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<document id="BAFBFF633C997818760F88875CC389AC" ID-DOI="10.5281/zenodo.202987" ID-GBIF-Dataset="71791e06-ffac-4895-8de0-e0c3557537cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="202987" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460273994055" checkinUser="plazi" docAuthor="Wang, Bo &amp; Zhang, Haichun" docDate="2011" docId="6674E4370C49FF8583AFFF006358B215" docLanguage="en" docName="zt02811p052.pdf" docOrigin="Zootaxa 2811" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lithohypna Nikolajev, Wang" docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="9A4D9C4F0C48FF878338FFDD6709B309" masterDocTitle="A new fossil genus of the family Glaphyridae (Coleoptera: Scarabaeoidea) from the Lower Cretaceous Yixian Formation" masterLastPageNumber="52" masterPageNumber="47" pageNumber="48" updateTime="1698247603497" updateUser="plazi">
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<mods:title id="8363B24EAF54ACC36F074C0F8003B4C0">A new fossil genus of the family Glaphyridae (Coleoptera: Scarabaeoidea) from the Lower Cretaceous Yixian Formation</mods:title>
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<mods:namePart id="B31B392A03B3899101C79AB3062251DD">Wang, Bo</mods:namePart>
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<mods:date id="E230C7430F4E1E85BCB986832F1DE1CF">2011</mods:date>
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<treatment id="6674E4370C49FF8583AFFF006358B215" ID-DOI="http://doi.org/10.5281/zenodo.6194289" ID-GBIF-Taxon="119395653" ID-Zenodo-Dep="6194289" LSID="urn:lsid:plazi:treatment:6674E4370C49FF8583AFFF006358B215" httpUri="http://treatment.plazi.org/id/6674E4370C49FF8583AFFF006358B215" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="48">
<subSubSection id="A6C706AA0C49FF8683AFFF006469B3FE" box="[151,864,221,247]" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="EE6255210C49FF8683AFFF006469B3FE" blockId="1.[151,864,221,247]" box="[151,864,221,247]" pageId="1" pageNumber="48">
<heading id="B52AE24D0C49FF8683AFFF006469B3FE" bold="true" box="[151,864,221,247]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="DCA989330C49FF8683AFFF006469B3FE" bold="true" box="[151,864,221,247]" pageId="1" pageNumber="48">
Genus
<taxonomicName id="29DD2EA20C49FF8683D6FF00655BB3FE" ID-CoL="5FMQ" authority="Nikolajev, Wang" authorityName="Nikolajev, Wang" box="[238,594,221,247]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683D6FF006673B3FE" bold="true" box="[238,378,221,247]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
Nikolajev, Wang
</taxonomicName>
, &amp; Zhang, new genus
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A6C706AA0C49FF8683AFFEF9649CB235" box="[151,917,291,316]" pageId="1" pageNumber="48" type="materials_examined">
<paragraph id="EE6255210C49FF8683AFFEF9649CB235" blockId="1.[151,1436,291,641]" box="[151,917,291,316]" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683AFFEF9663BB235" bold="true" box="[151,306,291,316]" pageId="1" pageNumber="48">
<typeStatus id="3166EB830C49FF8683AFFEF967DBB235" box="[151,210,292,316]" pageId="1" pageNumber="48">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="29DD2EA20C49FF868201FEF96540B235" box="[313,585,292,316]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="chifengensis" status="sp. nov.">
<emphasis id="DCA989330C49FF868201FEF96540B235" box="[313,585,292,316]" italics="true" pageId="1" pageNumber="48">Lithohypna chifengensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C79A34480C49FF868169FEF865DEB235" box="[593,727,292,316]" pageId="1" pageNumber="48" rank="species">new species</taxonomicNameLabel>
here designated.
</paragraph>
</subSubSection>
<subSubSection id="A6C706AA0C49FF8683FDFE95650EB28D" pageId="1" pageNumber="48" type="etymology">
<paragraph id="EE6255210C49FF8683FDFE95650EB28D" blockId="1.[151,1436,291,641]" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683FDFE956645B269" bold="true" box="[197,332,328,352]" pageId="1" pageNumber="48">Etymology.</emphasis>
The name is derived from lithos, which is Greek for stone, and the generic name
<taxonomicName id="29DD2EA20C49FF8687E4FE956626B28D" authority="Eschoscholtz, 1818" authorityName="Eschoscholtz" authorityYear="1818" class="Insecta" family="Glaphyridae" genus="Anthypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8687E4FE956241B269" box="[1244,1352,328,352]" italics="true" pageId="1" pageNumber="48">Anthypna</emphasis>
Eschoscholtz, 1818
</taxonomicName>
. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="A6C706AA0C49FF8683FDFE5265D9B5D8" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="EE6255210C49FF8683FDFE5267E4B155" blockId="1.[151,1436,291,641]" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683FDFE526636B2A1" bold="true" box="[197,319,399,424]" pageId="1" pageNumber="48">Diagnosis.</emphasis>
Winged. Body elongate, length about
<quantity id="2925F8C40C49FF8681CBFE52644CB2AE" box="[755,837,399,423]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="48" unit="mm" value="15.0">15 mm</quantity>
. Labrum large, approximately twice as wide as long, somewhat bilobed, sinuate in the middle. Clypeus with truncate, straight apical margin. Scutellum exposed, Ushaped. Elytra elongate, without striae, dehiscent at apex. Elytral lateral margin arcuate. Pygidium visible beyond elytra. Mesocoxae moderately separated. Mesofemur narrow. Protibia with three large denticles on the outer margin. Protarsus longer than protibia, non-lamellate medially. Metatibia with single, transverse carina on the outer margin.
</paragraph>
<paragraph id="EE6255210C49FF8683FDFDBA63D6B176" blockId="1.[151,1436,291,641]" box="[197,1247,615,641]" pageId="1" pageNumber="48">
The key differences between
<taxonomicName id="29DD2EA20C49FF868136FDB56587B189" box="[526,654,616,640]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868136FDB56587B189" box="[526,654,616,640]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
</taxonomicName>
and other
<taxonomicName id="29DD2EA20C49FF86803CFDBA6487B176" box="[772,910,615,639]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
genera is given in the table 1.
</paragraph>
<caption id="BAA205A90C49FF8683AFFD6D65B9B1FE" pageId="1" pageNumber="48">
<paragraph id="EE6255210C49FF8683AFFD6D65B9B1FE" blockId="1.[151,1391,688,1162]" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683AFFD6D660DB1CC" bold="true" box="[151,260,688,709]" pageId="1" pageNumber="48">TABLE 1.</emphasis>
<taxonomicName id="29DD2EA20C49FF868232FD6D668EB1CF" box="[266,391,688,710]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
generic characters—see Figs. 1–7 for further details of characters and character states. Genera Males characters
</paragraph>
</caption>
<paragraph id="EE6255210C49FF86813DFCD66617B769" blockId="1.[151,1391,688,1162]" pageId="1" pageNumber="48">
1 2 3 4 5 6 7
<date id="9A6373E10C49FF86876AFCD6621DB029" box="[1106,1300,779,800]" pageId="1" pageNumber="48" value="2010-09-08">8 9 10</date>
11
<taxonomicName id="29DD2EA20C49FF8683A7FCE966B7B040" authority="Eschscholtz, 1818" authorityName="Eschscholtz" authorityYear="1818" box="[159,446,819,842]" class="Insecta" family="Glaphyridae" genus="Anthypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FCE96609B043" box="[159,256,820,842]" italics="true" pageId="1" pageNumber="48">Anthypna</emphasis>
Eschscholtz, 1818
</taxonomicName>
b b b b a a c a a b a
<taxonomicName id="29DD2EA20C49FF8683A7FC8066BAB07B" authority="Latreille, 1807" authorityName="Latreille" authorityYear="1807" box="[159,435,861,883]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FC806611B07A" box="[159,280,861,883]" italics="true" pageId="1" pageNumber="48">Amphicoma</emphasis>
Latreille, 1807
</taxonomicName>
b a b b-c a b b b a b a
<taxonomicName id="29DD2EA20C49FF8683A7FC5A6519B095" authority="Nikolajev, 2005" authorityName="Nikolajev" authorityYear="2005" box="[159,528,902,925]" class="Insecta" family="Glaphyridae" genus="Cretoglaphyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FC5A6634B094" box="[159,317,903,925]" italics="true" pageId="1" pageNumber="48">Cretoglaphyrus</emphasis>
<bibRefCitation id="8A4C28D00C49FF86827BFC5B6519B095" author="Nikolajev" box="[323,528,902,924]" pageId="1" pageNumber="48" refString="Nikolajev, G. V. (2005) Sсarab beetles of the subfamily Glaphyrinae (Coleoptera, Sсarabaeidae) in the Lover Cretaceous of Transbaikalia [Plastinchatousye zhuki podsemeystva Glaphyrinae (Coleoptera, Sсarabaeidae) v nizhnem mele Zabaykal'ya]. Zhivotnyy mir Dal'nego Vostoka, 5, 57 - 66. (In Russian with English abstract)" type="journal article" year="2005" yearSuffix="a">Nikolajev, 2005 a</bibRefCitation>
</taxonomicName>
?? b a c?? a a b a
<taxonomicName id="29DD2EA20C49FF8683A7FC6D667CB0CC" authority="Truqui, 1848" authorityName="Truqui" authorityYear="1848" box="[159,373,944,966]" class="Insecta" family="Glaphyridae" genus="Eulasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FC6D67E4B0CF" box="[159,237,944,966]" italics="true" pageId="1" pageNumber="48">Eulasia</emphasis>
Truqui, 1848
</taxonomicName>
b b b b a a b a a b a
<taxonomicName id="29DD2EA20C49FF8683A7FC0466ACB0E6" authority="Latreille, 1807" authorityName="Latreille" authorityYear="1807" box="[159,421,985,1007]" class="Insecta" family="Glaphyridae" genus="Glaphyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FC046603B0E6" box="[159,266,985,1007]" italics="true" pageId="1" pageNumber="48">Glaphyrus</emphasis>
Latreille, 1807
</taxonomicName>
a b b a a-b b a a b a a-b
<taxonomicName id="29DD2EA20C49FF8683A7FBDF66CCB711" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[159,453,1026,1048]" class="Insecta" family="Glaphyridae" genus="Lichnanthe" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FBDF6618B711" box="[159,273,1026,1048]" italics="true" pageId="1" pageNumber="48">Lichnanthe</emphasis>
Burmeister, 1844
</taxonomicName>
b a a b a a a a a a a
<taxonomicName id="29DD2EA20C49FF8683A7FBF166CDB74B" authority="Nikolajev, Wang" authorityName="Nikolajev, Wang" box="[159,452,1067,1090]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FBF1661BB74B" box="[159,274,1068,1090]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
Nikolajev, Wang
</taxonomicName>
, &amp; a b a b a a?a a a a c Zhang, 2011
</paragraph>
<paragraph id="EE6255210C49FF8683A7FBA96253B783" blockId="1.[151,1391,688,1162]" box="[159,1370,1139,1162]" pageId="1" pageNumber="48">
<taxonomicName id="29DD2EA20C49FF8683A7FBA966EFB780" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[159,486,1139,1162]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8683A7FBA96616B783" box="[159,287,1140,1162]" italics="true" pageId="1" pageNumber="48">Pygopleurus</emphasis>
Motschulsky, 1860
</taxonomicName>
b b b a a a c a a b a
</paragraph>
<paragraph id="EE6255210C49FF8683AFFB766685B7D6" blockId="1.[151,1437,1194,1676]" pageId="1" pageNumber="48">1. Mesoepimeron: a—clearly visible from above between pronotum and elytron; b—completely covered by elytra and not visible from above.</paragraph>
<paragraph id="EE6255210C49FF8683AFFB3565E5B615" blockId="1.[151,1437,1194,1676]" pageId="1" pageNumber="48">2. Antennal club: a—second antennomere free (antennal club usually elongate); b—second antennomere partially enclosed by the first antennomere (antennal club usually short).</paragraph>
<paragraph id="EE6255210C49FF8683AFFAFB6303B632" blockId="1.[151,1437,1194,1676]" box="[151,1034,1317,1339]" pageId="1" pageNumber="48">3. Labrum: a—about 2.2 times wider than long; b—about 4–6 times wider than long.</paragraph>
<paragraph id="EE6255210C49FF8683AFFA9964DAB653" blockId="1.[151,1437,1194,1676]" box="[151,979,1347,1370]" pageId="1" pageNumber="48">4. Scutellum: a—triangular; b—U-shaped and broad; c—U-shaped and narrow.</paragraph>
<paragraph id="EE6255210C49FF8683AFFABF661EB69E" blockId="1.[151,1437,1194,1676]" pageId="1" pageNumber="48">5. Elytron: a—without longitudinal carinae; b—with low and wide longitudinal carinae; c—with sharp and thin longitudinal carinae.</paragraph>
<paragraph id="EE6255210C49FF8683AFFA7D648EB6BF" blockId="1.[151,1437,1194,1676]" box="[151,903,1440,1462]" pageId="1" pageNumber="48">
6. Protibia: a—shorter than protarsus (
<figureCitation id="76E649A40C49FF86810AFA7D6579B6BF" box="[562,624,1440,1462]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Fig. 7</figureCitation>
); b—longer than protarsus.
</paragraph>
<paragraph id="EE6255210C49FF8683AFFA6266D4B6FD" blockId="1.[151,1437,1194,1676]" pageId="1" pageNumber="48">
7. Protarsus: a—tarsomeres 1–4 unmodified (
<figureCitation id="76E649A40C49FF868143FA6365B3B6DD" box="[635,698,1470,1492]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Fig. 7</figureCitation>
); b—tarsomeres 1–4 lamellate on inside (
<figureCitation id="76E649A40C49FF86875AFA6363CDB6DC" box="[1122,1220,1470,1493]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Figs. 1–2</figureCitation>
); c—tarsomeres pectinate on inside (
<figureCitation id="76E649A40C49FF868257FA0066DBB6FD" box="[367,466,1501,1524]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Figs. 3–4</figureCitation>
).
</paragraph>
<paragraph id="EE6255210C49FF8683AFFA216421B518" blockId="1.[151,1437,1194,1676]" box="[151,808,1531,1554]" pageId="1" pageNumber="48">
8. Mesotibia: a—apex unmodified; b—apex lamellate (
<figureCitation id="76E649A40C49FF8681E5FA266413B518" box="[733,794,1531,1553]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="EE6255210C49FF8683AFF9C66517B539" blockId="1.[151,1437,1194,1676]" box="[151,542,1562,1584]" pageId="1" pageNumber="48">9. Metafemur: a—slender; b—wide.</paragraph>
<paragraph id="EE6255210C49FF8683AFF9E7648BB546" blockId="1.[151,1437,1194,1676]" box="[151,898,1593,1615]" pageId="1" pageNumber="48">10. Lateral margin of metatibia: a—with one carina; b—without carinae.</paragraph>
<paragraph id="EE6255210C49FF8683AFF9856649B585" blockId="1.[151,1437,1194,1676]" pageId="1" pageNumber="48">
11. Apex of metatibia: a—unmodified; b—with immobile spur; c—lamellate, like apex of mesotibia of
<taxonomicName id="29DD2EA20C49FF8687EAF9856242B567" box="[1234,1355,1624,1646]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF8687EAF9856242B567" box="[1234,1355,1624,1646]" italics="true" pageId="1" pageNumber="48">Amphicoma</emphasis>
</taxonomicName>
species (see
<figureCitation id="76E649A40C49FF8683CEF9AB663BB585" box="[246,306,1654,1676]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1105,1128]" captionTargetBox="[217,1378,350,1075]" captionTargetId="figure@2.[206,1381,339,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Extant Glaphyridae genera, leg features: 1 – 2 — Amphicoma, protarsus [1 — A. fairmairei (Semenov), 2 — A. latouchei (Fairmaire)]; 3 — Eulasia, protarsus of E. (Solskiola) analis (Solsky); 4 — Pygopleurus, protarsus of P. vulpes (Fabricius); 5 — Amphicoma, mesotibia apex of A. latouchei (Fairmaire); 6 – 7 — Glaphyrus festivus Ménétriés (6 — mesotibia apex, 7 — proleg) (after Medvedev [1960])." httpUri="https://zenodo.org/record/202988/files/figure.png" pageId="1" pageNumber="48">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="EE6255210C49FF8683FDF96465D9B5D8" blockId="1.[151,1437,1721,2034]" box="[197,720,1721,1746]" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683FDF964666DB5DB" bold="true" box="[197,356,1721,1746]" pageId="1" pageNumber="48">Composition.</emphasis>
Only the
<typeStatus id="3166EB830C49FF8682ECF967650DB5DB" box="[468,516,1722,1746]" pageId="1" pageNumber="48">type</typeStatus>
species is known.
</paragraph>
</subSubSection>
<subSubSection id="A6C706AA0C49FF8583FDF9006358B215" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48" type="discussion">
<paragraph id="EE6255210C49FF8583FDF9006358B215" blockId="1.[151,1437,1721,2034]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<emphasis id="DCA989330C49FF8683FDF9006627B5FC" bold="true" box="[197,302,1757,1781]" pageId="1" pageNumber="48">Remark.</emphasis>
<taxonomicName id="29DD2EA20C49FF86820FF90066BEB5FC" box="[311,439,1757,1781]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF86820FF90066BEB5FC" box="[311,439,1757,1781]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
</taxonomicName>
characters may be a complex of plesiomorphies, synapomorphies and autapomorphies. Characters such as a large labrum, long protarsus with unmodified tarsomeres
<quantity id="2925F8C40C49FF868736F8DC635FB410" box="[1038,1110,1793,1817]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="1" pageNumber="48" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
males, and unmodified apex of mesotibia in males are likely plesiomorphic. The mesoepimeron being clearly visible from above is a possible synapomorphy with the genera
<taxonomicName id="29DD2EA20C49FF868138F894657FB468" box="[512,630,1865,1889]" class="Insecta" family="Glaphyridae" genus="Glaphyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868138F894657FB468" box="[512,630,1865,1889]" italics="true" pageId="1" pageNumber="48">Glaphyrus</emphasis>
</taxonomicName>
and
<taxonomicName id="29DD2EA20C49FF868188F8946427B468" box="[688,814,1865,1889]" class="Insecta" family="Glaphyridae" genus="Lichnanthe" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868188F8946427B468" box="[688,814,1865,1889]" italics="true" pageId="1" pageNumber="48">Lichnanthe</emphasis>
</taxonomicName>
. The antennal club with second antennomere partially enclosed by the first antennomere is a possible synapomorphy with the genera
<emphasis id="DCA989330C49FF868720F8B063E1B48C" box="[1048,1256,1901,1925]" italics="true" pageId="1" pageNumber="48">
<taxonomicName id="29DD2EA20C49FF868720F8B0638FB48C" box="[1048,1158,1901,1925]" class="Insecta" family="Glaphyridae" genus="Anthypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">Anthypna</taxonomicName>
,
<taxonomicName id="29DD2EA20C49FF8687ABF8B063E1B48C" box="[1171,1256,1901,1925]" class="Insecta" family="Glaphyridae" genus="Eulasia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">Eulasia</taxonomicName>
</emphasis>
, and
<taxonomicName id="29DD2EA20C49FF868611F8B067B2B4A0" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868611F8B067B2B4A0" italics="true" pageId="1" pageNumber="48">Pygopleurus</emphasis>
</taxonomicName>
. Undoubtedly, the modified apex of the metatibia in males is an autapomorphy. A phylogenetics analysis will be needed to determine if the genus
<taxonomicName id="29DD2EA20C49FF868101F86865B1B4C4" box="[569,696,1973,1997]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868101F86865B1B4C4" box="[569,696,1973,1997]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
</taxonomicName>
is a sister group to any of the extant taxa of the same rank. The Mesozoic genus
<taxonomicName id="29DD2EA20C49FF86826CF804650AB4F8" box="[340,515,2009,2033]" class="Insecta" family="Glaphyridae" genus="Cretoglaphyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF86826CF804650AB4F8" box="[340,515,2009,2033]" italics="true" pageId="1" pageNumber="48">Cretoglaphyrus</emphasis>
</taxonomicName>
might also be a sister group to
<taxonomicName id="29DD2EA20C49FF86805EF80464EFB4F8" box="[870,998,2009,2033]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF86805EF80464EFB4F8" box="[870,998,2009,2033]" italics="true" pageId="1" pageNumber="48">Lithohypna</emphasis>
</taxonomicName>
as
<taxonomicName id="29DD2EA20C49FF868734F80463B2B4F8" box="[1036,1211,2009,2033]" class="Insecta" family="Glaphyridae" genus="Cretoglaphyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C49FF868734F80463B2B4F8" box="[1036,1211,2009,2033]" italics="true" pageId="1" pageNumber="48">Cretoglaphyrus</emphasis>
</taxonomicName>
has characters such as a short upper lip (autapomorphy or possible synapomorphy with the extant
<taxonomicName id="29DD2EA20C4AFF858738FF4A6382B3A6" box="[1024,1163,151,175]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
genera) and an unmodified apex of the mesotibiae and metatibiae in males (possible plesiomorphies). Until a thorough phylogenetic analysis provides better evidence for evolutionary relationships among
<taxonomicName id="29DD2EA20C4AFF8580B1FF02631DB3FE" box="[905,1044,223,247]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
taxa, the genus
<taxonomicName id="29DD2EA20C4AFF8587F2FF3D6243B3F1" box="[1226,1354,224,248]" class="Insecta" family="Glaphyridae" genus="Lithohypna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="DCA989330C4AFF8587F2FF3D6243B3F1" box="[1226,1354,224,248]" italics="true" pageId="2" pageNumber="49">Lithohypna</emphasis>
</taxonomicName>
should be considered as
<emphasis id="DCA989330C4AFF85826FFED866FBB215" box="[343,498,260,284]" italics="true" pageId="2" pageNumber="49">incertae sedis</emphasis>
and not placed in any of the
<taxonomicName id="29DD2EA20C4AFF85800EFED964C8B215" box="[822,961,260,284]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
subfamilies.
</paragraph>
</subSubSection>
</treatment>
</document>
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