specimenCount>
, same collecting data as holotype, deposited in
<specimenCode id="4F08132EFF945A24579DFDF7B94DAE00" box="[413,639,564,590]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="2" pageNumber="3" type="Museum">MCZ 48184-48195</specimenCode>
</materialsCitation>
(including
<specimenCount id="09A870DCFF945A245505FDF7B866AE00" box="[773,852,564,590]" pageId="2" pageNumber="3" type="male">2 male</specimenCount>
and
<specimenCount id="09A870DCFF945A24558BFDF7B8CEAE00" box="[907,1020,564,590]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
mounted on SEM stubs).
</paragraph>
<paragraph id="1F11BB55FF945A245738FD98BACEAF70" blockId="2.[264,1323,403,910]" pageId="2" pageNumber="3">
<emphasis id="2DDA6747FF945A245738FD98B9B6AE3B" bold="true" box="[312,644,603,629]" pageId="2" pageNumber="3">Other material examined.</emphasis>
<materialsCitation id="AFC6B108FF945A245490FD9FBF9EAED0" ID-GBIF-Occurrence="3320620305" collectingDate="1977-03-18" collectionCode="OMNZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="75" specimenCount-female="36" specimenCount-male="39" stateProvince="Otago">
<specimenCount id="09A870DCFF945A245490FD9FB9CFAE38" box="[656,765,604,630]" pageId="2" pageNumber="3" type="male">39 males</specimenCount>
and
<specimenCount id="09A870DCFF945A245540FD9FB8F6AE38" box="[832,964,604,630]" pageId="2" pageNumber="3" type="female">36 females</specimenCount>
from
<location id="1A71ED8EFF945A245215FD9FBF47AE38" LSID="urn:lsid:plazi:treatment:97070A43FF975A2A5708F9FAB9DCADEB:1A71ED8EFF945A245215FD9FBF47AE38" box="[1045,1141,604,630]" country="New Zealand" municipality="South Island" name="Waipori" pageId="2" pageNumber="3" stateProvince="Otago">Waipori</location>
(
<collectingRegion id="DD6A75B7FF945A24528AFD9FBFE4AE38" box="[1162,1238,604,630]" country="New Zealand" name="Otago" pageId="2" pageNumber="3">Otago</collectingRegion>
,
<collectingMunicipality id="FF75212FFF945A2452E6FD9FBA61AED0" pageId="2" pageNumber="3">South Island</collectingMunicipality>
, of
<collectingCountry id="67B9FBC5FF945A245781FD47B916AED0" box="[385,548,644,670]" name="New Zealand" pageId="2" pageNumber="3">New Zealand</collectingCountry>
),
<date id="6B109D95FF945A245439FD47B9C3AED0" box="[569,753,644,670]" pageId="2" pageNumber="3" value="1977-03-18">
<collectingDate id="7B54647DFF945A245439FD47B9C3AED0" box="[569,753,644,670]" pageId="2" pageNumber="3" value="1977-03-18">18 March 1977</collectingDate>
</date>
,
<collectorName id="B25BDE83FF945A2454FFFD47B847AED0" box="[767,885,644,670]" pageId="2" pageNumber="3">R. Forster</collectorName>
leg., deposited in
<collectionCode id="79BF2390FF945A245252FD47BF9EAED0" box="[1106,1196,644,670]" country="New Zealand" httpUri="http://grbio.org/cool/1rhp-izex" name="Otago Museum" pageId="2" pageNumber="3">OMNZ</collectionCode>
</materialsCitation>
;
<materialsCitation id="AFC6B108FF945A2452B9FD47B834AEA0" ID-GBIF-Occurrence="3320620307" collectingDate="1977-03-21" collectionCode="OMNZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="40" specimenCount-female="18" specimenCount-juvenile="6" specimenCount-male="16" stateProvince="Otago">
<specimenCount id="09A870DCFF945A2452B9FD47BE14AED0" box="[1209,1318,644,670]" pageId="2" pageNumber="3" type="male">16 males</specimenCount>
,
<specimenCount id="09A870DCFF945A245708FD6FBABAAE88" box="[264,392,684,710]" pageId="2" pageNumber="3" type="female">18 females</specimenCount>
and
<specimenCount id="09A870DCFF945A2457C0FD6FB90DAE88" box="[448,575,684,710]" pageId="2" pageNumber="3" type="juvenile">6 juveniles</specimenCount>
from
<location id="1A71ED8EFF945A245486FD6FB9D4AE88" LSID="urn:lsid:plazi:treatment:97070A43FF975A2A5708F9FAB9DCADEB:1A71ED8EFF945A245486FD6FB9D4AE88" box="[646,742,684,710]" country="New Zealand" municipality="South Island" name="Waipori" pageId="2" pageNumber="3" stateProvince="Otago">Waipori</location>
(
<collectingRegion id="DD6A75B7FF945A2454F5FD6FB873AE88" box="[757,833,684,710]" country="New Zealand" name="Otago" pageId="2" pageNumber="3">Otago</collectingRegion>
,
<collectingMunicipality id="FF75212FFF945A24554CFD6FB8D2AE88" box="[844,992,684,710]" pageId="2" pageNumber="3">South Island</collectingMunicipality>
of
<collectingCountry id="67B9FBC5FF945A245205FD6FBF95AE88" box="[1029,1191,684,710]" name="New Zealand" pageId="2" pageNumber="3">New Zealand</collectingCountry>
),
<date id="6B109D95FF945A2452BAFD6FBA77AEA0" pageId="2" pageNumber="3" value="1977-03-21">
<collectingDate id="7B54647DFF945A2452BAFD6FBA77AEA0" pageId="2" pageNumber="3" value="1977-03-21">21 March 1977</collectingDate>
</date>
,
<collectorName id="B25BDE83FF945A245754FD17BAF9AEA0" box="[340,459,724,750]" pageId="2" pageNumber="3">R. Forster</collectorName>
leg., deposited in
<collectionCode id="79BF2390FF945A2454ACFD17B834AEA0" box="[684,774,724,750]" country="New Zealand" httpUri="http://grbio.org/cool/1rhp-izex" name="Otago Museum" pageId="2" pageNumber="3">OMNZ</collectionCode>
</materialsCitation>
;
<specimenCount id="09A870DCFF945A245514FD17B8B0AEA0" box="[788,898,724,750]" pageId="2" pageNumber="3" type="male">19 males</specimenCount>
,
<specimenCount id="09A870DCFF945A245590FD17BF27AEA0" box="[912,1045,724,750]" pageId="2" pageNumber="3" type="female">19 females</specimenCount>
, and
<materialsCitation id="AFC6B108FF945A245257FD17BACAAF70" ID-GBIF-Occurrence="3320620306" collectingDate="1977-09-20" collectionCode="OMNZ" collectorName="R. Forster" country="New Zealand" location="Waipori" municipality="South Island" pageId="2" pageNumber="3" specimenCount="10" specimenCount-juvenile="10" stateProvince="Otago">
<specimenCount id="09A870DCFF945A245257FD17BFDAAEA0" box="[1111,1256,724,750]" pageId="2" pageNumber="3" type="juvenile">10 juveniles</specimenCount>
from
<location id="1A71ED8EFF945A245708FD3FBA5AAF58" LSID="urn:lsid:plazi:treatment:97070A43FF975A2A5708F9FAB9DCADEB:1A71ED8EFF945A245708FD3FBA5AAF58" box="[264,360,764,790]" country="New Zealand" municipality="South Island" name="Waipori" pageId="2" pageNumber="3" stateProvince="Otago">Waipori</location>
(
<collectingRegion id="DD6A75B7FF945A24577EFD3FBAF8AF58" box="[382,458,764,790]" country="New Zealand" name="Otago" pageId="2" pageNumber="3">Otago</collectingRegion>
,
<collectingMunicipality id="FF75212FFF945A2457DCFD3FB944AF58" box="[476,630,764,790]" pageId="2" pageNumber="3">South Island</collectingMunicipality>
of
<collectingCountry id="67B9FBC5FF945A2454A9FD3FB862AF58" box="[681,848,764,790]" name="New Zealand" pageId="2" pageNumber="3">New Zealand</collectingCountry>
),
<date id="6B109D95FF945A24556AFD3FBF6FAF58" box="[874,1117,764,790]" pageId="2" pageNumber="3" value="1977-09-20">
<collectingDate id="7B54647DFF945A24556AFD3FBF6FAF58" box="[874,1117,764,790]" pageId="2" pageNumber="3" value="1977-09-20">20 September 1977</collectingDate>
</date>
,
<collectorName id="B25BDE83FF945A24526FFD3FBFD9AF58" box="[1135,1259,764,790]" pageId="2" pageNumber="3">R. Forster</collectorName>
leg., deposited in
<collectionCode id="79BF2390FF945A24579EFCE7BACAAF70" box="[414,504,804,830]" country="New Zealand" httpUri="http://grbio.org/cool/1rhp-izex" name="Otago Museum" pageId="2" pageNumber="3">OMNZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57B4E8DEFF945A245738FC8FBF17AFC0" pageId="2" pageNumber="3" type="etymology">
<paragraph id="1F11BB55FF945A245738FC8FBF17AFC0" blockId="2.[264,1323,403,910]" pageId="2" pageNumber="3">
<emphasis id="2DDA6747FF945A245738FC8FBAF4AF28" bold="true" box="[312,454,844,870]" pageId="2" pageNumber="3">Etymology.</emphasis>
Latin adjective:
<taxonomicName id="D8AEC0D6FF945A245491FC8FB9EEAF2B" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[657,732,844,869]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF945A245491FC8FB9EEAF2B" box="[657,732,844,869]" italics="true" pageId="2" pageNumber="3">macra</emphasis>
</taxonomicName>
= lean, skinny. The specific epithet refers to the general appearance of the animal’s body, particularly in lateral view.
</paragraph>
</subSubSection>
<caption id="4BD1EBDDFF945A245708F8E4B95DAB2F" ID-DOI="http://doi.org/10.5281/zenodo.5014173" ID-Zenodo-Dep="5014173" httpUri="https://zenodo.org/record/5014173/files/figure.png" pageId="2" pageNumber="3" startId="2.[264,382,1831,1855]" targetBox="[492,1094,969,1804]" targetPageId="2">
<paragraph id="1F11BB55FF945A245708F8E4B95DAB2F" blockId="2.[264,1323,1831,1889]" pageId="2" pageNumber="3">
<emphasis id="2DDA6747FF945A245708F8E4BA83AB71" bold="true" box="[264,433,1831,1855]" pageId="2" pageNumber="3">FIGURES 1-2.</emphasis>
<taxonomicName id="D8AEC0D6FF945A2457BFF8E4B964AB70" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[447,598,1831,1854]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF945A2457BFF8E4B964AB70" box="[447,598,1831,1854]" italics="true" pageId="2" pageNumber="3">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF945A24545EF8E4B981AB70" box="[606,691,1831,1854]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm.
</paragraph>
</caption>
<subSubSection id="57B4E8DEFF955A255738FED8B8D1ADE0" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="1F11BB55FF955A255738FED8B8D1ADE0" blockId="3.[264,1324,283,1150]" pageId="3" pageNumber="4">
<emphasis id="2DDA6747FF955A255738FED8BA88AD7B" bold="true" box="[312,442,283,309]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
Pettalid with dorsal scutum flat (
<figureCitation id="8795A7D0FF955A25554CFEDFB889AD78" box="[844,955,284,310]" captionStart="FIGURES 6­7" captionStartId="5.[264,383,1773,1797]" captionTargetBox="[315,1271,876,1740]" captionTargetId="figure-202@5.[311,1276,873,1746]" captionTargetPageId="5" captionText="FIGURES 6­7. Rakaia macra sp. nov., lateral view. 6. Male holotype; 7. Female paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014179" httpUri="https://zenodo.org/record/5014179/files/figure.png" pageId="3" pageNumber="4">Figs 6, 7</figureCitation>
). Trochanter of palp with conspicuous ventral process (
<figureCitation id="8795A7D0FF955A25543DFE87B9A7AD10" box="[573,661,324,350]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
); chelicerae with distinct outer lateral ridge on the surface of the second article (
<figureCitation id="8795A7D0FF955A255442FEAFB986ADC8" box="[578,692,364,390]" captionStart="FIGURES 4­5" captionStartId="4.[264,383,1837,1861]" captionTargetBox="[265,1321,689,1810]" captionTargetId="figure-177@4.[265,1321,689,1810]" captionTargetPageId="4" captionText="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="3" pageNumber="4">Figs. 4, 5</figureCitation>
,
<figureCitation id="8795A7D0FF955A2554C2FEAFB9E0ADC8" box="[706,722,364,390]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="3" pageNumber="4">8</figureCitation>
), lacking a ventral process on the first article (
<figureCitation id="8795A7D0FF955A2552FCFEAFBA2AADE0" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
); dentition of the mobile digit of chelicerae uniform (
<figureCitation id="8795A7D0FF955A25558DFE57B8E6ADE0" box="[909,980,404,430]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57B4E8DEFF955A2F5738FE78B98AAD10" lastPageId="9" lastPageNumber="10" pageId="3" pageNumber="4" type="description">
<paragraph id="1F11BB55FF955A255738FE78BE22AF70" blockId="3.[264,1324,283,1150]" pageId="3" pageNumber="4">
<emphasis id="2DDA6747FF955A255738FE78BAE0AD9B" bold="true" box="[312,466,443,469]" pageId="3" pageNumber="4">Description:</emphasis>
Total length of male
<typeStatus id="C01505F7FF955A2554D3FE7FB80BAD98" box="[723,825,444,470]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
(female
<typeStatus id="C01505F7FF955A2555A4FE7FBF3BAD98" box="[932,1033,444,470]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
measurements in parentheses) 2.25 (2.60), width across ozophores 0.855 (0.769), greatest width 1.19 (1.29); length/width ratio 1.89 (2.02). Body brown­yellow (in alcohol) (
<figureCitation id="8795A7D0FF955A255210FDCFBFB0AE68" box="[1040,1154,524,550]" captionStart="FIGURES 1-2" captionStartId="2.[264,382,1831,1855]" captionTargetBox="[492,1094,969,1804]" captionTargetId="figure-213@2.[492,1094,969,1804]" captionTargetPageId="2" captionText="FIGURES 1-2. Rakaia macra sp. nov. 1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5014173" httpUri="https://zenodo.org/record/5014173/files/figure.png" pageId="3" pageNumber="4">Figs. 1, 2</figureCitation>
) with most of the dorsal surface and legs showing a tuberculate­granulate microstructure. Anterior margin of dorsal scutum with two projections delimiting the insertion of the chelicerae (
<figureCitation id="8795A7D0FF955A2552F0FD9FBA24AED0" captionStart="FIGURES 1-2" captionStartId="2.[264,382,1831,1855]" captionTargetBox="[492,1094,969,1804]" captionTargetId="figure-213@2.[492,1094,969,1804]" captionTargetPageId="2" captionText="FIGURES 1-2. Rakaia macra sp. nov. 1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5014173" httpUri="https://zenodo.org/record/5014173/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="8795A7D0FF955A255727FD47BA05AED0" box="[295,311,644,670]" captionStart="FIGURE 3" captionStartId="3.[264,368,1833,1857]" captionTargetBox="[267,1319,1258,1815]" captionTargetId="figure-331@3.[267,1319,1258,1815]" captionTargetPageId="3" captionText="FIGURE 3. Rakaia macra sp. nov. Dorsal view of male paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014175" httpUri="https://zenodo.org/record/5014175/files/figure.png" pageId="3" pageNumber="4">3</figureCitation>
); prosomal region trapezoidal. Eyes absent. Ozophores conical and positioned dorsally, somewhere between types 2 and 3 of
<bibRefCitation id="7B3FC6A4FF955A255520FD6FB8D6AE88" author="Juberthie, C." box="[800,996,684,710]" pageId="3" pageNumber="4" pagination="1371 - 1390" refId="ref4486" refString="Juberthie, C. (1970) Les genres d'opilions sironinae (Cyphophthalmes). Bulletin du Museum National d'Histoire Naturelle, 41 (6), 1371 - 1390." type="journal article" year="1970">Juberthie (1970)</bibRefCitation>
(
<figureCitation id="8795A7D0FF955A2555F7FD6FBF7FAE88" box="[1015,1101,684,710]" captionStart="FIGURE 3" captionStartId="3.[264,368,1833,1857]" captionTargetBox="[267,1319,1258,1815]" captionTargetId="figure-331@3.[267,1319,1258,1815]" captionTargetPageId="3" captionText="FIGURE 3. Rakaia macra sp. nov. Dorsal view of male paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014175" httpUri="https://zenodo.org/record/5014175/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
,
<figureCitation id="8795A7D0FF955A25525EFD6FBF4CAE88" box="[1118,1150,684,710]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="3" pageNumber="4">17</figureCitation>
). Transverse prosomal sulcus distinct but not conspicuous (
<figureCitation id="8795A7D0FF955A25552BFD17B841AEA0" box="[811,883,724,750]" captionStart="FIGURE 3" captionStartId="3.[264,368,1833,1857]" captionTargetBox="[267,1319,1258,1815]" captionTargetId="figure-331@3.[267,1319,1258,1815]" captionTargetPageId="3" captionText="FIGURE 3. Rakaia macra sp. nov. Dorsal view of male paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014175" httpUri="https://zenodo.org/record/5014175/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). Transverse opisthosomal sulci distinct by lacking granulation, longitudinal opisthosomal sulcus absent (
<figureCitation id="8795A7D0FF955A255249FD3FBFAEAF58" box="[1097,1180,764,790]" captionStart="FIGURES 1-2" captionStartId="2.[264,382,1831,1855]" captionTargetBox="[492,1094,969,1804]" captionTargetId="figure-213@2.[492,1094,969,1804]" captionTargetPageId="2" captionText="FIGURES 1-2. Rakaia macra sp. nov. 1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5014173" httpUri="https://zenodo.org/record/5014173/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="8795A7D0FF955A2552ACFD3FBF8EAF58" box="[1196,1212,764,790]" captionStart="FIGURE 3" captionStartId="3.[264,368,1833,1857]" captionTargetBox="[267,1319,1258,1815]" captionTargetId="figure-331@3.[267,1319,1258,1815]" captionTargetPageId="3" captionText="FIGURE 3. Rakaia macra sp. nov. Dorsal view of male paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014175" httpUri="https://zenodo.org/record/5014175/files/figure.png" pageId="3" pageNumber="4">3</figureCitation>
). Dorsal scutum flat (
<figureCitation id="8795A7D0FF955A25579BFCE7B93EAF70" box="[411,524,804,830]" captionStart="FIGURES 6­7" captionStartId="5.[264,383,1773,1797]" captionTargetBox="[315,1271,876,1740]" captionTargetId="figure-202@5.[311,1276,873,1746]" captionTargetPageId="5" captionText="FIGURES 6­7. Rakaia macra sp. nov., lateral view. 6. Male holotype; 7. Female paratype. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014179" httpUri="https://zenodo.org/record/5014179/files/figure.png" pageId="3" pageNumber="4">Figs. 6, 7</figureCitation>
); opisthosomal region reaching its maximum width at segment II.
</paragraph>
<paragraph id="1F11BB55FF955A255738FC8FB886A830" blockId="3.[264,1324,283,1150]" pageId="3" pageNumber="4">
Coxae of legs I and II mobile; coxae of remaining legs fused. Ventral prosomal complex of male with coxae II, III, and IV meeting in the midline (
<figureCitation id="8795A7D0FF955A2555FCFCB7BF74AFC0" box="[1020,1094,884,910]" captionStart="FIGURES 4­5" captionStartId="4.[264,383,1837,1861]" captionTargetBox="[265,1321,689,1810]" captionTargetId="figure-177@4.[265,1321,689,1810]" captionTargetPageId="4" captionText="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). Male gonostome small, subtriangular, slightly wider than long, bordered on posterior margin by the first opisthosomal sternite (
<figureCitation id="8795A7D0FF955A255413FC07B959AF90" box="[531,619,964,990]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="3" pageNumber="4">Fig. 19</figureCitation>
); male gonostome shorter than length of seam of contact of left and right coxae IV parallel to midline. Ventral prosomal complex of female with only coxae II and III meeting in the midline (
<figureCitation id="8795A7D0FF955A2554F6FBD7B863A860" box="[758,849,1044,1070]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="3" pageNumber="4">Fig. 20</figureCitation>
). Female gonostome roughly round in shape, with the edges of coxae of leg IV and first opisthosomal sternite forming a partial “tube” at the posterior margin of the opening (
<figureCitation id="8795A7D0FF955A255526FBA7B84AA830" box="[806,888,1124,1150]" captionStart="FIGURES 4­5" captionStartId="4.[264,383,1837,1861]" captionTargetBox="[265,1321,689,1810]" captionTargetId="figure-177@4.[265,1321,689,1810]" captionTargetPageId="4" captionText="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="3" pageNumber="4">Figs. 5</figureCitation>
,
<figureCitation id="8795A7D0FF955A255586FBA7B895A830" box="[902,935,1124,1150]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="3" pageNumber="4">20</figureCitation>
).
</paragraph>
<caption id="4BD1EBDDFF955A255708F8EABFA0AB0E" ID-DOI="http://doi.org/10.5281/zenodo.5014175" ID-Zenodo-Dep="5014175" box="[264,1170,1833,1857]" httpUri="https://zenodo.org/record/5014175/files/figure.png" pageId="3" pageNumber="4" startId="3.[264,368,1833,1857]" targetBox="[267,1319,1258,1815]" targetPageId="3">
<paragraph id="1F11BB55FF955A255708F8EABFA0AB0E" blockId="3.[264,1170,1833,1857]" box="[264,1170,1833,1857]" pageId="3" pageNumber="4">
<emphasis id="2DDA6747FF955A255708F8EABAB8AB0F" bold="true" box="[264,394,1833,1857]" pageId="3" pageNumber="4">FIGURE 3.</emphasis>
<taxonomicName id="D8AEC0D6FF955A255798F8E9B91CAB0F" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[408,558,1834,1857]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF955A255798F8E9B91CAB0F" box="[408,558,1834,1857]" italics="true" pageId="3" pageNumber="4">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF955A255435F8EAB9BAAB0E" box="[565,648,1833,1856]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
Dorsal view of male paratype. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="1F11BB55FF925A225738FEDFBF47AE38" blockId="4.[264,1324,284,630]" pageId="4" pageNumber="5">
Spiracles C­shaped (sensu
<bibRefCitation id="7B3FC6A4FF925A225478FEDFB8ABAD78" author="Giribet, G. &amp; Boyer, S. L." box="[632,921,284,310]" pageId="4" pageNumber="5" pagination="110 - 128" refId="ref4406" refString="Giribet, G. &amp; Boyer, S. L. (2002) A cladistic analysis of the cyphophthalmid genera (Opiliones, Cyphophthalmi). Journal of Arachnology, 30, 110 - 128." type="journal article" year="2002">Giribet and Boyer 2002</bibRefCitation>
), with one of the edges recurving internally as found in the “open circle”
<typeStatus id="C01505F7FF925A2254D4FE87B834AD10" box="[724,774,324,350]" pageId="4" pageNumber="5">type</typeStatus>
(
<figureCitation id="8795A7D0FF925A225517FE87B85FAD10" box="[791,877,324,350]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
). Sternal opisthosomal region without modifications or glandular pores (
<figureCitation id="8795A7D0FF925A2254A3FEAFB9C8ADC8" box="[675,762,364,390]" captionStart="FIGURES 1-2" captionStartId="2.[264,382,1831,1855]" captionTargetBox="[492,1094,969,1804]" captionTargetId="figure-213@2.[492,1094,969,1804]" captionTargetPageId="2" captionText="FIGURES 1-2. Rakaia macra sp. nov. 1. Dorsal view of male holotype; 2. Ventral view of male holotype. Total length: 2.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5014173" httpUri="https://zenodo.org/record/5014173/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
,
<figureCitation id="8795A7D0FF925A225509FEAFB809ADC8" box="[777,827,364,390]" captionStart="FIGURES 4­5" captionStartId="4.[264,383,1837,1861]" captionTargetBox="[265,1321,689,1810]" captionTargetId="figure-177@4.[265,1321,689,1810]" captionTargetPageId="4" captionText="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="4" pageNumber="5">4, 5</figureCitation>
). Anal region with sternites 8 and 9 and tergite IX free, not forming a corona analis (
<figureCitation id="8795A7D0FF925A225509FE57B858ADE0" box="[777,874,404,430]" captionStart="FIGURES 17­22" captionStartId="8.[264,382,1730,1754]" captionTargetBox="[264,1323,449,1704]" captionTargetId="figure-115@8.[268,1319,917,1704]" captionTargetPageId="8" captionText="FIGURES 17­22. Rakaia macra sp. nov. 17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="4" pageNumber="5">Figs. 21</figureCitation>
,
<figureCitation id="8795A7D0FF925A225576FE57B8A5ADE0" box="[886,919,404,430]" captionStart="FIGURES 23­24" captionStartId="9.[264,382,1303,1327]" captionTargetBox="[490,1097,609,1276]" captionTargetId="figure-276@9.[490,1097,609,1276]" captionTargetPageId="9" captionText="FIGURES 23­24. Rakaia macra sp. nov. male paratype anal region. 23. Posterior view of anal region, showing scopulae on margin of the anal plate and on lobes of tergite 8 (scale bar 200 µm); 24. Dorsal angle, showing lobes and scopulae of tergite 8 (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014189" httpUri="https://zenodo.org/record/5014189/files/figure.png" pageId="4" pageNumber="5">23</figureCitation>
). Area of contact of tergite IX and sternite 9 of “pettalid”
<typeStatus id="C01505F7FF925A22541AFE7FB97EAD98" box="[538,588,444,470]" pageId="4" pageNumber="5">type</typeStatus>
(
<bibRefCitation id="7B3FC6A4FF925A22545DFE7FB848AD98" author="Giribet, G. &amp; Boyer, S. L." box="[605,890,444,470]" pageId="4" pageNumber="5" pagination="110 - 128" refId="ref4406" refString="Giribet, G. &amp; Boyer, S. L. (2002) A cladistic analysis of the cyphophthalmid genera (Opiliones, Cyphophthalmi). Journal of Arachnology, 30, 110 - 128." type="journal article" year="2002">Giribet and Boyer 2002</bibRefCitation>
) in which tergite IX laterally covers sternite 9 and clearly meets sternite 8. Anal plate of male without longitudinal carina; posterior margin of anal plate slightly concave, bearing a scopula (
<figureCitation id="8795A7D0FF925A2255EFFDCFBF75AE68" box="[1007,1095,524,550]" captionStart="FIGURES 23­24" captionStartId="9.[264,382,1303,1327]" captionTargetBox="[490,1097,609,1276]" captionTargetId="figure-276@9.[490,1097,609,1276]" captionTargetPageId="9" captionText="FIGURES 23­24. Rakaia macra sp. nov. male paratype anal region. 23. Posterior view of anal region, showing scopulae on margin of the anal plate and on lobes of tergite 8 (scale bar 200 µm); 24. Dorsal angle, showing lobes and scopulae of tergite 8 (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014189" httpUri="https://zenodo.org/record/5014189/files/figure.png" pageId="4" pageNumber="5">Fig. 23</figureCitation>
). Scopulae present on each lobe of tergite VIII (
<figureCitation id="8795A7D0FF925A22546AFDF7B98CAE00" box="[618,702,564,590]" captionStart="FIGURES 23­24" captionStartId="9.[264,382,1303,1327]" captionTargetBox="[490,1097,609,1276]" captionTargetId="figure-276@9.[490,1097,609,1276]" captionTargetPageId="9" captionText="FIGURES 23­24. Rakaia macra sp. nov. male paratype anal region. 23. Posterior view of anal region, showing scopulae on margin of the anal plate and on lobes of tergite 8 (scale bar 200 µm); 24. Dorsal angle, showing lobes and scopulae of tergite 8 (scale bar 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014189" httpUri="https://zenodo.org/record/5014189/files/figure.png" pageId="4" pageNumber="5">Fig 24</figureCitation>
). Cuticle with granular surface in all ventral areas including coxae and anal plate. Anal glandular pore not observed in males.
</paragraph>
<caption id="4BD1EBDDFF925A225708F8EEB9C4AB29" ID-DOI="http://doi.org/10.5281/zenodo.5014177" ID-Zenodo-Dep="5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="4" pageNumber="5" startId="4.[264,383,1837,1861]" targetBox="[265,1321,689,1810]" targetPageId="4">
<paragraph id="1F11BB55FF925A225708F8EEB9C4AB29" blockId="4.[264,1323,1837,1895]" pageId="4" pageNumber="5">
<emphasis id="2DDA6747FF925A225708F8EEBA9DAB0B" bold="true" box="[264,431,1837,1861]" pageId="4" pageNumber="5">FIGURES 4­5.</emphasis>
<taxonomicName id="D8AEC0D6FF925A2257BEF8EDB966AB0B" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[446,596,1838,1861]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF925A2257BEF8EDB966AB0B" box="[446,596,1838,1861]" italics="true" pageId="4" pageNumber="5">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF925A22545CF8EEB99DAB0A" box="[604,687,1837,1860]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm).
</paragraph>
</caption>
<paragraph id="1F11BB55FF935A235738FEDFBFD5AE38" blockId="5.[264,1324,284,782]" pageId="5" pageNumber="6">
Chelicerae short, extremely robust, with very few long setae. Proximal article with hook­like dorsal crest (“dorsal ridge” of
<bibRefCitation id="7B3FC6A4FF935A2354FDFE87BF75AD10" author="Hansen, H. J. &amp; Sorensen, W." box="[765,1095,324,350]" pageId="5" pageNumber="6" refId="ref4443" refString="Hansen, H. J. &amp; Sorensen, W. (1904) &quot; On Two Orders of Arachnida: Opiliones, Especially the Suborder Cyphophthalmi, and Ricinulei, Namely the Family Cryptostemmatoidae. &quot; Cambridge University Press, Cambridge." type="book" year="1904">Hansen and Sørensen 1904</bibRefCitation>
and
<bibRefCitation id="7B3FC6A4FF935A23528AFE87BE14AD10" author="Forster, R. R." box="[1162,1318,324,350]" pageId="5" pageNumber="6" pagination="79 - 119" refId="ref4280" refString="Forster, R. R. (1948) The sub-order Cyphophthalmi Simon in New Zealand. Dominion Museum Records in Entomology, 1, 79 - 119." type="journal article" year="1948">Forster 1948</bibRefCitation>
; “dorsal transverse crest” of
<bibRefCitation id="7B3FC6A4FF935A235461FEAFB82AADC8" author="Juberthie, C." box="[609,792,364,390]" pageId="5" pageNumber="6" pagination="1371 - 1390" refId="ref4486" refString="Juberthie, C. (1970) Les genres d'opilions sironinae (Cyphophthalmes). Bulletin du Museum National d'Histoire Naturelle, 41 (6), 1371 - 1390." type="journal article" year="1970">Juberthie 1970</bibRefCitation>
), without ventral process, ornamented over almost its entire surface from the dorsal ridge to the second article (
<figureCitation id="8795A7D0FF935A23522DFE57BF44ADE0" box="[1069,1142,404,430]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
). Chelicerae of the protruding type described by Giribet (in press). Second article extremely wide, with a distinct longitudinal ridge on the outer lateral surface running towards the base of the mobile digit (
<figureCitation id="8795A7D0FF935A2357A7FDCFBACBAE68" box="[423,505,524,550]" captionStart="FIGURES 4­5" captionStartId="4.[264,383,1837,1861]" captionTargetBox="[265,1321,689,1810]" captionTargetId="figure-177@4.[265,1321,689,1810]" captionTargetPageId="4" captionText="FIGURES 4­5. Rakaia macra sp. nov. 4. Ventral view of male paratype (scale bar 1mm); 5. Ventral view of female paratype (scale bar 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5014177" httpUri="https://zenodo.org/record/5014177/files/figure.png" pageId="5" pageNumber="6">Figs. 5</figureCitation>
,
<figureCitation id="8795A7D0FF935A235406FDCFB924AE68" box="[518,534,524,550]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="5" pageNumber="6">8</figureCitation>
). Mobile digit with 12 uniform denticles of the same type (
<figureCitation id="8795A7D0FF935A2352D4FDCFBE2FAE68" box="[1236,1309,524,550]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
). Proximal cheliceral article of male
<typeStatus id="C01505F7FF935A2354ACFDF7B823AE00" box="[684,785,564,590]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
0.697 long, 0.303 wide; second article 0.947 long, 0.273 wide; mobile digit 0.121 long, 0.083 wide, 13% of second article length.
</paragraph>
<paragraph id="1F11BB55FF935A235738FD47BAFEAF40" blockId="5.[264,1324,284,782]" pageId="5" pageNumber="6">
Palp (
<figureCitation id="8795A7D0FF935A23577BFD47BAFDAED0" box="[379,463,644,670]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
) with a sharp, spiny ventral process on the trochanter (
<figureCitation id="8795A7D0FF935A23524CFD47BF93AED0" box="[1100,1185,644,670]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
). Measurements of palpal articles in male
<typeStatus id="C01505F7FF935A235480FD6AB9DAAE8D" box="[640,744,681,707]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
, length/width (L/W ratio), from trochanter to tarsus:
<geoCoordinate id="7A9ADD92FF935A23573EFD0CBA4DAEA7" box="[318,383,719,745]" degrees="0.275" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.275">0.275</geoCoordinate>
/
<geoCoordinate id="7A9ADD92FF935A235788FD0CBAF8AEA7" box="[392,458,719,745]" degrees="0.129" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="0.129">0.129</geoCoordinate>
(2.13);
<geoCoordinate id="7A9ADD92FF935A23542AFD0CB95EAEA7" box="[554,620,719,745]" degrees="0.422" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.422">0.422</geoCoordinate>
/
<geoCoordinate id="7A9ADD92FF935A235474FD0CB984AEA7" box="[628,694,719,745]" degrees="0.100" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="0.1">0.100</geoCoordinate>
(4.22);
<geoCoordinate id="7A9ADD92FF935A235515FD0CB867AEA7" box="[789,853,719,745]" degrees="0.311" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.311">0.311</geoCoordinate>
/
<geoCoordinate id="7A9ADD92FF935A23555EFD0CB8ACAEA7" box="[862,926,719,745]" degrees="0.111" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="0.111">0.111</geoCoordinate>
(2.80);
<geoCoordinate id="7A9ADD92FF935A2355FEFD0CBF0DAEA7" box="[1022,1087,719,745]" degrees="0.333" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="0.333">0.333</geoCoordinate>
/
<geoCoordinate id="7A9ADD92FF935A235248FD0CBFB8AEA7" box="[1096,1162,719,745]" degrees="0.222" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="0.222">0.222</geoCoordinate>
(1.5). Palpal claw 0.041 long.
</paragraph>
<caption id="4BD1EBDDFF935A235708F92EBA44AB69" ID-DOI="http://doi.org/10.5281/zenodo.5014179" ID-Zenodo-Dep="5014179" httpUri="https://zenodo.org/record/5014179/files/figure.png" pageId="5" pageNumber="6" startId="5.[264,383,1773,1797]" targetBox="[315,1271,876,1740]" targetPageId="5">
<paragraph id="1F11BB55FF935A235708F92EBA44AB69" blockId="5.[264,1323,1773,1831]" pageId="5" pageNumber="6">
<emphasis id="2DDA6747FF935A235708F92EBA82AB4B" bold="true" box="[264,432,1773,1797]" pageId="5" pageNumber="6">FIGURES 6­7.</emphasis>
<taxonomicName id="D8AEC0D6FF935A2357BEF92EB966AB4A" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[446,596,1773,1796]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF935A2357BEF92EB966AB4A" box="[446,596,1773,1796]" italics="true" pageId="5" pageNumber="6">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF935A23545BF92EB982AB4A" box="[603,688,1773,1796]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, lateral view. 6. Male holotype; 7. Female paratype. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="4BD1EBDDFF905A205708FC20B996A871" ID-DOI="http://doi.org/10.5281/zenodo.5014183" ID-Zenodo-Dep="5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="6" pageNumber="7" startId="6.[264,382,995,1019]" targetBox="[265,1321,310,970]" targetPageId="6">
<paragraph id="1F11BB55FF905A205708FC20B996A871" blockId="6.[264,1323,995,1087]" pageId="6" pageNumber="7">
<emphasis id="2DDA6747FF905A205708FC20BA8EAFB5" bold="true" box="[264,444,995,1019]" pageId="6" pageNumber="7">FIGURES 8­11.</emphasis>
<taxonomicName id="D8AEC0D6FF905A2057CCFC20B951AFB4" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[460,611,995,1018]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF905A2057CCFC20B951AFB4" box="[460,611,995,1018]" italics="true" pageId="6" pageNumber="7">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF905A20546BFC20B98DAFB4" box="[619,703,995,1018]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm).
</paragraph>
</caption>
<paragraph id="1F11BB55FF905A205738FB58B9E4A983" blockId="6.[264,1324,1179,1525]" pageId="6" pageNumber="7">
Legs with all claws smooth, without ventral dentition or lateral pegs (
<figureCitation id="8795A7D0FF905A205290FB58BE2CA8FB" box="[1168,1310,1179,1205]" captionStart="FIGURES 12­16" captionStartId="7.[264,383,1630,1654]" captionTargetBox="[270,1317,609,1603]" captionTargetId="figure-183@7.[270,1317,609,1603]" captionTargetPageId="7" captionText="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="6" pageNumber="7">Figs. 12­16</figureCitation>
). Surface of most articles clearly ornamented with granules, including metatarsi II and III; metatarsus of legs I and II with ornamentation only in the proximal half; tarsi without granules. Ventral side of tarsus I with a concentration of short sensory hairs occupying about half of the total tarsal length, not forming a distinct solea (
<figureCitation id="8795A7D0FF905A205219FAF8BF41A91B" box="[1049,1139,1339,1365]" captionStart="FIGURES 12­16" captionStartId="7.[264,383,1630,1654]" captionTargetBox="[270,1317,609,1603]" captionTargetId="figure-183@7.[270,1317,609,1603]" captionTargetPageId="7" captionText="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
). Tarsus IV of male entire, carrying a short, thick adenostyle projecting upward and slightly distally (
<figureCitation id="8795A7D0FF905A2052FBFAA0BA1AA9EB" captionStart="FIGURES 12­16" captionStartId="7.[264,383,1630,1654]" captionTargetBox="[270,1317,609,1603]" captionTargetId="figure-183@7.[270,1317,609,1603]" captionTargetPageId="7" captionText="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
). Leading edge of adenostyle base at about 40% of the tarsal length. Tarsus IV of the female without modifications (
<figureCitation id="8795A7D0FF905A205471FA70B9FAA983" box="[625,712,1459,1485]" captionStart="FIGURES 12­16" captionStartId="7.[264,383,1630,1654]" captionTargetBox="[270,1317,609,1603]" captionTargetId="figure-183@7.[270,1317,609,1603]" captionTargetPageId="7" captionText="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="6" pageNumber="7">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="1F11BB55FF905A205738FA18BF41A9BB" blockId="6.[264,1324,1179,1525]" box="[312,1139,1499,1525]" pageId="6" pageNumber="7">
Leg measurements of male
<typeStatus id="C01505F7FF905A20547CFA18B9D3A9BB" box="[636,737,1499,1525]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
in µm [length/width (L/W ratio)]:
</paragraph>
<paragraph id="1F11BB55FF905A20575AF9E8BE2FAA9B" blockId="6.[264,1315,1579,1749]" pageId="6" pageNumber="7">Tr Fe Pa Ti Mt Ta Total Leg I 224/210 (1.07) 605/171 (3.54) 468/171 (2.15) 447/171 (1.61) 289/145 (1.99) 500/184 (2.72) 2533 Leg II 132/140 (0.94) 421/132 (3.19) 228/140 (1.63) 281/140 (2.01) 211/105 (2.01) 360/123 (2.93) 1633 Leg III 226/170 (1.33) 396/151 (2.62) 255/151 (1.69) 321/179 (1/79) 245/132 (1.86) 415/142 (2.92) 1858 Leg IV 224/184 (1.22) 579/184 (3.15) 342/197 (1.74) 395/197 (2.01) 289/158 (1.83) 526/188 (2.80) 2355</paragraph>
<paragraph id="1F11BB55FF905A215738F8CDBAC6ADB0" blockId="6.[264,1323,1806,1872]" lastBlockId="7.[264,1324,284,510]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Penis small, typical of pettalids (
<figureCitation id="8795A7D0FF905A2054BBF8CDB875AB66" box="[699,839,1806,1832]" captionStart="FIGURES 25­28" captionStartId="10.[264,383,1797,1821]" captionTargetBox="[268,1323,738,1777]" captionTargetId="figure-205@10.[264,1323,729,1786]" captionTargetPageId="10" captionText="FIGURES 25­28. Rakaia macra sp. nov. apex of penis. 25. Dorsal view; 26. Dorsal view showing movable fingers (mf); 27. Ventral view; 28. Light microscope image, dorsal view. Scale bar 100 µm for all." figureDoi="http://doi.org/10.5281/zenodo.5014193" httpUri="https://zenodo.org/record/5014193/files/figure.png" pageId="6" pageNumber="7">Figs. 25­28</figureCitation>
). Setal formula 3, 6, 3 or 4 (two penises examined). Ventral side ornamented with tiny denticles along distolateral and distal margins. Three ventral setae set back far from distal margin, their bases separated by distances roughly equal to setal diameter. Rounded distal margin of penis with 6 apical setae. Dorsal side of penis with a group of 3 or 4 long setae on each side, their broad bases separated by distances roughly equal to setal diameter and arranged in a V. Gonopore complex with two movable fingers (m.f.) in the shape of pronounced hooks, with denticles at their bases (
<figureCitation id="8795A7D0FF915A215757FE27BAD5ADB0" box="[343,487,484,510]" captionStart="FIGURES 25­28" captionStartId="10.[264,383,1797,1821]" captionTargetBox="[268,1323,738,1777]" captionTargetId="figure-205@10.[264,1323,729,1786]" captionTargetPageId="10" captionText="FIGURES 25­28. Rakaia macra sp. nov. apex of penis. 25. Dorsal view; 26. Dorsal view showing movable fingers (mf); 27. Ventral view; 28. Light microscope image, dorsal view. Scale bar 100 µm for all." figureDoi="http://doi.org/10.5281/zenodo.5014193" httpUri="https://zenodo.org/record/5014193/files/figure.png" pageId="7" pageNumber="8">Figs. 26, 28</figureCitation>
).
</paragraph>
<caption id="4BD1EBDDFF915A215708F99DBF3AAA8F" ID-DOI="http://doi.org/10.5281/zenodo.5014185" ID-Zenodo-Dep="5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="7" pageNumber="8" startId="7.[264,383,1630,1654]" targetBox="[270,1317,609,1603]" targetPageId="7">
<paragraph id="1F11BB55FF915A215708F99DBF3AAA8F" blockId="7.[264,1323,1630,1729]" pageId="7" pageNumber="8">
<emphasis id="2DDA6747FF915A215708F99DBAF9AA38" bold="true" box="[264,459,1630,1654]" pageId="7" pageNumber="8">FIGURES 12­16.</emphasis>
<taxonomicName id="D8AEC0D6FF915A2157D9F99DB942AA3B" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[473,624,1630,1653]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF915A2157D9F99DB942AA3B" box="[473,624,1630,1653]" italics="true" pageId="7" pageNumber="8">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF915A215478F99DB9F9AA3B" box="[632,715,1630,1653]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm).
</paragraph>
</caption>
<paragraph id="1F11BB55FF915A2E5738F8E3B93AADC8" blockId="7.[264,1323,1824,1890]" lastBlockId="8.[264,1323,284,390]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Ovipositor (
<figureCitation id="8795A7D0FF915A2157C6F8E3B966AB74" box="[454,596,1824,1850]" captionStart="FIGURES 29­30" captionStartId="11.[264,383,1673,1697]" captionTargetBox="[493,1093,777,1642]" captionTargetId="figure-215@11.[493,1093,777,1642]" captionTargetPageId="11" captionText="FIGURES 29­30. Rakaia macra sp. nov. ovipositor. 29. Entire ovipositor structure (scale bar 400 µm); 30. Ovipositor tip (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014195" httpUri="https://zenodo.org/record/5014195/files/figure.png" pageId="7" pageNumber="8">Figs. 29, 30</figureCitation>
) wide, composed of two apical lobes and 29 circular articles, each of the latter furnished with eight setae. Apical lobes carrying several setae (increasing in length towards the tip); a long terminal seta rising from a small socket at the end of each lobe. Sensitive processes carrying a long, multibranched seta on the distolateral side of each terminal lobe.
</paragraph>
<caption id="4BD1EBDDFF9E5A2E5708F901B9C3AB0F" ID-DOI="http://doi.org/10.5281/zenodo.5014187" ID-Zenodo-Dep="5014187" httpUri="https://zenodo.org/record/5014187/files/figure.png" pageId="8" pageNumber="9" startId="8.[264,382,1730,1754]" targetBox="[264,1323,449,1704]" targetPageId="8">
<paragraph id="1F11BB55FF9E5A2E5708F901B9C3AB0F" blockId="8.[264,1323,1730,1857]" pageId="8" pageNumber="9">
<emphasis id="2DDA6747FF9E5A2E5708F901BAF9AA94" bold="true" box="[264,459,1730,1754]" pageId="8" pageNumber="9">FIGURES 17­22.</emphasis>
<taxonomicName id="D8AEC0D6FF9E5A2E57DCF900B946AA94" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[476,628,1731,1754]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9E5A2E57DCF900B946AA94" box="[476,628,1731,1754]" italics="true" pageId="8" pageNumber="9">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9E5A2E547CF901B9E2AA97" box="[636,720,1730,1753]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
17. Ozophore of male paratype (scale bar 50 µm); 18. Spiracle (scale bar 20 µm); 19. Gonostome of male paratype (scale bar 100 µm); 20. Gonostome of female paratype (scale bar 100 µm); 21. Anal region of male paratype (scale bar 400 µm); 22. Anal region of female paratype (scale bar 300 µm).
</paragraph>
</caption>
<paragraph id="1F11BB55FF9F5A2F5738FED8B98AAD10" blockId="9.[264,1324,283,550]" pageId="9" pageNumber="10">
<emphasis id="2DDA6747FF9F5A2F5738FED8BA88AD7B" bold="true" box="[312,442,283,309]" pageId="9" pageNumber="10">Variation:</emphasis>
Range of measurements: male (n = 6), female (n = 6) in parentheses: dorsal scutum length 2.24­2.50 (2.48­2.64).
</paragraph>
</subSubSection>
<subSubSection id="57B4E8DEFF9F5A2F5738FEA8B91DAE68" pageId="9" pageNumber="10" type="distribution">
<paragraph id="1F11BB55FF9F5A2F5738FEA8B91DAE68" blockId="9.[264,1324,283,550]" pageId="9" pageNumber="10">
<emphasis id="2DDA6747FF9F5A2F5738FEA8B945ADCB" bold="true" box="[312,631,363,389]" pageId="9" pageNumber="10">Distribution and habitat:</emphasis>
Known only from Waipori,
<collectingRegion id="DD6A75B7FF9F5A2F55D9FEAFBF16ADC8" box="[985,1060,364,390]" country="New Zealand" name="Otago" pageId="9" pageNumber="10">Otago</collectingRegion>
, South Island of
<collectingCountry id="67B9FBC5FF9F5A2F52F4FEAFBA58ADE0" name="New Zealand" pageId="9" pageNumber="10">New Zealand</collectingCountry>
. Collected by Ray Forster three times in 1977: March 18, March 21, and September 20. Although there is a scenic reserve at this site, no further information about these collections is available. A collection of specimens of an undescribed species is also known from this locality.
</paragraph>
</subSubSection>
<caption id="4BD1EBDDFF9F5A2F5708FAD4BF70A93D" ID-DOI="http://doi.org/10.5281/zenodo.5014189" ID-Zenodo-Dep="5014189" httpUri="https://zenodo.org/record/5014189/files/figure.png" pageId="9" pageNumber="10" startId="9.[264,382,1303,1327]" targetBox="[490,1097,609,1276]" targetPageId="9">
<paragraph id="1F11BB55FF9F5A2F5708FAD4BF70A93D" blockId="9.[264,1323,1303,1395]" pageId="9" pageNumber="10">
<emphasis id="2DDA6747FF9F5A2F5708FAD4BAFFA961" bold="true" box="[264,461,1303,1327]" pageId="9" pageNumber="10">FIGURES 23­24.</emphasis>
<taxonomicName id="D8AEC0D6FF9F5A2F57E0FAD4B94BA960" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[480,633,1303,1326]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9F5A2F57E0FAD4B94BA960" box="[480,633,1303,1326]" italics="true" pageId="9" pageNumber="10">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9F5A2F5482FAD4B9EAA960" box="[642,728,1303,1326]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
male paratype anal region. 23. Posterior view of anal region, showing scopulae on margin of the anal plate and on lobes of tergite 8 (scale bar 200 µm); 24. Dorsal angle, showing lobes and scopulae of tergite 8 (scale bar 300 µm).
</paragraph>
</caption>
<subSubSection id="57B4E8DEFF9F5A2A5738FA0DB9DCADEB" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="discussion">
<paragraph id="1F11BB55FF9F5A2F5738FA0DB999AB4F" blockId="9.[264,1324,1486,1873]" pageId="9" pageNumber="10">
<emphasis id="2DDA6747FF9F5A2F5738FA0DBA81A9A6" bold="true" box="[312,435,1486,1512]" pageId="9" pageNumber="10">Remarks:</emphasis>
<taxonomicName id="D8AEC0D6FF9F5A2F57BCFA0CB951A9A6" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[444,611,1487,1512]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9F5A2F57BCFA0CB951A9A6" box="[444,611,1487,1512]" italics="true" pageId="9" pageNumber="10">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9F5A2F546CFA0CB9FAA9A7" box="[620,712,1487,1513]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
clearly belongs to the family
<taxonomicName id="D8AEC0D6FF9F5A2F522FFA0CBF90A9A7" authorityName="Shear" authorityYear="1980" box="[1071,1186,1487,1513]" class="Arachnida" family="Pettalidae" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Pettalidae</taxonomicName>
because of the ozophore position and the modified anal region. However, the dentition of its chelicerae is unusual for this group; a homogeneous dentition has been previously reported only in the genera
<taxonomicName id="D8AEC0D6FF9F5A2F57F5F984B896AA2F" authority="Hansen &amp; Sorensen 1904" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[501,932,1607,1633]" class="Arachnida" family="Pettalidae" genus="Purcellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="2DDA6747FF9F5A2F57F5F984B953AA2E" box="[501,609,1607,1632]" italics="true" pageId="9" pageNumber="10">Purcellia</emphasis>
<bibRefCitation id="7B3FC6A4FF9F5A2F546CF984B896AA2F" author="Hansen, H. J. &amp; Sorensen, W." box="[620,932,1607,1633]" pageId="9" pageNumber="10" refId="ref4443" refString="Hansen, H. J. &amp; Sorensen, W. (1904) &quot; On Two Orders of Arachnida: Opiliones, Especially the Suborder Cyphophthalmi, and Ricinulei, Namely the Family Cryptostemmatoidae. &quot; Cambridge University Press, Cambridge." type="book" year="1904">Hansen &amp; Sørensen 1904</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D8AEC0D6FF9F5A2F55E7F984BA70AAC7" authority="Rosas Costa 1950" authorityName="Rosas Costa" authorityYear="1950" class="Arachnida" family="Pettalidae" genus="Parapurcellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="2DDA6747FF9F5A2F55E7F984BFB8AA2E" box="[999,1162,1607,1632]" italics="true" pageId="9" pageNumber="10">Parapurcellia</emphasis>
Rosas
<collectingRegion id="DD6A75B7FF9F5A2F52E8F984BE19AA2F" box="[1256,1323,1607,1633]" country="New Zealand" name="West Coast" pageId="9" pageNumber="10">Costa</collectingRegion>
1950
</taxonomicName>
among the
<taxonomicName id="D8AEC0D6FF9F5A2F57CEF9ACB977AAC7" authorityName="Shear" authorityYear="1980" box="[462,581,1647,1673]" class="Arachnida" family="Pettalidae" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Pettalidae</taxonomicName>
.
<taxonomicName id="D8AEC0D6FF9F5A2F5451F9ACB9C5AAC6" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[593,759,1647,1672]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9F5A2F5451F9ACB9C5AAC6" box="[593,759,1647,1672]" italics="true" pageId="9" pageNumber="10">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9F5A2F54FFF9ACB869AAC7" box="[767,859,1647,1673]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
resembles other
<taxonomicName id="D8AEC0D6FF9F5A2F5224F9ACBFE1AAC7" box="[1060,1235,1647,1673]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2DDA6747FF9F5A2F5224F9ACBF45AAC6" box="[1060,1143,1647,1672]" italics="true" pageId="9" pageNumber="10">Rakaia</emphasis>
species
</taxonomicName>
which, in ventral view, appear to have scopulae only on the posterior margin of the anal plate. These include
<taxonomicName id="D8AEC0D6FF9F5A2F57BBF97CB82BAA97" authority="Forster 1952" authorityName="Forster" authorityYear="1952" box="[443,793,1727,1753]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lindsayi">
<emphasis id="2DDA6747FF9F5A2F57BBF97CB944AA96" box="[443,630,1727,1752]" italics="true" pageId="9" pageNumber="10">Rakaia lindsayi</emphasis>
<bibRefCitation id="7B3FC6A4FF9F5A2F5480F97CB82BAA97" author="Forster, R. R." box="[640,793,1727,1753]" pageId="9" pageNumber="10" pagination="179 - 211" refId="ref4309" refString="Forster, R. R. (1952) Supplement to the sub-order Cyphophthalmi. Dominion Museum Records in Entomology, 1, 179 - 211." type="journal article" year="1952">Forster 1952</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D8AEC0D6FF9F5A2F5537F97CBF63AA97" authority="Forster 1948" authorityName="Forster" authorityYear="1948" box="[823,1105,1727,1753]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="2DDA6747FF9F5A2F5537F97CB898AA96" box="[823,938,1727,1752]" italics="true" pageId="9" pageNumber="10">R. magna</emphasis>
<bibRefCitation id="7B3FC6A4FF9F5A2F55B6F97CBF63AA97" author="Forster, R. R." box="[950,1105,1727,1753]" pageId="9" pageNumber="10" pagination="79 - 119" refId="ref4280" refString="Forster, R. R. (1948) The sub-order Cyphophthalmi Simon in New Zealand. Dominion Museum Records in Entomology, 1, 79 - 119." type="journal article" year="1948">Forster 1948</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D8AEC0D6FF9F5A2F5261F97CBA70AB4F" authority="Forster 1948" authorityName="Forster" authorityYear="1948" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="media">
<emphasis id="2DDA6747FF9F5A2F5261F97CBFFFAA96" box="[1121,1229,1727,1752]" italics="true" pageId="9" pageNumber="10">R. media</emphasis>
<bibRefCitation id="7B3FC6A4FF9F5A2F52D7F97CBA70AB4F" author="Forster, R. R." pageId="9" pageNumber="10" pagination="79 - 119" refId="ref4280" refString="Forster, R. R. (1948) The sub-order Cyphophthalmi Simon in New Zealand. Dominion Museum Records in Entomology, 1, 79 - 119." type="journal article" year="1948">Forster 1948</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="D8AEC0D6FF9F5A2F5783F924B994AB4F" authority="Forster 1948" authorityName="Forster" authorityYear="1948" box="[387,678,1767,1793]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="2DDA6747FF9F5A2F5783F924B934AB4E" box="[387,518,1767,1792]" italics="true" pageId="9" pageNumber="10">R. solitaria</emphasis>
<bibRefCitation id="7B3FC6A4FF9F5A2F540EF924B994AB4F" author="Forster, R. R." box="[526,678,1767,1793]" pageId="9" pageNumber="10" pagination="79 - 119" refId="ref4280" refString="Forster, R. R. (1948) The sub-order Cyphophthalmi Simon in New Zealand. Dominion Museum Records in Entomology, 1, 79 - 119." type="journal article" year="1948">Forster 1948</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="1F11BB55FF9F5A2C5738F8CCBA44AED0" blockId="9.[264,1324,1486,1873]" lastBlockId="10.[264,1324,284,670]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<taxonomicName id="D8AEC0D6FF9F5A2F5738F8CCBAECAB66" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[312,478,1807,1832]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9F5A2F5738F8CCBAECAB66" box="[312,478,1807,1832]" italics="true" pageId="9" pageNumber="10">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9F5A2F57E5F8CCB972AB67" box="[485,576,1807,1833]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from all other congeners by the presence of an outer lateral ridge on the second cheliceral article (
<figureCitation id="8795A7D0FF9F5A2F5558F8F4B8ACAB1F" box="[856,926,1847,1873]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
). Additionally, it is distinguished from
<taxonomicName id="D8AEC0D6FF9C5A2C574AFEDFBAFAAD7B" authorityName="Forster" authorityYear="1952" box="[330,456,284,309]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lindsayi">
<emphasis id="2DDA6747FF9C5A2C574AFEDFBAFAAD7B" box="[330,456,284,309]" italics="true" pageId="10" pageNumber="11">R. lindsayi</emphasis>
</taxonomicName>
by the uniform dentition in the mobile cheliceral digit (
<figureCitation id="8795A7D0FF9C5A2C5263FEDFBF9EAD78" box="[1123,1196,284,310]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
), the presence of a ventral process on the trochanter of the pedipalp (
<figureCitation id="8795A7D0FF9C5A2C55CCFE87BF10AD10" box="[972,1058,324,350]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="10" pageNumber="11">Fig. 11</figureCitation>
), and the absence of a ventral process on the chelicerae (
<figureCitation id="8795A7D0FF9C5A2C54ADFEAFB9CAADC8" box="[685,760,364,390]" captionStart="FIGURES 8­11" captionStartId="6.[264,382,995,1019]" captionTargetBox="[265,1321,310,970]" captionTargetId="figure-273@6.[265,1321,310,970]" captionTargetPageId="6" captionText="FIGURES 8­11. Rakaia macra sp. nov. 8. External view of left chelicera of male (scale bar 500 µm); 9. Dentition of chelicera (scale bar 100 µm); 10. Left palp of male (scale bar 500 µm); 11. Trochanter of palp (scale bar 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014183" httpUri="https://zenodo.org/record/5014183/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
).
<taxonomicName id="D8AEC0D6FF9C5A2C551FFEAFB8BDADCB" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[799,911,364,389]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9C5A2C551FFEAFB8BDADCB" box="[799,911,364,389]" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9C5A2C559BFEAFB8C8ADC8" box="[923,1018,364,390]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from
<taxonomicName id="D8AEC0D6FF9C5A2C5311FEAFBA6AADE3" authorityName="Forster" authorityYear="1948" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="2DDA6747FF9C5A2C5311FEAFBA6AADE3" italics="true" pageId="10" pageNumber="11">R. magna</emphasis>
</taxonomicName>
by size; specimens of
<taxonomicName id="D8AEC0D6FF9C5A2C5464FE57B9E6ADE3" authorityName="Forster" authorityYear="1948" box="[612,724,404,429]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="2DDA6747FF9C5A2C5464FE57B9E6ADE3" box="[612,724,404,429]" italics="true" pageId="10" pageNumber="11">R. magna</emphasis>
</taxonomicName>
are 3.0 mm or more in length, while all specimens of
<taxonomicName id="D8AEC0D6FF9C5A2C5728FE7FBAA6AD9B" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[296,404,444,469]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9C5A2C5728FE7FBAA6AD9B" box="[296,404,444,469]" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
are smaller than
<quantity id="D85616B0FF9C5A2C545FFE7FB9F5AD98" box="[607,711,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="11" unit="mm" value="2.7">2.70 mm</quantity>
in length.
<taxonomicName id="D8AEC0D6FF9C5A2C554FFE7FB889AD9B" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[847,955,444,469]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9C5A2C554FFE7FB889AD9B" box="[847,955,444,469]" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="D8AEC0D6FF9C5A2C52C2FE7FBE19AD9B" authorityName="Forster" authorityYear="1948" box="[1218,1323,444,469]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="media">
<emphasis id="2DDA6747FF9C5A2C52C2FE7FBE19AD9B" box="[1218,1323,444,469]" italics="true" pageId="10" pageNumber="11">R. media</emphasis>
</taxonomicName>
by the shape of the male tarsus IV;
<taxonomicName id="D8AEC0D6FF9C5A2C54ABFE27B821ADB3" authorityName="Forster" authorityYear="1948" box="[683,787,484,509]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="media">
<emphasis id="2DDA6747FF9C5A2C54ABFE27B821ADB3" box="[683,787,484,509]" italics="true" pageId="10" pageNumber="11">R. media</emphasis>
</taxonomicName>
has a pyriform male tarsus IV (sensu
<bibRefCitation id="7B3FC6A4FF9C5A2C52D8FE27BA74AE68" author="Forster, R. R." pageId="10" pageNumber="11" pagination="179 - 211" refId="ref4309" refString="Forster, R. R. (1952) Supplement to the sub-order Cyphophthalmi. Dominion Museum Records in Entomology, 1, 179 - 211." type="journal article" year="1952">Forster 1952</bibRefCitation>
), while the male tarsus IV of
<taxonomicName id="D8AEC0D6FF9C5A2C54A7FDCFB821AE6B" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[679,787,524,549]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9C5A2C54A7FDCFB821AE6B" box="[679,787,524,549]" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
tends to be elongate (sensu
<bibRefCitation id="7B3FC6A4FF9C5A2C5263FDCFBFCFAE68" author="Forster, R. R." box="[1123,1277,524,550]" pageId="10" pageNumber="11" pagination="179 - 211" refId="ref4309" refString="Forster, R. R. (1952) Supplement to the sub-order Cyphophthalmi. Dominion Museum Records in Entomology, 1, 179 - 211." type="journal article" year="1952">Forster 1952</bibRefCitation>
).
<taxonomicName id="D8AEC0D6FF9C5A2C5311FDCFBA61AE03" authorityName="Boyer &amp; Giribet" authorityYear="2003" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9C5A2C5311FDCFBA61AE03" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="D8AEC0D6FF9C5A2C545FFDF7B9D6AE03" authorityName="Forster" authorityYear="1948" box="[607,740,564,589]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="2DDA6747FF9C5A2C545FFDF7B9D6AE03" box="[607,740,564,589]" italics="true" pageId="10" pageNumber="11">R. solitaria</emphasis>
</taxonomicName>
by elongate vs. pyriform male tarsus IV, and the shape of the adenostyle, which is long and slender in
<taxonomicName id="D8AEC0D6FF9C5A2C5597FD9FBF2CAE3B" authorityName="Forster" authorityYear="1948" box="[919,1054,604,629]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="2DDA6747FF9C5A2C5597FD9FBF2CAE3B" box="[919,1054,604,629]" italics="true" pageId="10" pageNumber="11">R. solitaria</emphasis>
</taxonomicName>
but stout in
<taxonomicName id="D8AEC0D6FF9C5A2C52BCFD9FBE19AE3B" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[1212,1323,604,629]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9C5A2C52BCFD9FBE19AE3B" box="[1212,1323,604,629]" italics="true" pageId="10" pageNumber="11">R. macra</emphasis>
</taxonomicName>
(
<figureCitation id="8795A7D0FF9C5A2C5711FD47BA5AAED0" box="[273,360,644,670]" captionStart="FIGURES 12­16" captionStartId="7.[264,383,1630,1654]" captionTargetBox="[270,1317,609,1603]" captionTargetId="figure-183@7.[270,1317,609,1603]" captionTargetPageId="7" captionText="FIGURES 12­16. Rakaia macra sp. nov. tarsi. 12. Male left tarsus I (scale bar 300 µm) (12); 13. Male left tarsus II (scale bar 300 µm); 14. Male left tarsus III (scale bar 200 µm); 15. Male left tarsus IV (scale bar 300 µm); 16. Female left tarsus IV (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014185" httpUri="https://zenodo.org/record/5014185/files/figure.png" pageId="10" pageNumber="11">Fig. 15</figureCitation>
).
</paragraph>
<caption id="4BD1EBDDFF9C5A2C5708F8C6BA48AB2C" ID-DOI="http://doi.org/10.5281/zenodo.5014193" ID-Zenodo-Dep="5014193" httpUri="https://zenodo.org/record/5014193/files/figure.png" pageId="10" pageNumber="11" startId="10.[264,383,1797,1821]" targetBox="[268,1323,738,1777]" targetPageId="10">
<paragraph id="1F11BB55FF9C5A2C5708F8C6BA48AB2C" blockId="10.[264,1323,1797,1890]" pageId="10" pageNumber="11">
<emphasis id="2DDA6747FF9C5A2C5708F8C6BAF8AB53" bold="true" box="[264,458,1797,1821]" pageId="10" pageNumber="11">FIGURES 25­28.</emphasis>
<taxonomicName id="D8AEC0D6FF9C5A2C57D7F8C5B95CAB53" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[471,622,1798,1821]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9C5A2C57D7F8C5B95CAB53" box="[471,622,1798,1821]" italics="true" pageId="10" pageNumber="11">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9C5A2C5475F8C6B9F5AB52" box="[629,711,1797,1820]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
apex of penis. 25. Dorsal view; 26. Dorsal view showing movable fingers (mf); 27. Ventral view; 28. Light microscope image, dorsal view. Scale bar 100 µm for all.
</paragraph>
</caption>
<paragraph id="1F11BB55FF9D5A2D5738FEDFB9ECAED0" blockId="11.[264,1323,284,670]" pageId="11" pageNumber="12">
The penis of
<taxonomicName id="D8AEC0D6FF9D5A2D57D1FEDFB90FAD7B" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[465,573,284,309]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9D5A2D57D1FEDFB90FAD7B" box="[465,573,284,309]" italics="true" pageId="11" pageNumber="12">R. macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9D5A2D5444FEDFB9ACAD78" box="[580,670,284,310]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="8795A7D0FF9D5A2D54AEFEDFB804AD78" box="[686,822,284,310]" captionStart="FIGURES 25­28" captionStartId="10.[264,383,1797,1821]" captionTargetBox="[268,1323,738,1777]" captionTargetId="figure-205@10.[264,1323,729,1786]" captionTargetPageId="10" captionText="FIGURES 25­28. Rakaia macra sp. nov. apex of penis. 25. Dorsal view; 26. Dorsal view showing movable fingers (mf); 27. Ventral view; 28. Light microscope image, dorsal view. Scale bar 100 µm for all." figureDoi="http://doi.org/10.5281/zenodo.5014193" httpUri="https://zenodo.org/record/5014193/files/figure.png" pageId="11" pageNumber="12">Figs. 25­28</figureCitation>
) is very similar to that of several pettalids:
<taxonomicName id="D8AEC0D6FF9D5A2D5708FE87B97AAD10" authority="Juberthie 1989" authorityName="Juberthie" authorityYear="1989" box="[264,584,324,350]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="2DDA6747FF9D5A2D5708FE87BAA2AD13" box="[264,400,324,349]" italics="true" pageId="11" pageNumber="12">R. daviesae</emphasis>
<bibRefCitation id="7B3FC6A4FF9D5A2D5799FE87B97AAD10" author="Juberthie, C." box="[409,584,324,350]" pageId="11" pageNumber="12" pagination="499 - 507" refId="ref4552" refString="Juberthie, C. (1989) Rakaia daviesae sp. nov. (Opiliones, Cyphophthalmi, Pettalidae) from Australia. Memoirs of the Queensland Museum, 27, 499 - 507." type="journal article" year="1989">Juberthie 1989</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7B3FC6A4FF9D5A2D5459FE87B83CAD10" author="Juberthie, C." box="[601,782,324,350]" pageId="11" pageNumber="12" pagination="499 - 507" refId="ref4552" refString="Juberthie, C. (1989) Rakaia daviesae sp. nov. (Opiliones, Cyphophthalmi, Pettalidae) from Australia. Memoirs of the Queensland Museum, 27, 499 - 507." type="journal article" year="1989">Juberthie 1989</bibRefCitation>
: figure 7),
<taxonomicName id="D8AEC0D6FF9D5A2D5591FE87BA70ADC8" authority="Juberthie 1988" authorityName="Juberthie" authorityYear="1988" class="Arachnida" family="Pettalidae" genus="Austropurcellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="scoparia">
<emphasis id="2DDA6747FF9D5A2D5591FE87BF85AD13" box="[913,1207,324,349]" italics="true" pageId="11" pageNumber="12">Austropurcellia scoparia</emphasis>
<bibRefCitation id="7B3FC6A4FF9D5A2D52BFFE87BA70ADC8" author="Juberthie, C." pageId="11" pageNumber="12" pagination="133 - 140" refId="ref4517" refString="Juberthie, C. (1988) Un nouvel opilion cyphophthalme aveugle d'Australie: Austropurcellia gen. nov., scoparia n. sp. Memoires de Biospeologie, 15, 133 - 140." type="journal article" year="1988">Juberthie 1988</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7B3FC6A4FF9D5A2D5754FEAFB938ADC8" author="Juberthie, C." box="[340,522,364,390]" pageId="11" pageNumber="12" pagination="133 - 140" refId="ref4517" refString="Juberthie, C. (1988) Un nouvel opilion cyphophthalme aveugle d'Australie: Austropurcellia gen. nov., scoparia n. sp. Memoires de Biospeologie, 15, 133 - 140." type="journal article" year="1988">Juberthie 1988</bibRefCitation>
: figures 10­13),
<taxonomicName id="D8AEC0D6FF9D5A2D54D1FEAFBF61ADC8" authority="Shear 1993" authorityName="Shear" authorityYear="1993" box="[721,1107,364,390]" class="Arachnida" family="Pettalidae" genus="Chileogovea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jocasta">
<emphasis id="2DDA6747FF9D5A2D54D1FEAFB8F6ADCB" box="[721,964,364,389]" italics="true" pageId="11" pageNumber="12">Chileogovea jocasta</emphasis>
<bibRefCitation id="7B3FC6A4FF9D5A2D55CDFEAFBF61ADC8" author="Shear, W. A." box="[973,1107,364,390]" pageId="11" pageNumber="12" pagination="73 - 78" refId="ref4676" refString="Shear, W. A. (1993) The genus Chileogovea (Opiliones, Cyphophthalmi, Petallidae). Journal of Arachnology, 21, 73 - 78." type="journal article" year="1993">Shear 1993</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7B3FC6A4FF9D5A2D5266FEAFBFC1ADC8" author="Shear, W. A." box="[1126,1267,364,390]" pageId="11" pageNumber="12" pagination="73 - 78" refId="ref4676" refString="Shear, W. A. (1993) The genus Chileogovea (Opiliones, Cyphophthalmi, Petallidae). Journal of Arachnology, 21, 73 - 78." type="journal article" year="1993">Shear 1993</bibRefCitation>
: figures 9­10), and
<taxonomicName id="D8AEC0D6FF9D5A2D57C5FE57B856ADE0" authority="Roewer 1961" authorityName="Roewer" authorityYear="1961" box="[453,868,404,430]" class="Arachnida" family="Pettalidae" genus="Chileogovea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="2DDA6747FF9D5A2D57C5FE57B98EADE3" box="[453,700,404,429]" italics="true" pageId="11" pageNumber="12">Chileogovea oedipus</emphasis>
Roewer 1961
</taxonomicName>
(
<bibRefCitation id="7B3FC6A4FF9D5A2D5575FE57BF33ADE0" author="Shear, W. A." box="[885,1025,404,430]" pageId="11" pageNumber="12" pagination="73 - 78" refId="ref4676" refString="Shear, W. A. (1993) The genus Chileogovea (Opiliones, Cyphophthalmi, Petallidae). Journal of Arachnology, 21, 73 - 78." type="journal article" year="1993">Shear 1993</bibRefCitation>
: figures 15­16), but setal formula differs among the various species. For example, there are 8 or 9 dorsal setae in
<taxonomicName id="D8AEC0D6FF9D5A2D5311FE7FBA5CADB3" authorityName="Juberthie" authorityYear="1989" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="2DDA6747FF9D5A2D5311FE7FBA5CADB3" italics="true" pageId="11" pageNumber="12">R. daviesae</emphasis>
</taxonomicName>
but only 3 or
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<taxonomicName id="D8AEC0D6FF9D5A2D5449FE27B987ADB3" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[585,693,484,509]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9D5A2D5449FE27B987ADB3" box="[585,693,484,509]" italics="true" pageId="11" pageNumber="12">R. macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9D5A2D54BCFE27B827ADB3" box="[700,789,484,510]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
The ventral setae of the penis are also found in different numbers in the two species:
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<taxonomicName id="D8AEC0D6FF9D5A2D551FFDCFB895AE6B" authorityName="Juberthie" authorityYear="1989" box="[799,935,524,549]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="2DDA6747FF9D5A2D551FFDCFB895AE6B" box="[799,935,524,549]" italics="true" pageId="11" pageNumber="12">R. daviesae</emphasis>
</taxonomicName>
, but
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<taxonomicName id="D8AEC0D6FF9D5A2D521BFDCFBFBAAE6B" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[1051,1160,524,549]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9D5A2D521BFDCFBFBAAE6B" box="[1051,1160,524,549]" italics="true" pageId="11" pageNumber="12">R. macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9D5A2D5291FDCFBFDEAE6B" box="[1169,1260,524,550]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
The mobile digit with two prominent hooks is present in all of the species listed above, but is absent in the pettalid species
<taxonomicName id="D8AEC0D6FF9D5A2D546BFD9FBFBEAE38" authority="Hansen &amp; Sorensen 1904" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[619,1164,604,630]" class="Arachnida" family="Pettalidae" genus="Purcellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="illustrans">
<emphasis id="2DDA6747FF9D5A2D546BFD9FB862AE3B" box="[619,848,604,629]" italics="true" pageId="11" pageNumber="12">Purcellia illustrans</emphasis>
<bibRefCitation id="7B3FC6A4FF9D5A2D5559FD9FBFBEAE38" author="Hansen, H. J. &amp; Sorensen, W." box="[857,1164,604,630]" pageId="11" pageNumber="12" refId="ref4443" refString="Hansen, H. J. &amp; Sorensen, W. (1904) &quot; On Two Orders of Arachnida: Opiliones, Especially the Suborder Cyphophthalmi, and Ricinulei, Namely the Family Cryptostemmatoidae. &quot; Cambridge University Press, Cambridge." type="book" year="1904">Hansen &amp; Sørensen 1904</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7B3FC6A4FF9D5A2D529DFD9FBA88AED0" author="Hansen, H. J. &amp; Sorensen, W." pageId="11" pageNumber="12" refId="ref4443" refString="Hansen, H. J. &amp; Sorensen, W. (1904) &quot; On Two Orders of Arachnida: Opiliones, Especially the Suborder Cyphophthalmi, and Ricinulei, Namely the Family Cryptostemmatoidae. &quot; Cambridge University Press, Cambridge." type="book" year="1904">Hansen and Sørensen 1904</bibRefCitation>
: Plate IV, figures 1o­1q).
</paragraph>
<caption id="4BD1EBDDFF9D5A2D5708F94AB9E4AA8D" ID-DOI="http://doi.org/10.5281/zenodo.5014195" ID-Zenodo-Dep="5014195" httpUri="https://zenodo.org/record/5014195/files/figure.png" pageId="11" pageNumber="12" startId="11.[264,383,1673,1697]" targetBox="[493,1093,777,1642]" targetPageId="11">
<paragraph id="1F11BB55FF9D5A2D5708F94AB9E4AA8D" blockId="11.[264,1323,1673,1731]" pageId="11" pageNumber="12">
<emphasis id="2DDA6747FF9D5A2D5708F94ABAF9AAEF" bold="true" box="[264,459,1673,1697]" pageId="11" pageNumber="12">FIGURES 29­30.</emphasis>
<taxonomicName id="D8AEC0D6FF9D5A2D57D8F94AB95DAAEE" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[472,623,1673,1696]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9D5A2D57D8F94AB95DAAEE" box="[472,623,1673,1696]" italics="true" pageId="11" pageNumber="12">Rakaia macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9D5A2D5476F94AB9FAAAEE" box="[630,712,1673,1696]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
ovipositor. 29. Entire ovipositor structure (scale bar 400 µm); 30. Ovipositor tip (scale bar 200 µm).
</paragraph>
</caption>
<paragraph id="1F11BB55FF9D5A2D5738F8D7BE19AB60" blockId="11.[264,1323,1812,1875]" box="[312,1323,1812,1838]" pageId="11" pageNumber="12">
The ovipositors of
<taxonomicName id="D8AEC0D6FF9D5A2D541DF8D7B9F6AB63" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[541,708,1812,1837]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="2DDA6747FF9D5A2D541DF8D7B9F6AB63" box="[541,708,1812,1837]" italics="true" pageId="11" pageNumber="12">Rakaia macra</emphasis>
</taxonomicName>
and
<taxonomicName id="D8AEC0D6FF9D5A2D5501F8D7B8F6AB63" authorityName="Juberthie" authorityYear="1989" box="[769,964,1812,1837]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="2DDA6747FF9D5A2D5501F8D7B8F6AB63" box="[769,964,1812,1837]" italics="true" pageId="11" pageNumber="12">Rakaia daviesae</emphasis>
</taxonomicName>
appear nearly identical (
<figureCitation id="8795A7D0FF9D5A2D52F0F8D7BE19AB60" box="[1264,1323,1812,1838]" captionStart="FIGURES 29­30" captionStartId="11.[264,383,1673,1697]" captionTargetBox="[493,1093,777,1642]" captionTargetId="figure-215@11.[493,1093,777,1642]" captionTargetPageId="11" captionText="FIGURES 29­30. Rakaia macra sp. nov. ovipositor. 29. Entire ovipositor structure (scale bar 400 µm); 30. Ovipositor tip (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014195" httpUri="https://zenodo.org/record/5014195/files/figure.png" pageId="11" pageNumber="12">Figs.</figureCitation>
</paragraph>
<paragraph id="1F11BB55FF9D5A2A5708F8FAB9DCADEB" blockId="11.[264,1323,1812,1875]" lastBlockId="12.[264,1323,284,422]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
29­30,
<bibRefCitation id="7B3FC6A4FF9D5A2D575BF8FAB93EAB1D" author="Juberthie, C." box="[347,524,1849,1875]" pageId="11" pageNumber="12" pagination="499 - 507" refId="ref4552" refString="Juberthie, C. (1989) Rakaia daviesae sp. nov. (Opiliones, Cyphophthalmi, Pettalidae) from Australia. Memoirs of the Queensland Museum, 27, 499 - 507." type="journal article" year="1989">Juberthie 1989</bibRefCitation>
: figure 7) except in the number of circular articles:
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<taxonomicName id="D8AEC0D6FF9D5A2D52A4F8F9BE19AB1D" authorityName="Juberthie" authorityYear="1989" box="[1188,1323,1850,1875]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="daviesae">
<emphasis id="2DDA6747FF9D5A2D52A4F8F9BE19AB1D" box="[1188,1323,1850,1875]" italics="true" pageId="11" pageNumber="12">R. daviesae</emphasis>
</taxonomicName>
but
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<taxonomicName id="D8AEC0D6FF9A5A2A5777FEDFBAD1AD7B" authority="Boyer &amp; Giribet, 2003" authorityName="Boyer &amp; Giribet" authorityYear="2003" box="[375,483,284,309]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macra" status="sp. nov.">
<emphasis id="2DDA6747FF9A5A2A5777FEDFBAD1AD7B" box="[375,483,284,309]" italics="true" pageId="12" pageNumber="13">R. macra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E9DA3CFF9A5A2A57EAFEDFB971AD7B" box="[490,579,284,310]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
The sensitive processes and general configuration of the distal end of the first article and lobes appears identical to that illustrated for
<taxonomicName id="D8AEC0D6FF9A5A2A528BFE81BFC6AD15" authorityName="Juberthie" authorityYear="1960" box="[1163,1268,322,347]" class="Arachnida" family="Sironidae" genus="Metasiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="2DDA6747FF9A5A2A528BFE81BFC6AD15" box="[1163,1268,322,347]" italics="true" pageId="12" pageNumber="13">Metasiro</emphasis>
</taxonomicName>
and described as typical of
<emphasis id="2DDA6747FF9A5A2A541BFEA4B8FEADCE" box="[539,972,359,384]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="D8AEC0D6FF9A5A2A541BFEA4B980ADCE" authorityName="Roewer" authorityYear="1961" box="[539,690,359,384]" class="Arachnida" family="Pettalidae" genus="Chileogovea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Chileogovea</taxonomicName>
,
<taxonomicName id="D8AEC0D6FF9A5A2A54C0FEA4B854ADCE" box="[704,870,359,384]" class="Arachnida" family="Pettalidae" genus="Parapurcellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Parapurcellia</taxonomicName>
,
<taxonomicName id="D8AEC0D6FF9A5A2A5572FEA4B8FAADCE" authorityName="Hirst" authorityYear="1925" box="[882,968,359,384]" class="Arachnida" family="Pettalidae" genus="Rakaia" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Rakaia</taxonomicName>
,
</emphasis>
and the majority of
<taxonomicName id="D8AEC0D6FF9A5A2A52C1FEA4BA07ADE8" class="Arachnida" family="Sironidae" genus="Siro" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2DDA6747FF9A5A2A52C1FEA4BFC3ADCE" box="[1217,1265,359,384]" italics="true" pageId="12" pageNumber="13">Siro</emphasis>
species
</taxonomicName>
(
<figureCitation id="8795A7D0FF9A5A2A5746FE4FBAAEADE8" box="[326,412,396,422]" captionStart="FIGURES 29­30" captionStartId="11.[264,383,1673,1697]" captionTargetBox="[493,1093,777,1642]" captionTargetId="figure-215@11.[493,1093,777,1642]" captionTargetPageId="11" captionText="FIGURES 29­30. Rakaia macra sp. nov. ovipositor. 29. Entire ovipositor structure (scale bar 400 µm); 30. Ovipositor tip (scale bar 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.5014195" httpUri="https://zenodo.org/record/5014195/files/figure.png" pageId="12" pageNumber="13">Fig. 30</figureCitation>
,
<bibRefCitation id="7B3FC6A4FF9A5A2A57A9FE4FB968ADE8" author="Juberthie, C." box="[425,602,396,422]" pageId="12" pageNumber="13" pagination="1371 - 1390" refId="ref4486" refString="Juberthie, C. (1970) Les genres d'opilions sironinae (Cyphophthalmes). Bulletin du Museum National d'Histoire Naturelle, 41 (6), 1371 - 1390." type="journal article" year="1970">Juberthie 1970</bibRefCitation>
: figure 12c).
</paragraph>
</subSubSection>
</treatment>
</document>
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