297 lines
33 KiB
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297 lines
33 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.82.e111148" ID-Pensoft-Pub="1864-8312-82-151" ID-Pensoft-UUID="8D187145BFCA55A1A73960CDE5BD124B" ID-ZooBank="43AC036E93CC4D79939A07DF54BE1A2D" ModsDocID="1864-8312-82-151" checkinTime="1711188240520" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent" docDate="2024" docId="A5AC297D1FAF55B89C817FFB71BFF7D8" docLanguage="en" docName="ArthropodSystPhyl 82: 151-181" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e111148" docTitle="Megacoxa Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024, gen. nov." docType="treatment" docUuid="7AAE84F9-483D-42A6-A210-2C0BB64CD140" docUuidSource="ZooBank" docVersion="1" id="8D187145BFCA55A1A73960CDE5BD124B" lastPageNumber="151" masterDocId="8D187145BFCA55A1A73960CDE5BD124B" masterDocTitle="Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities" masterLastPageNumber="181" masterPageNumber="151" pageNumber="151" updateTime="1711188240520" updateUser="pensoft">
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<mods:title id="AA38B9AC40BD442E6EC408B309D8FF34">Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities</mods:title>
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<mods:namePart id="2291EE2BD2A8A1CCC36F92486BAD76F8">Brazidec, Manuel</mods:namePart>
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<mods:affiliation id="995F9FAE40FE94DDB2363AF0D762F185">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="4855AC605487A99ED991325F9C50FEE8" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="306EE23BE76A0B0A4145CDD1EBBB754F">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="8158E09EE9B5F646E18DDB94914B2BDA">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark</mods:affiliation>
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<mods:namePart id="4B52C00D1B87B1E52E89BD2D7CA09FA0">Boudinot, Brendon E.</mods:namePart>
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<mods:affiliation id="931B98EE6F999AA31D2B2971C8349FF3">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany</mods:affiliation>
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<mods:namePart id="EF594D055178C84BF7843034DEB87270">Richter, Adrian</mods:namePart>
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<mods:affiliation id="723B58A013EC5A02DE742D723A4BFBC0">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan</mods:affiliation>
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<mods:namePart id="F6FE4EE1AAF9775CB37C9F8A70D63DAC">Hammel, Joerg U.</mods:namePart>
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<mods:affiliation id="26A0766AFE40B8F65B9B029CBEE6E6FE">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="38A6AE1FF8F5BFEF1B75F3C1F07D45FA">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="BCCD7FB4848FCE7794F457082CA588E1" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="86974D95F99D47DB25E05624A9498DA5">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:namePart id="9756A8E8707CA981182C73378090F5B8">Fan, Yong</mods:namePart>
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<mods:affiliation id="11EFA54231196957B39F5CF3AC6CE847">Fushun Amber Institute, Fushun 113005, China</mods:affiliation>
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<mods:namePart id="A3426FD1E59F767C6771684D0711E20F">Wang, Zhen</mods:namePart>
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<mods:affiliation id="F6407911572E6C69FD37C0461BC9086B">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="E6728FC5939D0239DCA6C9E91D0CE598">Wu, Qiong</mods:namePart>
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<mods:affiliation id="08910CAEE3294B3B01A1ABEAD857FF85">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="6486D0440E2A30CB0D3B668CFC92D1A5">Wang, Bo</mods:namePart>
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<mods:affiliation id="F924FA156870EC496EE1B6119FA49F44">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="13C1E531BAF0D316337C4C21881498D7">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="75582938070FEB10451F6928487D2FB8" type="ORCID">https://orcid.org/0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="B9640244F59857D5368CB88D0BF5CB6D">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France</mods:affiliation>
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<mods:typeOfResource id="6997DC34730B628F8FBF86CBCC7B48DB">text</mods:typeOfResource>
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<mods:title id="E862EE1DFBCEE5A8AEBD2076F67E84A0">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="3D2A05D04DD43C93E510F73E047A4204">2024</mods:date>
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<mods:number id="9E0197A2D70477DCAE493D65DF2EE6F7">2024-03-22</mods:number>
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<mods:number id="73F34252124A8717EE1373AE094020C8">82</mods:number>
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<mods:start id="9914EA2FA261355378C65E90993F8518">151</mods:start>
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<mods:end id="5D7F6345303A410FCF445B20D3362032">181</mods:end>
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<mods:url id="5F63218AD0F2675C294E7C46B406D80B">http://dx.doi.org/10.3897/asp.82.e111148</mods:url>
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<treatment id="A5AC297D1FAF55B89C817FFB71BFF7D8" LSID="urn:lsid:zoobank.org:act:7AAE84F9-483D-42A6-A210-2C0BB64CD140" httpUri="http://treatment.plazi.org/id/A5AC297D1FAF55B89C817FFB71BFF7D8" lastPageNumber="151" pageId="0" pageNumber="151">
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<subSubSection id="16571A25DAF045615710FE811C646152" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="0E09488AF0030CC08E0CE52649012395" pageId="0" pageNumber="151">
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3.1.1.3. Genus †
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<taxonomicName id="7490FD6D43B6BD523A12C646A654A4A4" LSID="https://zoobank.org/7AAE84F9-483D-42A6-A210-2C0BB64CD140" authority="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Megacoxa" order="Hymenoptera" pageId="0" pageNumber="151" rank="genus" status="gen. nov.">Megacoxa</taxonomicName>
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<taxonomicNameLabel id="A34202377DD6FDC16341424506A30E10" pageId="0" pageNumber="151">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="92B18EA8EF31B5465314FB3594777550" pageId="0" pageNumber="151" type="type species">
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<paragraph id="C33417453D8B38D5311BAD08DE9BE6A1" pageId="0" pageNumber="151">Type species.</paragraph>
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<paragraph id="91814F31D79C25E75EF26AF745186414" pageId="0" pageNumber="151">
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†
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<taxonomicName id="605DB62828C17BD7865AC863F55AF382" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" kingdom="Animalia" lsidName="Megacoxa janzeni" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="janzeni">
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<emphasis id="841521CE0B845E588E3F5F2C284B3526" italics="true" pageId="0" pageNumber="151">Megacoxa janzeni</emphasis>
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</taxonomicName>
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<emphasis id="2092ECC1CABF055C1E73156245C2FC51" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="A238D4A36B82AFC8EFCBCA13B56671B5" pageId="0" pageNumber="151" type="etymology">
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<paragraph id="6FB51580093D6AAD955CC60378EE9818" pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph id="C2E3AA3B81032FCB75D978DD09E7F2E1" pageId="0" pageNumber="151">
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The genus name refers to the size of the metacoxa, which is the key diagnostic feature of the genus; besides, the prefix
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<emphasis id="B347534DD61C0A8BD1A5E617428860AB" italics="true" pageId="0" pageNumber="151">Mega</emphasis>
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- alludes to the family name
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<taxonomicName id="EC646807AF52596E4B23E748D640CB2D" authorityName="Schletterer" authorityYear="1889" class="Insecta" family="Megalyridae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="family">Megalyridae</taxonomicName>
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. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="B539D613F5D8FEB5B2120FD7EE7A2CD2" pageId="0" pageNumber="151" type="diagnosis">
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<paragraph id="A0A000BE866256EB532E52CA8FBE1000" pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph id="A3490084A9C00F1A92658F2F0E32CD82" pageId="0" pageNumber="151">
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Compound eye oval; scape wider than long; thin occipital carina slightly crenulate. Parapsidal line present and located on posterolateral part of mesoscutum (Figs
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<figureCitation id="1B57EE41EE2EE1E6D6D471C972298E2E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. † Megacoxa chandrahrasa gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female IGR. BU- 068, habitus in dorsal view; B habitus in lateral view; C mesosoma in lateral view; D head in frontal view; E paratype female CNU-HYM-MA 2016207, habitus in lateral view; F head and mesosoma in dorsolateral view; G hind legs; H ovipositor and its sheaths. Scale bars: 1 mm (A, B, E, F); 0.5 mm (C, D, G, H). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure6" httpUri="https://binary.pensoft.net/fig/1007135" pageId="0" pageNumber="151">6C</figureCitation>
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,
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<figureCitation id="7208483CD23EE077EF58D7C19BFEBE5A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. † Megacoxa janzeni gen. et sp. nov., mid-Cretaceous, Kachin amber; A holotype female IGR. BU- 069, habitus in lateral view; B mesosoma in lateral view; C head in frontal view; D paratype male NIGP 203547, habitus in lateral view; E head and mesosoma in lateral view (scale bars: A, D = 2 mm; E = 1 mm; B, C = 0.5 mm). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure7" httpUri="https://binary.pensoft.net/fig/1007136" pageId="0" pageNumber="151">7B and E</figureCitation>
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); inner axillar groove crenulate (Fig.
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<figureCitation id="BEF1371AB5EA617DB328899203A8F730" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. † Megacoxa synchrotron gen. et sp. nov., holotype male CASENT 0753237, mid-Cretaceous, Kachin amber (E - H: reconstructed using synchrotron radiation-based micro-computed tomography). A habitus in lateral view; B head and anterior mesosoma in lateral view; C metacoxa in lateral view; D habitus in dorsal view; E habitus in dorsal view; F mesosoma in dorsal view; G mesosoma in lateral view; H head in frontal view, with antennae numerically removed. Scale bars: 1 mm (A, D, E); 0.5 mm (B, F, G, H); 0.25 mm (C). Abbreviations: AG = inner axillar groove; Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure8" httpUri="https://binary.pensoft.net/fig/1007137" pageId="0" pageNumber="151">8F</figureCitation>
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). Fore wing venation with R1, M+Cu, Sc+R, A and Cu fully pigmented; A connected to medial cell with cu-a; Rs closing marginal cell without bend (Figs
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<figureCitation id="73EA27765BB1CF106947E4A64DF27839" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H)." figureDoi="10.3897/asp.82.e111148.figure2" httpUri="https://binary.pensoft.net/fig/1007131" pageId="0" pageNumber="151">2D-F</figureCitation>
|
|
). Metacoxa much enlarged, with dorsal surface rounded to angular, outer metacoxal surface posteriorly foveate, inner metacoxal surface almost entirely accommodated in anterolateral, shallow concavity of metasoma (Figs
|
|
<figureCitation id="AF046001B46A5668D74270233A0529AB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. † Megacoxa chandrahrasa gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female IGR. BU- 068, habitus in dorsal view; B habitus in lateral view; C mesosoma in lateral view; D head in frontal view; E paratype female CNU-HYM-MA 2016207, habitus in lateral view; F head and mesosoma in dorsolateral view; G hind legs; H ovipositor and its sheaths. Scale bars: 1 mm (A, B, E, F); 0.5 mm (C, D, G, H). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure6" httpUri="https://binary.pensoft.net/fig/1007135" pageId="0" pageNumber="151">6C</figureCitation>
|
|
,
|
|
<figureCitation id="680F8F50A2B1219FB3D9A0B64053D95F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. † Megacoxa janzeni gen. et sp. nov., mid-Cretaceous, Kachin amber; A holotype female IGR. BU- 069, habitus in lateral view; B mesosoma in lateral view; C head in frontal view; D paratype male NIGP 203547, habitus in lateral view; E head and mesosoma in lateral view (scale bars: A, D = 2 mm; E = 1 mm; B, C = 0.5 mm). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure7" httpUri="https://binary.pensoft.net/fig/1007136" pageId="0" pageNumber="151">7B</figureCitation>
|
|
,
|
|
<figureCitation id="0F93C7C85D03D0FC5E9EFE689A2B0582" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. † Megacoxa synchrotron gen. et sp. nov., holotype male CASENT 0753237, mid-Cretaceous, Kachin amber (E - H: reconstructed using synchrotron radiation-based micro-computed tomography). A habitus in lateral view; B head and anterior mesosoma in lateral view; C metacoxa in lateral view; D habitus in dorsal view; E habitus in dorsal view; F mesosoma in dorsal view; G mesosoma in lateral view; H head in frontal view, with antennae numerically removed. Scale bars: 1 mm (A, D, E); 0.5 mm (B, F, G, H); 0.25 mm (C). Abbreviations: AG = inner axillar groove; Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure8" httpUri="https://binary.pensoft.net/fig/1007137" pageId="0" pageNumber="151">8C and G</figureCitation>
|
|
). Metatibia and metabasitarsus with ventral row of comb-like setae; two mesotibial and two metatibial spurs (Figs
|
|
<figureCitation id="5D38034F17FD5E8429AB76C84F601161" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. † Megacoxa chandrahrasa gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female IGR. BU- 068, habitus in dorsal view; B habitus in lateral view; C mesosoma in lateral view; D head in frontal view; E paratype female CNU-HYM-MA 2016207, habitus in lateral view; F head and mesosoma in dorsolateral view; G hind legs; H ovipositor and its sheaths. Scale bars: 1 mm (A, B, E, F); 0.5 mm (C, D, G, H). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure6" httpUri="https://binary.pensoft.net/fig/1007135" pageId="0" pageNumber="151">6A</figureCitation>
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and
|
|
<figureCitation id="FC0713B0AC4F73C136E6E4F255E831DF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. † Megacoxa janzeni gen. et sp. nov., mid-Cretaceous, Kachin amber; A holotype female IGR. BU- 069, habitus in lateral view; B mesosoma in lateral view; C head in frontal view; D paratype male NIGP 203547, habitus in lateral view; E head and mesosoma in lateral view (scale bars: A, D = 2 mm; E = 1 mm; B, C = 0.5 mm). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure7" httpUri="https://binary.pensoft.net/fig/1007136" pageId="0" pageNumber="151">7A</figureCitation>
|
|
). Metasoma slightly shorter than mesosoma, tergites and sternites beveled (Figs
|
|
<figureCitation id="7B01D7D3FE507F04E685C034C54BB87C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. † Megacoxa chandrahrasa gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female IGR. BU- 068, habitus in dorsal view; B habitus in lateral view; C mesosoma in lateral view; D head in frontal view; E paratype female CNU-HYM-MA 2016207, habitus in lateral view; F head and mesosoma in dorsolateral view; G hind legs; H ovipositor and its sheaths. Scale bars: 1 mm (A, B, E, F); 0.5 mm (C, D, G, H). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure6" httpUri="https://binary.pensoft.net/fig/1007135" pageId="0" pageNumber="151">6B</figureCitation>
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and
|
|
<figureCitation id="68ECA93E0752829618C671729E52A501" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. † Megacoxa janzeni gen. et sp. nov., mid-Cretaceous, Kachin amber; A holotype female IGR. BU- 069, habitus in lateral view; B mesosoma in lateral view; C head in frontal view; D paratype male NIGP 203547, habitus in lateral view; E head and mesosoma in lateral view (scale bars: A, D = 2 mm; E = 1 mm; B, C = 0.5 mm). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure7" httpUri="https://binary.pensoft.net/fig/1007136" pageId="0" pageNumber="151">7A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="29697A7A6E024B6E9CC8D9DE9BD10AE9" doi="10.3897/asp.82.e111148.figure6" httpUri="https://binary.pensoft.net/fig/1007135" pageId="0" pageNumber="151" start="Figure 6" startId="F6">
|
|
<paragraph id="CA5E38808315AD94110B0679AD4C8F2A" pageId="0" pageNumber="151">
|
|
<emphasis id="565ADB9EB3067C453CFD4D265CE4A31D" bold="true" pageId="0" pageNumber="151">Figure 6.</emphasis>
|
|
†
|
|
<taxonomicName id="3E1B87660BA5C03D0EEED8EC0435D680" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" kingdom="Animalia" lsidName="Megacoxa chandrahrasa" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="chandrahrasa">
|
|
<emphasis id="9188A9A9A23675A2752AE24DBCCF67D5" italics="true" pageId="0" pageNumber="151">Megacoxa chandrahrasa</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., mid-Cretaceous, Kachin amber.
|
|
<emphasis id="4C29BE97242A1C6252DEF0567F806A9F" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
holotype female IGR.BU-068, habitus in dorsal view;
|
|
<emphasis id="0F0DA72D5C319C5DB457172F9C0A4EA4" bold="true" pageId="0" pageNumber="151">B</emphasis>
|
|
habitus in lateral view;
|
|
<emphasis id="463487D6CEC2D07AC030735F711ED228" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
mesosoma in lateral view;
|
|
<emphasis id="22FAC6950910309270085562BD68243C" bold="true" pageId="0" pageNumber="151">D</emphasis>
|
|
head in frontal view;
|
|
<emphasis id="D4D671A18D2491B5D8B327F3B1EBEBD1" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
paratype female CNU-HYM-MA2016207, habitus in lateral view;
|
|
<emphasis id="B508DDC445BDBCA7C43EBAB51AF149CA" bold="true" pageId="0" pageNumber="151">F</emphasis>
|
|
head and mesosoma in dorsolateral view;
|
|
<emphasis id="57337A052214004838A3755768F21658" bold="true" pageId="0" pageNumber="151">G</emphasis>
|
|
hind legs;
|
|
<emphasis id="A97C24D40355A103ECE217F63F8ED486" bold="true" pageId="0" pageNumber="151">H</emphasis>
|
|
ovipositor and its sheaths. Scale bars: 1 mm (
|
|
<emphasis id="4EF3EA16BFFA33D47268BCE12EC69808" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
,
|
|
<emphasis id="2765D4BEA36E889FE4F986E2C103C2AE" bold="true" pageId="0" pageNumber="151">B, E, F</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="87E069DC7A87C4BB4793C6C116AE1006" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
,
|
|
<emphasis id="FF7E7656EFD4B40D50FB69A5DE4BDF1D" bold="true" pageId="0" pageNumber="151">D, G, H</emphasis>
|
|
). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="CC542B41B7275192F2B48F91AD15EF2F" doi="10.3897/asp.82.e111148.figure7" httpUri="https://binary.pensoft.net/fig/1007136" pageId="0" pageNumber="151" start="Figure 7" startId="F7">
|
|
<paragraph id="A44C13A3B4C524104EDA317A96730B67" pageId="0" pageNumber="151">
|
|
<emphasis id="C7334BCA6C870F205012F9143DD05B29" bold="true" pageId="0" pageNumber="151">Figure 7.</emphasis>
|
|
†
|
|
<taxonomicName id="552C61FF843E2B85DF396B006E8A07F1" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" kingdom="Animalia" lsidName="Megacoxa janzeni" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="janzeni">
|
|
<emphasis id="A4AD55D30F604F530FA33409E76867FE" italics="true" pageId="0" pageNumber="151">Megacoxa janzeni</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., mid-Cretaceous, Kachin amber;
|
|
<emphasis id="7667C8D18F5FCF19F38ADB67B767F190" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
holotype female IGR.BU-069, habitus in lateral view;
|
|
<emphasis id="B96FB194CC2684D7D739B2E92A53E0E1" bold="true" pageId="0" pageNumber="151">B</emphasis>
|
|
mesosoma in lateral view;
|
|
<emphasis id="03D7D5CF84DE417CA8AA3A7907A0A4DF" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
head in frontal view;
|
|
<emphasis id="DBCCF67B0E470F70BE23A3B4A8D76C28" bold="true" pageId="0" pageNumber="151">D</emphasis>
|
|
paratype male NIGP203547, habitus in lateral view;
|
|
<emphasis id="39919A1244F13FBE33B195B3C4C573C0" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
head and mesosoma in lateral view (scale bars:
|
|
<emphasis id="3290C52AECA30AEF2E4325269C4C9B08" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
,
|
|
<emphasis id="E4E6AB39A41FC6D0CA6507A1F6288EEC" bold="true" pageId="0" pageNumber="151">D</emphasis>
|
|
= 2 mm;
|
|
<emphasis id="6ACDE17861ED6618611500B8465956F9" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
= 1 mm;
|
|
<emphasis id="756630FDA10BDC38E95D0EA07441081A" bold="true" pageId="0" pageNumber="151">B</emphasis>
|
|
,
|
|
<emphasis id="CEBFE08E9EA54CB1A1A80B6F50AB4222" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
= 0.5 mm). Abbreviations: Cx = hind coxa; MeS = mesometapectal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="E213D850E3B04BA39CB75EA4E58027E1" doi="10.3897/asp.82.e111148.figure8" httpUri="https://binary.pensoft.net/fig/1007137" pageId="0" pageNumber="151" start="Figure 8" startId="F8">
|
|
<paragraph id="6B2631BD4C36A2CFD8014F39F34EE68A" pageId="0" pageNumber="151">
|
|
<emphasis id="1F4966085D9F518B6708D8A53619AAE0" bold="true" pageId="0" pageNumber="151">Figure 8.</emphasis>
|
|
†
|
|
<taxonomicName id="FFCD3810B94DB965927FE090B9DCBF53" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" kingdom="Animalia" lsidName="Megacoxa synchrotron" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="synchrotron">
|
|
<emphasis id="DCF10040634805B637D3A58D50DFF335" italics="true" pageId="0" pageNumber="151">Megacoxa synchrotron</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., holotype male CASENT0753237, mid-Cretaceous, Kachin amber (
|
|
<emphasis id="F1F32E1C4C337C0A447ADEA2D0141BE7" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
-
|
|
<emphasis id="FBD76EF4A71EAF23145C93A93CDC50A3" bold="true" pageId="0" pageNumber="151">H</emphasis>
|
|
: reconstructed using synchrotron radiation-based micro-computed tomography).
|
|
<emphasis id="B8E84B4566C3F1B903BB4003DCC8EC88" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
habitus in lateral view;
|
|
<emphasis id="778527166AA8AD0052F66CEE99EA9402" bold="true" pageId="0" pageNumber="151">B</emphasis>
|
|
head and anterior mesosoma in lateral view;
|
|
<emphasis id="DE7B73009CFE42E26DD36F7740B06EDA" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
metacoxa in lateral view;
|
|
<emphasis id="8E562622FC91893B8B0CAAC72B5F5B2E" bold="true" pageId="0" pageNumber="151">D</emphasis>
|
|
habitus in dorsal view;
|
|
<emphasis id="61A8D2CB6BD9E89D8B120ED9F75BD5E3" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
habitus in dorsal view;
|
|
<emphasis id="AFD5E708D0184CC5E0B90D585B36306E" bold="true" pageId="0" pageNumber="151">F</emphasis>
|
|
mesosoma in dorsal view;
|
|
<emphasis id="163AE57DC592E25532C1EDC052DB0D06" bold="true" pageId="0" pageNumber="151">G</emphasis>
|
|
mesosoma in lateral view;
|
|
<emphasis id="33C3C637F060D27C9164C0F37D40639D" bold="true" pageId="0" pageNumber="151">H</emphasis>
|
|
head in frontal view, with antennae numerically removed. Scale bars: 1 mm (
|
|
<emphasis id="73333A56BB60CA888F6733ECB577833C" bold="true" pageId="0" pageNumber="151">A</emphasis>
|
|
,
|
|
<emphasis id="78C640CA2938229F4A5F283CA70A9C9C" bold="true" pageId="0" pageNumber="151">D</emphasis>
|
|
,
|
|
<emphasis id="07F1557B358AE41D720A486C5BFE6412" bold="true" pageId="0" pageNumber="151">E</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="3E51879A26BAC548AB0FC58C88E34228" bold="true" pageId="0" pageNumber="151">B</emphasis>
|
|
,
|
|
<emphasis id="D124EEEB1AE06D563D750A74A9C65B85" bold="true" pageId="0" pageNumber="151">F</emphasis>
|
|
,
|
|
<emphasis id="92D74B500CDDF908DB987884D90771F5" bold="true" pageId="0" pageNumber="151">G</emphasis>
|
|
,
|
|
<emphasis id="E67D14E58897C549728830ECA74541AE" bold="true" pageId="0" pageNumber="151">H</emphasis>
|
|
); 0.25 mm (
|
|
<emphasis id="EF40E66747E0FA8F610484CE21F5872D" bold="true" pageId="0" pageNumber="151">C</emphasis>
|
|
). Abbreviations: AG = inner axillar groove; Cx = hind coxa; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; PpL = parapsidal line; sp = anterior thoracic spiracle.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="3A23F6C6FB8B30F92682E5AFA4574B44" pageId="0" pageNumber="151" type="comments">
|
|
<paragraph id="589E4A1B96A1CA6797F9171D04DB6C35" pageId="0" pageNumber="151">Comments.</paragraph>
|
|
<paragraph id="62EF5D558BE14CFBEC5AF17E2F229BF0" pageId="0" pageNumber="151">
|
|
The enlarged metacoxa accommodated in the anterior metasomal concavity is the key diagnostic feature for †
|
|
<taxonomicName id="AA6D15889C1778B3FA130DABD09F5773" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Megacoxa" kingdom="Animalia" lsidName="Megacoxa" order="Hymenoptera" pageId="0" pageNumber="151" rank="genus">
|
|
<emphasis id="A12D535B8FD01DCB7EEF3D59413434CF" italics="true" pageId="0" pageNumber="151">Megacoxa</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="D79BD2BA9C6AC822931730743FCD8EA8" bold="true" pageId="0" pageNumber="151">gen. nov.</emphasis>
|
|
This condition is present in all five specimens assigned to this genus and the metacoxae are clearly symmetrical in the synchrotron-scanned specimen, thus ruling out a preservational artefact.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |