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<document id="A8360FD08B4BB13090267BE23B6F32AA" ID-CLB-Dataset="30935" ID-DOI="10.11646/zootaxa.4375.2.1" ID-GBIF-Dataset="c8d905a3-01d4-4d5f-a904-fad72add2ae1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1158445" ID-ZooBank="7A4595B8-9FDF-41A9-9C39-A8A4A83FF302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1516789970485" checkinUser="plazi" docAuthor="Phan, Quoc Toan, Kompier, Tom, Karube, Haruki &amp; Hayashi, Fumio" docDate="2018" docId="006F87D3FFF9EC07B594F9A5FC43D236" docLanguage="en" docName="zootaxa.4375.2.1.pdf" docOrigin="Zootaxa 4375 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euphaea saola Phan &amp; Kompier &amp; Karube &amp; Hayashi 2018, sp. nov." docType="treatment" docUuid="2C91F371-197F-4D4C-9258-446CEA8FAC1B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="182" masterDocId="FC56FFABFFE5EC18B503FF87FFBFD376" masterDocTitle="A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea" masterLastPageNumber="190" masterPageNumber="151" pageNumber="179" updateTime="1698516995768" updateUser="ExternalLinkService">
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<mods:title id="5EA4305EC5BDC8113761D89A27E8E81A">A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea</mods:title>
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<mods:namePart id="859C37C4C993BA7ED717568AA93777D5">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="90B66470FBEF5E43D09C4E2B870FDC5E">Kompier, Tom</mods:namePart>
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<mods:namePart id="6E9FF4DB73AD5E479F865C40BAA78003">Karube, Haruki</mods:namePart>
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<heading id="D33181A9FFF9EC04B594F9A5FD32D54B" bold="true" box="[151,653,1570,1597]" fontSize="11" level="1" pageId="28" pageNumber="179" reason="1">
<emphasis id="BAB2EAD7FFF9EC04B594F9A5FD32D54B" bold="true" box="[151,653,1570,1597]" pageId="28" pageNumber="179">
<taxonomicName id="4FC64D46FFF9EC04B594F9A5FD9DD54B" ID-CoL="9X2NV" authority="Phan &amp; Hayashi" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[151,546,1570,1597]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="saola" status="sp. nov.">
<emphasis id="BAB2EAD7FFF9EC04B594F9A5FEF4D54A" bold="true" box="[151,331,1570,1596]" italics="true" pageId="28" pageNumber="179">Euphaea saola</emphasis>
Phan &amp; Hayashi
</taxonomicName>
,
<taxonomicNameLabel id="A18157ACFFF9EC04B72CF9A4FD32D54B" box="[559,653,1571,1597]" pageId="28" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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(
<figureCitation id="10FD2A40FFF9EC04B59CF9C2FEBBD528" box="[159,260,1605,1630]" captionStart="FIGURE 14" captionStartId="23.[151,250,1889,1911]" captionTargetBox="[220,1341,852,1868]" captionTargetId="figure@23.[184,1403,844,1868]" captionTargetPageId="23" captionText="FIGURE 14. Dorsal view of wings of male Euphaea spp. (14a) E. hirta; (14b) E. masoni; (14c) E. guerini; (14d) E. saola sp. nov." httpUri="https://zenodo.org/record/3096097/files/figure.png" pageId="28" pageNumber="179">Figs 14d</figureCitation>
,
<figureCitation id="10FD2A40FFF9EC04B40CF9C2FE85D528" box="[271,314,1605,1630]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="28" pageNumber="179">15a</figureCitation>
,
<figureCitation id="10FD2A40FFF9EC04B445F9C2FECED528" box="[326,369,1605,1630]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="28" pageNumber="179">16e</figureCitation>
,
<figureCitation id="10FD2A40FFF9EC04B47FF9C2FE44D528" box="[380,507,1605,1630]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="28" pageNumber="179">
18ab,
<quantity id="4F3E9B20FFF9EC04B4CCF9C1FE44D528" box="[463,507,1606,1630]" metricMagnitude="-2" metricUnit="kg" metricValue="1.8" pageId="28" pageNumber="179" unit="g" value="18.0">18g</quantity>
</figureCitation>
,
<figureCitation id="10FD2A40FFF9EC04B704F9C2FDF2D528" box="[519,589,1605,1630]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="28" pageNumber="179">20ad</figureCitation>
)
</paragraph>
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<subSubSection id="C0DC654EFFF9EC04B594F90AFA26D5B4" pageId="28" pageNumber="179" type="reference_group">
<paragraph id="887936C5FFF9EC04B594F90AFA26D5B4" blockId="28.[151,1436,1677,1730]" pageId="28" pageNumber="179">
<taxonomicName id="4FC64D46FFF9EC04B594F90AFEFCD5D5" ID-CoL="3CLZ7" authorityName=": Rambur" authorityYear="1842" box="[151,323,1677,1699]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFF9EC04B594F90AFEFCD5D5" box="[151,323,1677,1699]" italics="true" pageId="28" pageNumber="179">Euphaea guerini</emphasis>
</taxonomicName>
[nec.
<bibRefCitation id="EC574B34FFF9EC04B48BF90AFD9FD5D5" author="Rambur" box="[392,544,1677,1699]" pageId="28" pageNumber="179" refString="Rambur, P. (1842) Histoire naturelle des insectes: Nevropteres. Librairie Encyclopedique de Roret, Paris, 534 pp., 12 pls." type="book" year="1842">Rambur, 1842</bibRefCitation>
]: van
<bibRefCitation id="EC574B34FFF9EC04B762F90AFCDDD5D5" author="Tol" box="[609,866,1677,1699]" pageId="28" pageNumber="179" refString="van Tol, J. &amp; Rozendaal, F. G. (1995) Records of Calopterygoidea from Vietnam, with descriptions of two new species (Zygoptera: Amphipterygidae, Calopterygidae, Chlorocyphidae, Euphaeidae). Odonatologica, 24, 89 - 107." type="journal article" year="1995">Tol &amp; Rozendaal (1995)</bibRefCitation>
, p. 102 (
<figureCitation id="10FD2A40FFF9EC04B6C1F90AFBB1D5D5" box="[962,1038,1677,1699]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="28" pageNumber="179">Fig. 20</figureCitation>
) [Illustration of S910 of unspecified locality in
<collectingCountry id="F0D17655FFF9EC04B430F92BFE30D5B4" box="[307,399,1708,1730]" name="Vietnam" pageId="28" pageNumber="179">Vietnam</collectingCountry>
];
<bibRefCitation id="EC574B34FFF9EC04B49DF92BFD44D5B4" author="Karjalainen" box="[414,763,1708,1730]" pageId="28" pageNumber="179" refString="Karjalainen, S. &amp; Hamalainen, M. (2013) Demoiselle Damselflies: Winged Jewels of Silvery streams. Caloptera Publishing, Helsinki, 223 pp. [bilingual, in Finnish and English]" type="book" year="2013">Karjalainen &amp; Hämäläinen (2013)</bibRefCitation>
, p. 202) [Photo of male from Lak Sao, Bolikhamxay Prov.,
<collectingCountry id="F0D17655FFF9EC04B05BF92BFA30D5B4" box="[1368,1423,1708,1730]" name="Laos" pageId="28" pageNumber="179">Laos</collectingCountry>
].
</paragraph>
</subSubSection>
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<materialsCitation id="38AE3C98FFF9EC04B594F969FE2CD405" ID-GBIF-Occurrence="1805098036" collectingDate="2015-09-18" collectionCode="TMUZ" collectorName="A Luoi Nature Reserve &amp; F. Hayashi &amp; Deposited &amp; Phan &amp; Hayashi &amp; H. Karube" country="Japan" county="Thua Thien-Hue Prov." location="Thua Thien-Hue Prov." municipality="Systematic Zoology Institute in Tokyo Metropolitan University" pageId="28" pageNumber="179" specimenCount="5" specimenCount-male="5" stateProvince="Tokyo" typeStatus="holotype">
<emphasis id="BAB2EAD7FFF9EC04B594F969FEB4D471" bold="true" box="[151,267,1774,1799]" pageId="28" pageNumber="179">
<typeStatus id="577D8867FFF9EC04B594F969FEB9D471" box="[151,262,1774,1799]" pageId="28" pageNumber="179" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9EC0FD4CFFF9EC04B412F968FE97D470" box="[273,296,1775,1798]" pageId="28" pageNumber="179" type="male"></specimenCount>
,
<collectorName id="25335313FFF9EC04B437F968FD85D470" box="[308,570,1774,1799]" pageId="28" pageNumber="179">A Luoi Nature Reserve</collectorName>
,
<location id="8D19601EFFF9EC04B747F969FCFED470" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B747F969FCFED470" box="[580,833,1774,1799]" country="Japan" county="Thua Thien-Hue Prov." municipality="Systematic Zoology Institute in Tokyo Metropolitan University" name="Thua Thien-Hue Prov." pageId="28" pageNumber="179" stateProvince="Tokyo">Thua Thien-Hue Prov.</location>
,
<collectingDate id="EC3CE9EDFFF9EC04B64FF969FC72D471" box="[844,973,1774,1799]" pageId="28" pageNumber="179" value="2015-09-18">
<date id="FC781005FFF9EC04B64FF969FC72D471" box="[844,973,1774,1799]" pageId="28" pageNumber="179" value="2015-09-18">18.IX.2015</date>
</collectingDate>
, Q.T.
<location id="8D19601EFFF9EC04B112F969FBF6D471" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B112F969FBF6D471" box="[1041,1097,1774,1799]" country="Japan" county="Thua Thien-Hue Prov." municipality="Systematic Zoology Institute in Tokyo Metropolitan University" name="Phan" pageId="28" pageNumber="179" stateProvince="Tokyo">Phan</location>
and
<collectorName id="25335313FFF9EC04B17DF968FB4AD471" box="[1150,1269,1774,1799]" pageId="28" pageNumber="179">F. Hayashi</collectorName>
leg.
<collectorName id="25335313FFF9EC04B028F969FA22D471" box="[1323,1437,1774,1799]" pageId="28" pageNumber="179">Deposited</collectorName>
at
<location id="8D19601EFFF9EC04B5B1F895FCB2D45D" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B5B1F895FCB2D45D" box="[178,781,1810,1835]" country="Japan" county="Thua Thien-Hue Prov." municipality="Systematic Zoology Institute in Tokyo Metropolitan University" name="Zoological Collection of Systematic Zoology Institute" pageId="28" pageNumber="179" stateProvince="Tokyo">
Zoological Collection of
<collectingMunicipality id="681DACBFFFF9EC04B4C9F895FCB2D45D" box="[458,781,1810,1835]" pageId="28" pageNumber="179">Systematic Zoology Institute</collectingMunicipality>
</location>
in Tokyo Metropolitan University,
<collectingRegion id="4A02F827FFF9EC04B19BF895FB5DD45D" box="[1176,1250,1810,1835]" country="Japan" name="Tokyo" pageId="28" pageNumber="179">Tokyo</collectingRegion>
,
<collectingCountry id="F0D17655FFF9EC04B1EEF894FA92D45C" box="[1261,1325,1811,1834]" name="Japan" pageId="28" pageNumber="179">Japan</collectingCountry>
(
<collectionCode id="EED7AE00FFF9EC04B038F894FA2ED45D" box="[1339,1425,1811,1835]" pageId="28" pageNumber="179">TMUZ</collectionCode>
).
<emphasis id="BAB2EAD7FFF9EC04B5C4F8B1FE36D439" bold="true" box="[199,393,1846,1871]" pageId="28" pageNumber="179">
Male
<typeStatus id="577D8867FFF9EC04B40FF8B1FE3BD439" box="[268,388,1846,1871]" pageId="28" pageNumber="179" type="paratype">paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9EC0FD4CFFF9EC04B492F8B0FE0AD439" box="[401,437,1847,1871]" pageId="28" pageNumber="179" type="male">4♂</specimenCount>
,
<collectorName id="25335313FFF9EC04B4C0F8B0FD72D438" box="[451,717,1846,1871]" pageId="28" pageNumber="179">A Luoi Nature Reserve</collectorName>
,
<collectingCounty id="61184E49FFF9EC04B7DAF8B1FC66D438" box="[729,985,1846,1871]" pageId="28" pageNumber="179">Thua Thien-Hue Prov.</collectingCounty>
,
<collectingDate id="EC3CE9EDFFF9EC04B6E6F8B1FBD8D439" box="[997,1127,1846,1871]" pageId="28" pageNumber="179" value="2015-09-18">
<date id="FC781005FFF9EC04B6E6F8B1FBD8D439" box="[997,1127,1846,1871]" pageId="28" pageNumber="179" value="2015-09-18">18.IX.2015</date>
</collectingDate>
, Q.T.
<collectorName id="25335313FFF9EC04B1AEF8B1FB57D439" box="[1197,1256,1846,1871]" pageId="28" pageNumber="179">Phan</collectorName>
, F.
<collectorName id="25335313FFF9EC04B013F8B1FAD3D439" box="[1296,1388,1846,1871]" pageId="28" pageNumber="179">Hayashi</collectorName>
and
<collectorName id="25335313FFF9EC04B594F8DCFEB3D405" box="[151,268,1882,1907]" pageId="28" pageNumber="179">H. Karube</collectorName>
leg. (FHC).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFF9EC05B5C4F8F9FB36D212" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179" type="description">
<paragraph id="887936C5FFF9EC05B5C4F8F9FB36D212" blockId="28.[151,1437,1774,2015]" lastBlockId="29.[151,1436,151,357]" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179">
<materialsCitation id="38AE3C98FFF9EC04B5C4F8F9FAA6D4CD" ID-GBIF-Occurrence="1805098045" collectingDate="2015-09-18" collectingDateMax="2016-06" collectingDateMin="2015-09-18" collectorName="F. Hayashi &amp; T. Kompier" country="Vietnam" county="Female" location="Thua Thien-Hue Prov." municipality="Luoi Nature Reserve" pageId="28" pageNumber="179" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<emphasis id="BAB2EAD7FFF9EC04B5C4F8F9FE28D4E1" bold="true" box="[199,407,1918,1943]" pageId="28" pageNumber="179">
<collectingCounty id="61184E49FFF9EC04B5C4F8F9FEA1D4E1" box="[199,286,1918,1943]" pageId="28" pageNumber="179">Female</collectingCounty>
<typeStatus id="577D8867FFF9EC04B426F8F9FE2DD4E1" box="[293,402,1918,1943]" pageId="28" pageNumber="179" type="paratype">paratype</typeStatus>
.
</emphasis>
<specimenCount id="9EC0FD4CFFF9EC04B49DF8F8FE7ED4E1" box="[414,449,1919,1943]" pageId="28" pageNumber="179" type="female">1♀</specimenCount>
, A
<collectingMunicipality id="681DACBFFFF9EC04B4E9F8F9FD69D4E0" box="[490,726,1918,1943]" pageId="28" pageNumber="179">Luoi Nature Reserve</collectingMunicipality>
,
<location id="8D19601EFFF9EC04B7E2F8F9FC5FD4E0" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B7E2F8F9FC5FD4E0" box="[737,992,1918,1943]" country="Vietnam" county="Female" municipality="Luoi Nature Reserve" name="Thua Thien-Hue Prov." pageId="28" pageNumber="179">Thua Thien-Hue Prov.</location>
,
<collectingDate id="EC3CE9EDFFF9EC04B6E9F8F9FBD3D4E1" box="[1002,1132,1918,1943]" pageId="28" pageNumber="179" value="2015-09-18">
<date id="FC781005FFF9EC04B6E9F8F9FBD3D4E1" box="[1002,1132,1918,1943]" pageId="28" pageNumber="179" value="2015-09-18">18.IX.2015</date>
</collectingDate>
,
<collectorName id="25335313FFF9EC04B17BF8F8FB50D4E1" box="[1144,1263,1918,1943]" pageId="28" pageNumber="179">F. Hayashi</collectorName>
leg. (FHC).
<emphasis id="BAB2EAD7FFF9EC04B5C4F825FDB4D4CD" bold="true" box="[199,523,1954,1979]" pageId="28" pageNumber="179">
<location id="8D19601EFFF9EC04B5C4F825FEB0D4CD" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B5C4F825FEB0D4CD" box="[199,271,1954,1979]" country="Vietnam" county="Female" municipality="Luoi Nature Reserve" name="Other" pageId="28" pageNumber="179">Other</location>
materials examined.
</emphasis>
[
<collectingCountry id="F0D17655FFF9EC04B719F825FDC0D4CD" box="[538,639,1954,1979]" name="Vietnam" pageId="28" pageNumber="179">Vietnam</collectingCountry>
]
<specimenCount id="9EC0FD4CFFF9EC04B78CF824FD0BD4CD" box="[655,692,1955,1979]" pageId="28" pageNumber="179" type="male">2♂</specimenCount>
,
<location id="8D19601EFFF9EC04B7C1F824FC2CD4CC" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF9EC04B7C1F824FC2CD4CC" box="[706,915,1954,1979]" country="Vietnam" county="Female" municipality="Luoi Nature Reserve" name="Quang Binh Prov." pageId="28" pageNumber="179">Quang Binh Prov.</location>
,
<collectingDate id="EC3CE9EDFFF9EC04B6A3F825FBBFD4CD" box="[928,1024,1954,1979]" pageId="28" pageNumber="179" value="2016-06">
<date id="FC781005FFF9EC04B6A3F825FBBFD4CD" box="[928,1024,1954,1979]" pageId="28" pageNumber="179" value="2016-06">VI.2016</date>
</collectingDate>
,
<collectorName id="25335313FFF9EC04B10EF824FB2FD4CD" box="[1037,1168,1954,1979]" pageId="28" pageNumber="179">T. Kompier</collectorName>
leg. (TKC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF9EC04B024F824FDEBD4A9" ID-GBIF-Occurrence="1805098051" collectingDate="2016-06" collectorName="T. Kompier" country="Vietnam" location="Quang Tri" pageId="28" pageNumber="179" specimenCount="1" stateProvince="Quang Tri" typeStatus="paratype">
<collectingRegion id="4A02F827FFF9EC04B024F824FA23D4CD" box="[1319,1436,1954,1979]" country="Vietnam" name="Quang Tri" pageId="28" pageNumber="179">Quang Tri</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF9EC04B5E3F841FE80D4A9" box="[224,319,1990,2015]" pageId="28" pageNumber="179" value="2016-06">
<date id="FC781005FFF9EC04B5E3F841FE80D4A9" box="[224,319,1990,2015]" pageId="28" pageNumber="179" value="2016-06">VI.2016</date>
</collectingDate>
,
<collectorName id="25335313FFF9EC04B44FF840FE71D4A9" box="[332,462,1990,2015]" pageId="28" pageNumber="179">T. Kompier</collectorName>
leg. (TKC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF9EC04B762F841FB5ED4A9" ID-GBIF-Occurrence="1805098059" box="[609,1249,1990,2015]" collectingDate="2016-06" collectorName="T. Kompier" country="Vietnam" location="Thua Thien-Hue" pageId="28" pageNumber="179" specimenCount="1" stateProvince="Thua Thien-Hue" typeStatus="paratype">
<collectingRegion id="4A02F827FFF9EC04B762F841FCA4D4A9" box="[609,795,1990,2015]" country="Vietnam" name="Thua Thien-Hue" pageId="28" pageNumber="179">Thua Thien-Hue</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF9EC04B66EF841FC73D4A9" box="[877,972,1990,2015]" pageId="28" pageNumber="179" value="2016-06">
<date id="FC781005FFF9EC04B66EF841FC73D4A9" box="[877,972,1990,2015]" pageId="28" pageNumber="179" value="2016-06">VI.2016</date>
</collectingDate>
,
<collectorName id="25335313FFF9EC04B6DBF840FBE5D4A9" box="[984,1114,1990,2015]" pageId="28" pageNumber="179">T. Kompier</collectorName>
leg. (TKC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF9EC05B1EDF840FC5BD3C6" ID-GBIF-Occurrence="1805098067" collectingDate="2015-09-18" collectorName="A Luoi Nature Reserve &amp; F. Hayashi" country="Vietnam" lastPageId="29" lastPageNumber="180" location="Thua Thien-Hue" pageId="28" pageNumber="179" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Thua Thien-Hue" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF9EC04B1EDF840FAAED4A9" box="[1262,1297,1991,2015]" pageId="28" pageNumber="179" type="male">3♂</specimenCount>
<specimenCount id="9EC0FD4CFFF9EC04B01AF840FA83D4A9" box="[1305,1340,1991,2015]" pageId="28" pageNumber="179" type="female">1♀</specimenCount>
,
<collectorName id="25335313FFF9EC05B048F840FEF8D3D9" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179">A Luoi Nature Reserve</collectorName>
,
<collectingRegion id="4A02F827FFF8EC05B451FF10FDB3D3C6" box="[338,524,151,176]" country="Vietnam" name="Thua Thien-Hue" pageId="29" pageNumber="180">Thua Thien-Hue</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B75FFF10FD62D3C6" box="[604,733,151,176]" pageId="29" pageNumber="180" value="2015-09-18">
<date id="FC781005FFF8EC05B75FFF10FD62D3C6" box="[604,733,151,176]" pageId="29" pageNumber="180" value="2015-09-18">18.IX.2015</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B7EAFF1FFCDED3C6" box="[745,865,151,176]" pageId="29" pageNumber="180">F. Hayashi</collectorName>
leg. (FHC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B6F3FF1FFDD7D3A2" ID-GBIF-Occurrence="1805098070" collectingDate="2015-05-26" collectionCode="PQTC" collectorName="Q. T. Phan" country="Vietnam" location="Dong Giang District" pageId="29" pageNumber="180" specimenCount="3" specimenCount-male="3" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B6F3FF1FFBAAD3C6" box="[1008,1045,152,176]" pageId="29" pageNumber="180" type="male">3♂</specimenCount>
,
<location id="8D19601EFFF8EC05B122FF1FFAB6D3C6" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B122FF1FFAB6D3C6" box="[1057,1289,151,176]" country="Vietnam" name="Dong Giang District" pageId="29" pageNumber="180" stateProvince="Quang Nam">Dong Giang District</location>
,
<collectingRegion id="4A02F827FFF8EC05B017FF1FFA23D3D9" box="[1300,1436,152,176]" country="Vietnam" name="Quang Nam" pageId="29" pageNumber="180">Quang Nam</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B5DCFF3BFEEAD3A3" box="[223,341,188,213]" pageId="29" pageNumber="180" value="2015-05-26">
<date id="FC781005FFF8EC05B5DCFF3BFEEAD3A3" box="[223,341,188,213]" pageId="29" pageNumber="180" value="2015-05-26">26.V.2015</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B462FF3BFE6DD3A3" box="[353,466,188,213]" pageId="29" pageNumber="180">Q.T. Phan</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B711FF3BFDDFD3A2" box="[530,608,188,212]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B777FF3BFA80D3A2" ID-GBIF-Occurrence="1805098077" box="[628,1343,188,213]" collectingDate="2016-06-21" collectorName="T. Kompier" country="Vietnam" location="Quang Nam" pageId="29" pageNumber="180" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B777FF3BFD28D3A2" box="[628,663,188,212]" pageId="29" pageNumber="180" type="male">3♂</specimenCount>
<specimenCount id="9EC0FD4CFFF8EC05B79DFF3BFD7ED3A2" box="[670,705,188,212]" pageId="29" pageNumber="180" type="female">1♀</specimenCount>
,
<collectingRegion id="4A02F827FFF8EC05B7D3FF3BFCE7D3A2" box="[720,856,188,212]" country="Vietnam" name="Quang Nam" pageId="29" pageNumber="180">Quang Nam</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B6AAFF3BFB93D3A3" box="[937,1068,188,213]" pageId="29" pageNumber="180" value="2016-06-21">
<date id="FC781005FFF8EC05B6AAFF3BFB93D3A3" box="[937,1068,188,213]" pageId="29" pageNumber="180" value="2016-06-21">21.VI.2016</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B13BFF3AFB06D3A3" box="[1080,1209,188,213]" pageId="29" pageNumber="180">T. Kompier</collectorName>
leg. (TKC)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B048FF3BFC16D38E" ID-GBIF-Occurrence="1805098084" collectingDate="2017-05-22" collectionCode="PQTC" collectorName="Q. T. Phan" country="Vietnam" location="Ba Na Nature Reserve" pageId="29" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="Da Nang" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B048FF3BFACFD3A2" box="[1355,1392,188,212]" pageId="29" pageNumber="180" type="male">1♂</specimenCount>
,
<location id="8D19601EFFF8EC05B07EFF3AFECFD381" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B07EFF3AFECFD381" country="Vietnam" name="Ba Na Nature Reserve" pageId="29" pageNumber="180" stateProvince="Da Nang">Ba Na Nature Reserve</location>
,
<collectingRegion id="4A02F827FFF8EC05B47FFF67FE5DD38E" box="[380,482,224,248]" country="Vietnam" name="Da Nang" pageId="29" pageNumber="180">Da Nang</collectingRegion>
city,
<collectingDate id="EC3CE9EDFFF8EC05B723FF67FD29D38E" box="[544,662,224,248]" pageId="29" pageNumber="180" value="2017-05-22">
<date id="FC781005FFF8EC05B723FF67FD29D38E" box="[544,662,224,248]" pageId="29" pageNumber="180" value="2017-05-22">22.V.2017</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B7A2FF67FCACD38E" box="[673,787,223,248]" pageId="29" pageNumber="180">Q.T. Phan</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B650FF67FC1ED38E" box="[851,929,224,248]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B6B6FF67FD1AD26A" ID-GBIF-Occurrence="1805098093" collectingDate="2017-05-22" collectionCode="PQTC" collectorName="Q. T. Phan" country="Vietnam" location="Chu Mom Ray National Park" pageId="29" pageNumber="180" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B6B6FF67FC67D38E" box="[949,984,224,248]" pageId="29" pageNumber="180" type="male">4♂</specimenCount>
<specimenCount id="9EC0FD4CFFF8EC05B6E3FF67FBBCD38E" box="[992,1027,224,248]" pageId="29" pageNumber="180" type="female">1♀</specimenCount>
,
<location id="8D19601EFFF8EC05B112FF58FADFD38E" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B112FF58FADFD38E" box="[1041,1376,223,248]" country="Vietnam" name="Chu Mom Ray National Park" pageId="29" pageNumber="180" stateProvince="Kon Tum">Chu Mom Ray National Park</location>
,
<collectingRegion id="4A02F827FFF8EC05B068FF67FF74D26A" country="Vietnam" name="Kon Tum" pageId="29" pageNumber="180">Kon Tum</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B41FFE83FE2ED26A" box="[284,401,260,284]" pageId="29" pageNumber="180" value="2017-05-22">
<date id="FC781005FFF8EC05B41FFE83FE2ED26A" box="[284,401,260,284]" pageId="29" pageNumber="180" value="2017-05-22">22.V.2017</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B49EFE83FDB1D26B" box="[413,526,260,285]" pageId="29" pageNumber="180">Q.T. Phan</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B74CFE83FD22D26A" box="[591,669,260,284]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B7B2FE83FED9D236" ID-GBIF-Occurrence="1805098097" collectingDate="2017-04-16" collectionCode="PQTC" collectorName="Q. T. Phan" country="Vietnam" location="Hon Ba Nature Reserve" pageId="29" pageNumber="180" specimenCount="10" specimenCount-male="10" stateProvince="Khanh Hoa" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B7B2FE83FD5BD26A" box="[689,740,260,284]" pageId="29" pageNumber="180" type="male">10♂</specimenCount>
,
<location id="8D19601EFFF8EC05B7F2FE82FBBDD26A" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B7F2FE82FBBDD26A" box="[753,1026,261,284]" country="Vietnam" name="Hon Ba Nature Reserve" pageId="29" pageNumber="180" stateProvince="Khanh Hoa">Hon Ba Nature Reserve</location>
,
<collectingRegion id="4A02F827FFF8EC05B10DFE83FB30D26A" box="[1038,1167,260,285]" country="Vietnam" name="Khanh Hoa" pageId="29" pageNumber="180">Khanh Hoa</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B1E3FE83FAE0D26B" box="[1248,1375,260,285]" pageId="29" pageNumber="180" value="2017-04-16">
<date id="FC781005FFF8EC05B1E3FE83FAE0D26B" box="[1248,1375,260,285]" pageId="29" pageNumber="180" value="2017-04-16">16.IV.2017</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B068FE83FF70D236" pageId="29" pageNumber="180">Q.T. Phan</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B412FEAFFEE0D236" box="[273,351,296,320]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B470FEAFFAD9D236" ID-GBIF-Occurrence="1805098104" box="[371,1382,295,320]" collectingDate="2017-06-28" collectionCode="PQTC" collectorName="Q. T. Phan" country="Vietnam" location="Bach Ma National Park" pageId="29" pageNumber="180" specimenCount="4" specimenCount-male="4" stateProvince="Thua Thien-Hue" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B470FEAFFE26D236" box="[371,409,296,320]" pageId="29" pageNumber="180" type="male">4♂</specimenCount>
,
<location id="8D19601EFFF8EC05B4A6FEA0FD0BD236" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B4A6FEA0FD0BD236" box="[421,692,295,320]" country="Vietnam" name="Bach Ma National Park" pageId="29" pageNumber="180" stateProvince="Thua Thien-Hue">Bach Ma National Park</location>
,
<collectingRegion id="4A02F827FFF8EC05B7C3FEA0FCC4D236" box="[704,891,295,320]" country="Vietnam" name="Thua Thien-Hue" pageId="29" pageNumber="180">Thua Thien-Hue</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B6CEFEAFFBEFD236" box="[973,1104,296,320]" pageId="29" pageNumber="180" value="2017-06-28">
<date id="FC781005FFF8EC05B6CEFEAFFBEFD236" box="[973,1104,296,320]" pageId="29" pageNumber="180" value="2017-06-28">28.VI.2017</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B15FFEAFFB71D236" box="[1116,1230,295,320]" pageId="29" pageNumber="180">Q.T. Phan</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B013FEAFFAE1D236" box="[1296,1374,296,320]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38AE3C98FFF8EC05B071FEAFFB3AD212" ID-GBIF-Occurrence="1805098110" collectingDate="2017-04-30" collectionCode="PQTC" collectorName="Hoang Quang Duy" country="Vietnam" location="Chu Yang Sin National Park" pageId="29" pageNumber="180" specimenCount="8" specimenCount-male="8" stateProvince="Dak Lak" typeStatus="paratype">
<specimenCount id="9EC0FD4CFFF8EC05B071FEAFFA28D236" box="[1394,1431,296,320]" pageId="29" pageNumber="180" type="male">8♂</specimenCount>
,
<location id="8D19601EFFF8EC05B594FECBFE68D213" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B594FECBFE68D213" box="[151,471,332,357]" country="Vietnam" name="Chu Yang Sin National Park" pageId="29" pageNumber="180" stateProvince="Dak Lak">Chu Yang Sin National Park</location>
,
<collectingRegion id="4A02F827FFF8EC05B4E1FECBFDFBD213" box="[482,580,332,357]" country="Vietnam" name="Dak Lak" pageId="29" pageNumber="180">Dak Lak</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B790FECBFCADD212" box="[659,786,332,356]" pageId="29" pageNumber="180" value="2017-04-30">
<date id="FC781005FFF8EC05B790FECBFCADD212" box="[659,786,332,356]" pageId="29" pageNumber="180" value="2017-04-30">30.IV.2017</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B61EFECAFC4FD213" box="[797,1008,332,357]" pageId="29" pageNumber="180">Hoang Quang Duy</collectorName>
leg. (
<collectionCode id="EED7AE00FFF8EC05B133FECBFBC1D212" box="[1072,1150,332,356]" pageId="29" pageNumber="180">PQTC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DCB9664DFFF8EC05B594FA5DFCC3D55B" httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="29" pageNumber="180" targetBox="[321,1248,424,1466]" targetPageId="29">
<paragraph id="887936C5FFF8EC05B594FA5DFCC3D55B" blockId="29.[151,1435,1498,1581]" pageId="29" pageNumber="180">
<emphasis id="BAB2EAD7FFF8EC05B594FA5DFE9ED699" bold="true" box="[151,289,1498,1520]" pageId="29" pageNumber="180">FIGURE 20.</emphasis>
<taxonomicName id="4FC64D46FFF8EC05B424FA5DFE3ED686" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[295,385,1498,1520]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="BAB2EAD7FFF8EC05B424FA5DFE3ED686" box="[295,385,1498,1520]" italics="true" pageId="29" pageNumber="180">Euphaea</emphasis>
</taxonomicName>
spp. [20ad]
<taxonomicName id="4FC64D46FFF8EC05B713FA5DFD17D686" authority="Phan &amp; Kompier &amp; Karube &amp; Hayashi, 2018" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[528,680,1498,1520]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="saola" status="sp. nov.">
<emphasis id="BAB2EAD7FFF8EC05B713FA5DFD17D686" box="[528,680,1498,1520]" italics="true" pageId="29" pageNumber="180">Euphaea saola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A18157ACFFF8EC05B7ACFA5DFCBFD699" box="[687,768,1498,1519]" pageId="29" pageNumber="180" rank="species">
<emphasis id="BAB2EAD7FFF8EC05B7ACFA5DFCBFD699" bold="true" box="[687,768,1498,1519]" pageId="29" pageNumber="180">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype ♂ and paratype ♀ and [20e]
<taxonomicName id="4FC64D46FFF8EC05B195FA5CFADFD686" authority=", Hon Ba." authorityName="Hon Ba." box="[1174,1376,1499,1520]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFF8EC05B195FA5CFABDD686" box="[1174,1282,1499,1520]" italics="true" pageId="29" pageNumber="180">E. guerini,</emphasis>
Hon Ba.
</taxonomicName>
(20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view. Scale bar is 10 mm in 20a.
</paragraph>
</caption>
<subSubSection id="C0DC654EFFF8EC05B5C4F9D3FDABD5E7" pageId="29" pageNumber="180" type="materials_examined">
<paragraph id="887936C5FFF8EC05B5C4F9D3FDABD5E7" blockId="29.[151,1437,1620,2006]" pageId="29" pageNumber="180">
<materialsCitation id="38AE3C98FFF8EC05B5C4F9D3FDABD5E7" ID-GBIF-Occurrence="1805098118" collectingDate="2016-08-05" collectingDateMax="2016-08-08" collectingDateMin="2016-08-05" collectorName="T. Kompier." country="Vietnam" county="Other" location="Bach Ma" municipality="Some" pageId="29" pageNumber="180" specimenCount="1" stateProvince="Thua Thien-Hue">
<emphasis id="BAB2EAD7FFF8EC05B5C4F9D3FCABD51B" bold="true" box="[199,788,1620,1645]" pageId="29" pageNumber="180">
<collectingCounty id="61184E49FFF8EC05B5C4F9D3FEB0D51B" box="[199,271,1620,1645]" pageId="29" pageNumber="180">Other</collectingCounty>
materials confirmed by field observations.
</emphasis>
[
<collectingCountry id="F0D17655FFF8EC05B621F9D3FC38D51B" box="[802,903,1620,1645]" name="Vietnam" pageId="29" pageNumber="180">Vietnam</collectingCountry>
]
<collectingMunicipality id="681DACBFFFF8EC05B695F9D2FC69D51B" box="[918,982,1621,1645]" pageId="29" pageNumber="180">Some</collectingMunicipality>
individuals,
<location id="8D19601EFFF8EC05B169F9D3FB6BD51B" LSID="urn:lsid:plazi:treatment:006F87D3FFF9EC07B594F9A5FC43D236:8D19601EFFF8EC05B169F9D3FB6BD51B" box="[1130,1236,1620,1645]" country="Vietnam" county="Other" municipality="Some" name="Bach Ma" pageId="29" pageNumber="180" stateProvince="Thua Thien-Hue">Bach Ma</location>
,
<collectingRegion id="4A02F827FFF8EC05B1E2F9D3FA23D51B" box="[1249,1436,1620,1645]" country="Vietnam" name="Thua Thien-Hue" pageId="29" pageNumber="180">Thua Thien-Hue</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFF8EC05B5DCF9FFFE3DD5E7" box="[223,386,1656,1681]" pageId="29" pageNumber="180" value="2016-08-05" valueMax="2016-08-08" valueMin="2016-08-05">
<date id="FC781005FFF8EC05B5DCF9FFFE3DD5E7" box="[223,386,1656,1681]" pageId="29" pageNumber="180" value="2016-08-05" valueMax="2016-08-08" valueMin="2016-08-05">58.VIII.2016</date>
</collectingDate>
,
<collectorName id="25335313FFF8EC05B48EF9FEFDABD5E7" box="[397,532,1656,1681]" pageId="29" pageNumber="180">T. Kompier.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFF8EC05B5C4F91AFEBED41F" pageId="29" pageNumber="180" type="etymology">
<paragraph id="887936C5FFF8EC05B5C4F91AFEBED41F" blockId="29.[151,1437,1620,2006]" pageId="29" pageNumber="180">
<emphasis id="BAB2EAD7FFF8EC05B5C4F91AFEF1D5C0" bold="true" box="[199,334,1693,1718]" pageId="29" pageNumber="180">Etymology.</emphasis>
The specific name “
<taxonomicName id="4FC64D46FFF8EC05B753F91AFD33D5C0" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[592,652,1693,1718]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFF8EC05B753F91AFD33D5C0" box="[592,652,1693,1718]" italics="true" pageId="29" pageNumber="180">saola</emphasis>
</taxonomicName>
” of this species is derived from the name of the bovid “Saola” (
<taxonomicName id="4FC64D46FFF8EC05B5A3F947FBE0D5AF" authority="Dung, Giao, Chinh, Tuoc, Arctander &amp; MacKinnon, 1993" authorityName="Dung, Giao, Chinh, Tuoc, Arctander &amp; MacKinnon" authorityYear="1993" box="[160,1119,1728,1753]" class="Mammalia" family="Bovidae" genus="Pseudoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="29" pageNumber="180" phylum="Chordata" rank="species" species="nghetinhensis">
<emphasis id="BAB2EAD7FFF8EC05B5A3F947FE06D5AF" box="[160,441,1728,1753]" italics="true" pageId="29" pageNumber="180">Pseudoryx nghetinhensis</emphasis>
Dung, Giao, Chinh, Tuoc, Arctander &amp; MacKinnon, 1993
</taxonomicName>
). This is one of the worlds rarest large mammals, a forest-dwelling bovine found only in the Truong Son mountain range of
<collectingCountry id="F0D17655FFF8EC05B1C8F962FA95D588" box="[1227,1322,1765,1790]" name="Vietnam" pageId="29" pageNumber="180">Vietnam</collectingCountry>
and
<collectingCountry id="F0D17655FFF8EC05B05CF961FA26D58B" box="[1375,1433,1766,1789]" name="Laos" pageId="29" pageNumber="180">Laos</collectingCountry>
. The
<typeStatus id="577D8867FFF8EC05B5C8F88EFF44D457" box="[203,251,1801,1825]" pageId="29" pageNumber="180">type</typeStatus>
locality of
<taxonomicName id="4FC64D46FFF8EC05B483F88FFD96D457" authority="Phan &amp; Kompier &amp; Karube &amp; Hayashi, 2018" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[384,553,1800,1825]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="saola" status="sp. nov.">
<emphasis id="BAB2EAD7FFF8EC05B483F88FFD96D457" box="[384,553,1800,1825]" italics="true" pageId="29" pageNumber="180">Euphaea saola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A18157ACFFF8EC05B731F88FFD32D457" box="[562,653,1800,1825]" pageId="29" pageNumber="180" rank="species">
<emphasis id="BAB2EAD7FFF8EC05B731F88FFD32D457" bold="true" box="[562,653,1800,1825]" pageId="29" pageNumber="180">sp. nov.</emphasis>
</taxonomicNameLabel>
is A Luoi Nature Reserve which is a part of the Saola Conservation Area, central
<collectingCountry id="F0D17655FFF8EC05B42CF8AAFE2DD430" box="[303,402,1837,1862]" name="Vietnam" pageId="29" pageNumber="180">Vietnam</collectingCountry>
. The general distribution of the new species largely coincides with that of this extraordinary mammal.
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFF8EC06B5C4F8F2FC1FD72E" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180" type="description">
<paragraph id="887936C5FFF8EC05B5C4F8F2FA3CD4C7" blockId="29.[151,1437,1620,2006]" pageId="29" pageNumber="180">
<emphasis id="BAB2EAD7FFF8EC05B5C4F8F2FE52D4F8" bold="true" box="[199,493,1909,1934]" pageId="29" pageNumber="180">
Description of
<typeStatus id="577D8867FFF8EC05B47CF8F2FE57D4F8" box="[383,488,1909,1934]" pageId="29" pageNumber="180" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="BAB2EAD7FFF8EC05B4FBF8F2FC15D4F8" box="[504,938,1909,1934]" italics="true" pageId="29" pageNumber="180">Head, prothorax, synthorax, and legs</emphasis>
(
<figureCitation id="10FD2A40FFF8EC05B6BDF8F2FB9DD4FB" box="[958,1058,1909,1934]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="29" pageNumber="180">Fig. 20a</figureCitation>
). Entirely black excluding some narrow yellow stripes along the border of mesepimeron and metepisterum and on lower margin of metepimeron.
</paragraph>
<paragraph id="887936C5FFF8EC06B5C4F839FD17D26B" blockId="29.[151,1437,1620,2006]" lastBlockId="30.[151,1437,151,2013]" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180">
<emphasis id="BAB2EAD7FFF8EC05B5C4F839FEB5D4A3" box="[199,266,1982,2005]" italics="true" pageId="29" pageNumber="180">Wings</emphasis>
(
<figureCitation id="10FD2A40FFF8EC05B419F83AFEC3D4A0" box="[282,380,1981,2006]" captionStart="FIGURE 14" captionStartId="23.[151,250,1889,1911]" captionTargetBox="[220,1341,852,1868]" captionTargetId="figure@23.[184,1403,844,1868]" captionTargetPageId="23" captionText="FIGURE 14. Dorsal view of wings of male Euphaea spp. (14a) E. hirta; (14b) E. masoni; (14c) E. guerini; (14d) E. saola sp. nov." httpUri="https://zenodo.org/record/3096097/files/figure.png" pageId="29" pageNumber="180">Fig. 14d</figureCitation>
). Similar to those of
<taxonomicName id="4FC64D46FFF8EC05B766F839FD6AD4A3" box="[613,725,1982,2005]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFF8EC05B766F839FD6AD4A3" box="[613,725,1982,2005]" italics="true" pageId="29" pageNumber="180">E. guerini</emphasis>
</taxonomicName>
with most of FW area darkened, excluding the basal part of the wing (but covering all cells along the subcosta), and the apical one-sixth, although the apical margin of the FW is also darkened. HW are black with metallic blue covering a large part of their underside, and with metallic green on the upperside, as in
<taxonomicName id="4FC64D46FFFBEC06B480FF66FE49D38E" box="[387,502,225,248]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B480FF66FE49D38E" box="[387,502,225,248]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
. FW has 2829 Ax and 3233 Px, and HW have 2223 Ax and 3537 Px. The cubital space has 23 crossveins in both wings.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FEA0FD06D2FE" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FEA0FE8ED236" box="[199,305,295,320]" italics="true" pageId="30" pageNumber="181">Abdomen</emphasis>
. Entirely black. Lateroventral margin of S3 and S78 with fine setae in addition to the tuft of long setae on the base of S9 like
<taxonomicName id="4FC64D46FFFBEC06B4C8FECAFD84D212" box="[459,571,333,356]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B4C8FECAFD84D212" box="[459,571,333,356]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B749FECBFD16D213" box="[586,681,332,357]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="30" pageNumber="181">Fig. 15a</figureCitation>
). Dorsum of S10 raised into large carinal spine posteriorly and with a small hump near its anterior margin (
<figureCitation id="10FD2A40FFFBEC06B749FEE8FD13D2FE" box="[586,684,367,392]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20b</figureCitation>
).
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FE13FB7CD2DA" blockId="30.[151,1437,151,2013]" box="[199,1219,404,429]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FE13FEDCD2DB" box="[199,355,404,429]" italics="true" pageId="30" pageNumber="181">Genital ligula</emphasis>
. With two thick flagella, perpendicular to terminal segment as in
<figureCitation id="10FD2A40FFFBEC06B141FE13FB00D2DA" box="[1090,1215,404,429]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Figure 20c</figureCitation>
.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FE30FED6D283" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FE30FE35D2A6" box="[199,394,439,464]" italics="true" pageId="30" pageNumber="181">Anal appendages</emphasis>
. Typical of the genus with cerci short and robust, paraprocts are very short, slightly curved upward (
<figureCitation id="10FD2A40FFFBEC06B5F9FE5BFEE3D283" box="[250,348,476,501]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20b</figureCitation>
).
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FD87FC1FD16E" blockId="30.[151,1437,151,2013]" box="[199,928,511,536]" pageId="30" pageNumber="181">Measurements (in mm). HW 29; abdomen (incl. appendages) 37.</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FDA3FC22D116" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FDA3FD9FD14B" bold="true" box="[199,544,548,573]" pageId="30" pageNumber="181">
Variation in
<typeStatus id="577D8867FFFBEC06B463FDA3FE74D14B" box="[352,459,548,573]" pageId="30" pageNumber="181" type="paratype">paratype</typeStatus>
males.
</emphasis>
Four
<typeStatus id="577D8867FFFBEC06B765FDA2FD6ED14B" box="[614,721,549,573]" pageId="30" pageNumber="181" type="paratype">paratypes</typeStatus>
show little morphological variation, excluding size;
<quantity id="4F3E9B20FFFBEC06B02FFDA3FF7CD129" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="2.95" metricValueMin="2.75" pageId="30" pageNumber="181" unit="mm" value="28.5" valueMax="29.5" valueMin="27.5">27.529.5 mm</quantity>
in HW length and
<quantity id="4F3E9B20FFFBEC06B49BFDC0FDAED129" box="[408,529,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="30" pageNumber="181" unit="mm" value="37.0" valueMax="38.0" valueMin="36.0">3638 mm</quantity>
in abdomen including appendages.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FDEBFC3BD1BA" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FDEBFD88D1F3" bold="true" box="[199,567,620,645]" pageId="30" pageNumber="181">
Description of
<typeStatus id="577D8867FFFBEC06B475FDEBFE5DD1F3" box="[374,482,620,645]" pageId="30" pageNumber="181" type="paratype">paratype</typeStatus>
female
</emphasis>
.
<emphasis id="BAB2EAD7FFFBEC06B746FDEBFD3DD1F3" box="[581,642,620,645]" italics="true" pageId="30" pageNumber="181">Head</emphasis>
(
<figureCitation id="10FD2A40FFFBEC06B792FDEBFD4DD1F3" box="[657,754,620,645]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20d</figureCitation>
). Postclypeus, frons, antennae, and the dorsum of head matte black. Anteclypeus shining black. Labrum and base of mandibles dark yellow but margin and mid-dorsal part black. Genae dark yellow and this extends to the level of antennae.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FD50FDF8D0F6" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FD50FEA7D186" box="[199,280,727,752]" italics="true" pageId="30" pageNumber="181">Thorax</emphasis>
(
<figureCitation id="10FD2A40FFFBEC06B424FD50FE38D186" box="[295,391,727,752]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16e</figureCitation>
). Anterior lobe of prothorax entirely black; middle and posterior lobes black with tiny orange spots; middle lobe raised into two large separately humps; distal margin of posterior lobe rounded, simple structurally (
<figureCitation id="10FD2A40FFFBEC06B424FC98FE34D04E" box="[295,395,799,824]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">
Fig.
<quantity id="4F3E9B20FFFBEC06B45EFCA7FE34D04E" box="[349,395,800,824]" metricMagnitude="-2" metricUnit="kg" metricValue="1.8" pageId="30" pageNumber="181" unit="g" value="18.0">18g</quantity>
</figureCitation>
). Synthorax black with narrow orange stripes as follows: mesepisternum has two separated narrow stripes on either side; mesepimeron entirely black; metepisternum and metepimeron both with two yellow stripes along distal and lower margins.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FC0AFEECD0D3" blockId="30.[151,1437,151,2013]" box="[199,339,908,933]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FC0AFF43D0D2" box="[199,252,909,932]" italics="true" pageId="30" pageNumber="181">Legs</emphasis>
. Black.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FC36FC61D09B" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FC36FEB5D0BE" box="[199,266,945,968]" italics="true" pageId="30" pageNumber="181">Wings</emphasis>
. Hyaline tinted amber on proximal half of FW and complete HW. FW with 2829 Ax and 3132 Px, HW with 21 Ax and 29 Px. Cubital space with 13 crossveins in all wings.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FC70FBD8D743" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FC70FE8ED766" box="[199,305,1015,1040]" italics="true" pageId="30" pageNumber="181">Abdomen</emphasis>
(
<figureCitation id="10FD2A40FFFBEC06B441FC70FE1AD766" box="[322,421,1015,1040]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16e</figureCitation>
). Black with lateral yellow markings as follows: S1 with a large spot, S23 with a very narrow stripe, S45 with only tiny oval posterior spot, and S610 and ovipositor black.
</paragraph>
<paragraph id="887936C5FFFBEC06B5C4FBC7FC1FD72E" blockId="30.[151,1437,151,2013]" box="[199,928,1087,1112]" pageId="30" pageNumber="181">Measurements (in mm). HW 29; abdomen (incl. appendages) 33.</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFFBEC07B5C4FBE3FDA9D3A2" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181" type="discussion">
<paragraph id="887936C5FFFBEC06B5C4FBE3FAFCD596" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="181">
<emphasis id="BAB2EAD7FFFBEC06B5C4FBE3FE84D70B" bold="true" box="[199,315,1124,1149]" pageId="30" pageNumber="181">Remarks.</emphasis>
<taxonomicName id="4FC64D46FFFBEC06B442FBE3FE57D70B" authority="Phan &amp; Kompier &amp; Karube &amp; Hayashi, 2018" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[321,488,1124,1149]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola" status="sp. nov.">
<emphasis id="BAB2EAD7FFFBEC06B442FBE3FE57D70B" box="[321,488,1124,1149]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A18157ACFFFBEC06B4ECFBE3FDF7D70B" box="[495,584,1124,1149]" pageId="30" pageNumber="181" rank="species">
<emphasis id="BAB2EAD7FFFBEC06B4ECFBE3FDF7D70B" bold="true" box="[495,584,1124,1149]" pageId="30" pageNumber="181">sp. nov.</emphasis>
</taxonomicNameLabel>
is very similar to the population of
<taxonomicName id="4FC64D46FFFBEC06B6D5FBE2FBFAD70A" box="[982,1093,1125,1148]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B6D5FBE2FBFAD70A" box="[982,1093,1125,1148]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
in
<collectingCountry id="F0D17655FFFBEC06B16BFBE3FB78D70B" box="[1128,1223,1124,1149]" name="Vietnam" pageId="30" pageNumber="181">Vietnam</collectingCountry>
with similar wingcolor pattern of the male, but can be distinguished by more distinct dark apical margin and more distinct hyaline base in FW (
<figureCitation id="10FD2A40FFFBEC06B431FB2BFE29D7B3" box="[306,406,1196,1221]" captionStart="FIGURE 14" captionStartId="23.[151,250,1889,1911]" captionTargetBox="[220,1341,852,1868]" captionTargetId="figure@23.[184,1403,844,1868]" captionTargetPageId="23" captionText="FIGURE 14. Dorsal view of wings of male Euphaea spp. (14a) E. hirta; (14b) E. masoni; (14c) E. guerini; (14d) E. saola sp. nov." httpUri="https://zenodo.org/record/3096097/files/figure.png" pageId="30" pageNumber="181">Fig. 14d</figureCitation>
) compared to
<taxonomicName id="4FC64D46FFFBEC06B747FB2AFD07D7B2" box="[580,696,1197,1220]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B747FB2AFD07D7B2" box="[580,696,1197,1220]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B7CFFB2BFC8ED7B2" box="[716,817,1196,1221]" captionStart="FIGURE 14" captionStartId="23.[151,250,1889,1911]" captionTargetBox="[220,1341,852,1868]" captionTargetId="figure@23.[184,1403,844,1868]" captionTargetPageId="23" captionText="FIGURE 14. Dorsal view of wings of male Euphaea spp. (14a) E. hirta; (14b) E. masoni; (14c) E. guerini; (14d) E. saola sp. nov." httpUri="https://zenodo.org/record/3096097/files/figure.png" pageId="30" pageNumber="181">Fig. 14c</figureCitation>
). Abdominal segments S3 and S8 bear long setae in
<taxonomicName id="4FC64D46FFFBEC06B594FB48FEFED79E" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[151,321,1231,1256]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B594FB48FEFED79E" box="[151,321,1231,1256]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B452FB48FE0CD79E" box="[337,435,1231,1256]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="30" pageNumber="181">Fig. 15a</figureCitation>
), but very short setae in
<taxonomicName id="4FC64D46FFFBEC06B7D7FB56FCF9D79E" box="[724,838,1233,1256]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B7D7FB56FCF9D79E" box="[724,838,1233,1256]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B654FB48FC05D79E" box="[855,954,1231,1256]" captionStart="FIGURE15" captionStartId="24.[151,250,929,951]" captionTargetBox="[186,1410,221,879]" captionTargetId="figure@24.[153,1433,193,907]" captionTargetPageId="24" captionText="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." httpUri="https://zenodo.org/record/3096107/files/figure.png" pageId="30" pageNumber="181">Fig. 15b</figureCitation>
). In
<taxonomicName id="4FC64D46FFFBEC06B6F1FB48FB24D79E" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[1010,1179,1231,1256]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B6F1FB48FB24D79E" box="[1010,1179,1231,1256]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
, the dorsum of S10 is raised in two places: there is a small hump in lateral view on the anterior part, but posteriorly it rises somewhat obliquely upward (
<figureCitation id="10FD2A40FFFBEC06B46FFA90FE74D646" box="[364,459,1303,1328]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20b</figureCitation>
) forming a conspicuous carinal spine. S10 of
<taxonomicName id="4FC64D46FFFBEC06B6D3FA90FB33D646" box="[976,1164,1303,1328]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B6D3FA90FB33D646" box="[976,1164,1303,1328]" italics="true" pageId="30" pageNumber="181">Euphaea guerini</emphasis>
</taxonomicName>
lacks the anterior hump and the large posterior carinal spine rises strongly upwards, almost perpendicular to the abdomen (
<figureCitation id="10FD2A40FFFBEC06B1E8FABBFAF3D622" box="[1259,1356,1340,1365]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20e</figureCitation>
). Cerci are short and robust, the posterior dorsal spine of S10 extending beyond their base posteriorly in lateral view in
<taxonomicName id="4FC64D46FFFBEC06B594FA03FEFED6EB" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[151,321,1412,1437]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B594FA03FEFED6EB" box="[151,321,1412,1437]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B451FA03FE09D6EB" box="[338,438,1412,1437]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20b</figureCitation>
), but cerci of
<taxonomicName id="4FC64D46FFFBEC06B759FA02FD73D6EA" box="[602,716,1413,1436]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B759FA02FD73D6EA" box="[602,716,1413,1436]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
are narrower, petiolated at base, and the dorsal spine does not extend posteriorly of their base, so they are visible in dorsal view (
<figureCitation id="10FD2A40FFFBEC06B6AEFA20FBAED6B6" box="[941,1041,1447,1472]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Fig. 20e</figureCitation>
). Vesicle of
<taxonomicName id="4FC64D46FFFBEC06B1A6FA20FAEFD6B6" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[1189,1360,1447,1472]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B1A6FA20FAEFD6B6" box="[1189,1360,1447,1472]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
heartshaped, pointed at its posterior margin (
<figureCitation id="10FD2A40FFFBEC06B753FA4BFD76D693" box="[592,713,1484,1509]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">Fig. 18a, b</figureCitation>
), but it is ax-shaped, with posterior margin rounded in
<taxonomicName id="4FC64D46FFFBEC06B02FFA4AFA23D692" box="[1324,1436,1485,1508]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B02FFA4AFA23D692" box="[1324,1436,1485,1508]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B59CFA68FF40D57E" box="[159,255,1519,1544]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">Fig. 18c</figureCitation>
). The shape of the posterior lobe of the prothorax in females of
<taxonomicName id="4FC64D46FFFBEC06B6C4FA68FBD1D57E" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[967,1134,1519,1544]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B6C4FA68FBD1D57E" box="[967,1134,1519,1544]" italics="true" pageId="30" pageNumber="181">Euphaea saola</emphasis>
</taxonomicName>
differs markedly from that of
<taxonomicName id="4FC64D46FFFBEC06B5B6F992FE9AD55A" box="[181,293,1557,1580]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B5B6F992FE9AD55A" box="[181,293,1557,1580]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
by the smoothly rounded lateral edges and minor indentation of the posterior margin (
<figureCitation id="10FD2A40FFFBEC06B1EAF993FAF6D55A" box="[1257,1353,1556,1581]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">
Fig.
<quantity id="4F3E9B20FFFBEC06B01EF993FAF6D55A" box="[1309,1353,1556,1580]" metricMagnitude="-2" metricUnit="kg" metricValue="1.8" pageId="30" pageNumber="181" unit="g" value="18.0">18g</quantity>
</figureCitation>
), while the lateral edges curve out and backwards in
<taxonomicName id="4FC64D46FFFBEC06B7A7F9BEFCA8D526" box="[676,791,1593,1616]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B7A7F9BEFD0AD526" box="[676,693,1593,1616]" italics="true" pageId="30" pageNumber="181">E</emphasis>
.
<emphasis id="BAB2EAD7FFFBEC06B7C5F9BEFCA8D526" box="[710,791,1593,1616]" italics="true" pageId="30" pageNumber="181">guerini</emphasis>
</taxonomicName>
, and the posterior margin is completely round centrally (
<figureCitation id="10FD2A40FFFBEC06B59CF9DBFEBDD503" box="[159,258,1628,1653]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">Fig. 18h</figureCitation>
). In addition, (i) thorax black with very narrow stripes in
<taxonomicName id="4FC64D46FFFBEC06B68EF9DAFC56D503" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[909,1001,1628,1653]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B68EF9DAFC56D503" box="[909,1001,1628,1653]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B6FAF9DBFBE5D503" box="[1017,1114,1628,1653]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16e</figureCitation>
), but stripes relatively broad in
<taxonomicName id="4FC64D46FFFBEC06B5B6F906FE99D5EE" box="[181,294,1665,1688]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B5B6F906FE99D5EE" box="[181,294,1665,1688]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B434F9F8FE25D5EE" box="[311,410,1663,1688]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16d</figureCitation>
); (ii) abdomen black with narrow yellow stripes on S13, tiny spots on S45, and entirely black on other segments in
<taxonomicName id="4FC64D46FFFBEC06B4CBF922FD9BD5CB" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[456,548,1700,1725]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B4CBF922FD9BD5CB" box="[456,548,1700,1725]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B730F923FD2DD5CB" box="[563,658,1700,1725]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16e</figureCitation>
), but with stripes on S16 and a small yellow spot on S
<quantity id="4F3E9B20FFFBEC06B1F8F923FA99D5CB" box="[1275,1318,1700,1725]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="30" pageNumber="181" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="4FC64D46FFFBEC06B02FF922FA23D5CA" box="[1324,1436,1701,1724]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B02FF922FA23D5CA" box="[1324,1436,1701,1724]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B59CF940FEBED596" box="[159,257,1735,1760]" captionStart="FIGURE 16" captionStartId="24.[151,250,1970,1992]" captionTargetBox="[286,1291,1018,1921]" captionTargetId="figure@24.[281,1305,1015,1949]" captionTargetPageId="24" captionText="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." httpUri="https://zenodo.org/record/3096109/files/figure.png" pageId="30" pageNumber="181">Fig. 16d</figureCitation>
); and (iii) wings entirely dark amber in
<taxonomicName id="4FC64D46FFFBEC06B7BCF94EFCA5D596" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[703,794,1735,1760]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B7BCF94EFCA5D596" box="[703,794,1735,1760]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
, but hyaline with pale yellow color in
<taxonomicName id="4FC64D46FFFBEC06B1CFF94EFA83D596" box="[1228,1340,1737,1760]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B1CFF94EFA83D596" box="[1228,1340,1737,1760]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="887936C5FFFBEC07B5C4F96AFDA9D3A2" blockId="30.[151,1437,151,2013]" lastBlockId="31.[151,1436,151,320]" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181">
It is difficult to establish what published records/info of
<taxonomicName id="4FC64D46FFFBEC06B670F96BFB8AD472" box="[883,1077,1772,1797]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B670F96BFB8AD472" box="[883,1077,1772,1797]" italics="true" pageId="30" pageNumber="181">Euphaea guerini</emphasis>
</taxonomicName>
refer to
<taxonomicName id="4FC64D46FFFBEC06B1A5F96AFAB7D473" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[1190,1288,1772,1797]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B1A5F96AFAB7D473" box="[1190,1288,1772,1797]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
. Van
<bibRefCitation id="EC574B34FFFBEC06B057F96BFEDED45E" author="Tol" pageId="30" pageNumber="181" refString="van Tol, J. &amp; Rozendaal, F. G. (1995) Records of Calopterygoidea from Vietnam, with descriptions of two new species (Zygoptera: Amphipterygidae, Calopterygidae, Chlorocyphidae, Euphaeidae). Odonatologica, 24, 89 - 107." type="journal article" year="1995">Tol &amp; Rozendaal (1995)</bibRefCitation>
provided figures of the last abdominal segments and vesicle of
<taxonomicName id="4FC64D46FFFBEC06B14BF888FAB9D45E" box="[1096,1286,1807,1832]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B14BF888FAB9D45E" box="[1096,1286,1807,1832]" italics="true" pageId="30" pageNumber="181">Euphaea guerini</emphasis>
</taxonomicName>
from central
<collectingCountry id="F0D17655FFFBEC06B594F8B3FF46D43B" box="[151,249,1844,1869]" name="Vietnam" pageId="30" pageNumber="181">Vietnam</collectingCountry>
, but they did not specify the locality of each specimen.
<figureCitation id="10FD2A40FFFBEC06B68EF8B3FC43D43A" box="[909,1020,1844,1869]" captionStart="FIGURE 20" captionStartId="29.[151,250,1498,1520]" captionTargetBox="[321,1248,424,1466]" captionTargetId="figure@29.[317,1269,409,1477]" captionTargetPageId="29" captionText="FIGURE 20. Euphaea spp. [20ad] Euphaea saola sp. nov., holotype ♂ and paratype ♀ and [20e] E. guerini, Hon Ba. (20a) habitus male; (20b) S910 and appendages, lateral view; (20c) genital ligula, dorsal view; (20d) head female, antero-dorsal view; (19e) S10 and appendages, lateral view.Scale bar is 10 mm in 20a." httpUri="https://zenodo.org/record/3096129/files/figure.png" pageId="30" pageNumber="181">Figure 20</figureCitation>
of
<taxonomicName id="4FC64D46FFFBEC06B132F8B3FB50D43A" box="[1073,1263,1844,1869]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B132F8B3FB50D43A" box="[1073,1263,1844,1869]" italics="true" pageId="30" pageNumber="181">Euphaea guerini</emphasis>
</taxonomicName>
in van
<bibRefCitation id="EC574B34FFFBEC06B055F8B3FEE1D406" author="Tol" pageId="30" pageNumber="181" refString="van Tol, J. &amp; Rozendaal, F. G. (1995) Records of Calopterygoidea from Vietnam, with descriptions of two new species (Zygoptera: Amphipterygidae, Calopterygidae, Chlorocyphidae, Euphaeidae). Odonatologica, 24, 89 - 107." type="journal article" year="1995">Tol &amp; Rozendaal (1995)</bibRefCitation>
matches
<taxonomicName id="4FC64D46FFFBEC06B4CBF8DEFD9CD406" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[456,547,1879,1904]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B4CBF8DEFD9CD406" box="[456,547,1879,1904]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
based on the resemblance of the dorsal prominence on S10, but their
<figureCitation id="10FD2A40FFFBEC06B02DF8D0FAC1D406" box="[1326,1406,1879,1904]" captionStart="FIGURE21" captionStartId="34.[151,250,1855,1877]" captionTargetBox="[287,1305,1025,1832]" captionTargetId="figure@34.[281,1305,1023,1833]" captionTargetPageId="34" captionText="FIGURE21. Distribution maps of 3 Anisopleura and 7 Bayaderaspecies in Vietnam.Closed circles represent the provinces in which the distribution is confirmed. Lines drawn in A. bipugio show the northern, central, and southern areas of Vietnam separated by the gaps of mainmountainous regions." httpUri="https://zenodo.org/record/3096131/files/figure.png" pageId="30" pageNumber="181">Fig. 21</figureCitation>
of vesicle is similar to that of
<taxonomicName id="4FC64D46FFFBEC06B4C1F8FAFD8DD4E2" box="[450,562,1917,1940]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC06B4C1F8FAFD8DD4E2" box="[450,562,1917,1940]" italics="true" pageId="30" pageNumber="181">E. guerini</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B743F8FBFD22D4E2" box="[576,669,1916,1941]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">Fig. 18c</figureCitation>
) with apical margin rounded, not pointed as in
<taxonomicName id="4FC64D46FFFBEC06B1A9F8FAFAB9D4E3" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[1194,1286,1916,1941]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B1A9F8FAFAB9D4E3" box="[1194,1286,1916,1941]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
(
<figureCitation id="10FD2A40FFFBEC06B017F8FBFA31D4E3" box="[1300,1422,1916,1941]" captionStart="FIGURE 18" captionStartId="26.[151,250,1894,1916]" captionTargetBox="[335,1233,745,1853]" captionTargetId="figure@26.[313,1273,735,1872]" captionTargetPageId="26" captionText="FIGURE 18. Euphaea spp. [18af] vesicle of male and [18gj] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc." httpUri="https://zenodo.org/record/3096117/files/figure.png" pageId="30" pageNumber="181">Fig. 18a, b</figureCitation>
). Hämäläinen (
<emphasis id="BAB2EAD7FFFBEC06B434F826FE3CD4CF" box="[311,387,1952,1977]" italics="true" pageId="30" pageNumber="181">in litt.</emphasis>
) also confirmed that
<emphasis id="BAB2EAD7FFFBEC06B794F827FCDAD4CE" box="[663,869,1952,1977]" italics="true" pageId="30" pageNumber="181">
<taxonomicName id="4FC64D46FFFBEC06B798F827FCDAD4CE" box="[667,869,1952,1977]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">Euphaea guerini</taxonomicName>
</emphasis>
from Lak Sao, Bolikhamxay Prov.,
<collectingCountry id="F0D17655FFFBEC06B03AF826FAD0D4CE" box="[1337,1391,1953,1976]" name="Laos" pageId="30" pageNumber="181">Laos</collectingCountry>
by
<bibRefCitation id="EC574B34FFFBEC06B594F843FD9DD4AA" author="Karjalainen" box="[151,546,1988,2013]" pageId="30" pageNumber="181" refString="Karjalainen, S. &amp; Hamalainen, M. (2013) Demoiselle Damselflies: Winged Jewels of Silvery streams. Caloptera Publishing, Helsinki, 223 pp. [bilingual, in Finnish and English]" type="book" year="2013">Karjalainen &amp; Hämäläinen (2013)</bibRefCitation>
actually refers to
<taxonomicName id="4FC64D46FFFBEC06B7FFF842FCE4D4AB" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[764,859,1988,2013]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFBEC06B7FFF842FCE4D4AB" box="[764,859,1988,2013]" italics="true" pageId="30" pageNumber="181">E. saola</emphasis>
</taxonomicName>
. Similarly, earlier published records of
<taxonomicName id="4FC64D46FFFBEC07B03BF843FF57D3C6" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="31" lastPageNumber="182" order="Odonata" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="BAB2EAD7FFFBEC07B03BF843FF57D3C6" italics="true" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181">Euphaea guerini</emphasis>
</taxonomicName>
in
<collectingCountry id="F0D17655FFFAEC07B414FF10FE32D3C6" box="[279,397,151,176]" name="Cambodia" pageId="31" pageNumber="182">Cambodia</collectingCountry>
, central to southern
<collectingCountry id="F0D17655FFFAEC07B770FF1FFD13D3D9" box="[627,684,152,175]" name="Laos" pageId="31" pageNumber="182">Laos</collectingCountry>
, and central
<collectingCountry id="F0D17655FFFAEC07B63EFF10FC23D3C6" box="[829,924,151,176]" name="Vietnam" pageId="31" pageNumber="182">Vietnam</collectingCountry>
could likely include or refer to
<taxonomicName id="4FC64D46FFFAEC07B006FF1EFADDD3C6" authorityName="Phan &amp; Kompier &amp; Karube &amp; Hayashi" authorityYear="2018" box="[1285,1378,151,176]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="saola">
<emphasis id="BAB2EAD7FFFAEC07B006FF1EFADDD3C6" box="[1285,1378,151,176]" italics="true" pageId="31" pageNumber="182">E. saola</emphasis>
</taxonomicName>
(e.g. Fig. 44, p.
<quantity id="4F3E9B20FFFAEC07B413FF3BFEF6D3A3" box="[272,329,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="31" pageNumber="182" unit="in" value="31.0">31 in</quantity>
<bibRefCitation id="EC574B34FFFAEC07B453FF3BFDB5D3A2" author="Phan" box="[336,522,188,213]" pageId="31" pageNumber="182" refString="Phan, Q. T., Do, M. C. &amp; Hamalainen, M. (2011) Xuan Son National Park, a paradise for Caloptera damselflies in northern Vietnam. IDF - Report, Newsletter of the International Dragonfly Fund, 32, 1 - 34." type="journal article" year="2011">
Phan
<emphasis id="BAB2EAD7FFFAEC07B48CFF3AFE78D3A3" box="[399,455,188,213]" italics="true" pageId="31" pageNumber="182">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFFAEC07B5C4FF58FC43D236" pageId="31" pageNumber="182" type="distribution">
<paragraph id="887936C5FFFAEC07B5C4FF58FC43D236" blockId="31.[151,1436,151,320]" pageId="31" pageNumber="182">
<emphasis id="BAB2EAD7FFFAEC07B5C4FF58FEDFD38E" bold="true" box="[199,352,223,248]" pageId="31" pageNumber="182">Distribution.</emphasis>
<collectingCountry id="F0D17655FFFAEC07B469FF58FE76D38E" box="[362,457,223,248]" name="Vietnam" pageId="31" pageNumber="182">Vietnam</collectingCountry>
(
<collectingRegion id="4A02F827FFFAEC07B4D9FF67FD85D38E" box="[474,570,223,248]" country="Vietnam" name="Ha Tinh" pageId="31" pageNumber="182">
<collectingRegion id="4A02F827FFFAEC07B4D9FF67FE43D38E" box="[474,508,224,248]" country="Bhutan" name="Ha" pageId="31" pageNumber="182">Ha</collectingRegion>
Tinh
</collectingRegion>
[Hämäläinen
<emphasis id="BAB2EAD7FFFAEC07B7DDFF66FC9BD38E" box="[734,804,223,248]" italics="true" pageId="31" pageNumber="182">in litt.</emphasis>
],
<collectingRegion id="4A02F827FFFAEC07B63DFF67FC74D38E" box="[830,971,223,248]" country="Vietnam" name="Quang Binh" pageId="31" pageNumber="182">Quang Binh</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B6D9FF67FBEDD38E" box="[986,1106,223,248]" country="Vietnam" name="Quang Tri" pageId="31" pageNumber="182">Quang Tri</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B163FF58FA9FD38E" box="[1120,1312,223,248]" country="Vietnam" name="Thua Thien-Hue" pageId="31" pageNumber="182">Thua Thien-Hue</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B02DFF67FA27D38E" box="[1326,1432,224,248]" country="Vietnam" name="Da Nang" pageId="31" pageNumber="182">Da Nang</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B594FE83FE9DD26A" box="[151,290,260,284]" country="Vietnam" name="Quang Nam" pageId="31" pageNumber="182">Quang Nam</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B42CFE83FE08D26B" box="[303,439,260,285]" country="Vietnam" name="Quang Ngai" pageId="31" pageNumber="182">Quang Ngai</collectingRegion>
[Hämäläinen
<emphasis id="BAB2EAD7FFFAEC07B75BFE82FD23D26B" box="[600,668,260,285]" italics="true" pageId="31" pageNumber="182">in litt.</emphasis>
],
<collectingRegion id="4A02F827FFFAEC07B7B7FE83FCA7D26B" box="[692,792,260,285]" country="Vietnam" name="Dak Lak" pageId="31" pageNumber="182">Dak Lak</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B626FE83FCC2D26B" box="[805,893,260,285]" country="Vietnam" name="Gia Lai" pageId="31" pageNumber="182">Gia Lai</collectingRegion>
,
<collectingRegion id="4A02F827FFFAEC07B68AFE82FC4AD26A" box="[905,1013,261,284]" country="Vietnam" name="Kon Tum" pageId="31" pageNumber="182">Kon Tum</collectingRegion>
and
<collectingRegion id="4A02F827FFFAEC07B12DFE83FB10D26A" box="[1070,1199,260,285]" country="Vietnam" name="Khanh Hoa" pageId="31" pageNumber="182">Khanh Hoa</collectingRegion>
[Hämäläinen
<emphasis id="BAB2EAD7FFFAEC07B04CFE82FA2CD26B" box="[1359,1427,260,285]" italics="true" pageId="31" pageNumber="182">in litt.</emphasis>
] Provinces),
<collectingCountry id="F0D17655FFFAEC07B41FFEAFFEEDD249" box="[284,338,296,319]" name="Laos" pageId="31" pageNumber="182">Laos</collectingCountry>
(Bolikhamxay Province) (
<bibRefCitation id="EC574B34FFFAEC07B77EFEA0FC4FD236" author="Karjalainen" box="[637,1008,295,320]" pageId="31" pageNumber="182" refString="Karjalainen, S. &amp; Hamalainen, M. (2013) Demoiselle Damselflies: Winged Jewels of Silvery streams. Caloptera Publishing, Helsinki, 223 pp. [bilingual, in Finnish and English]" type="book" year="2013">Karjalainen &amp; Hämäläinen 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>