220 lines
20 KiB
XML
220 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.983.52630" ID-GBIF-Dataset="d3b857f7-25c1-4b21-9d39-4510ab675fca" ID-PMC="PMC7655787" ID-Pensoft-Pub="1313-2970-983-1" ID-Pensoft-UUID="C890C7FD4B2D57A8B1A062305ED42D53" ID-PubMed="33244289" ID-ZooBank="7DED4CE2934C4539945F758930C927F9" ModsDocID="1313-2970-983-1" checkinTime="1604475085656" checkinUser="pensoft" docAuthor="Smith, Sarah M., Beaver, Roger A. & Cognato, Anthony I." docDate="2020" docId="665ABE3EB56B5A68ADCA3FE194EFCA72" docLanguage="en" docName="ZooKeys 983: 1-442" docOrigin="ZooKeys 983" docSource="http://dx.doi.org/10.3897/zookeys.983.52630" docTitle="Debus emarginatus" docType="treatment" docVersion="4" id="C890C7FD4B2D57A8B1A062305ED42D53" lastPageNumber="1" masterDocId="C890C7FD4B2D57A8B1A062305ED42D53" masterDocTitle="A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China" masterLastPageNumber="442" masterPageNumber="1" pageNumber="1" updateTime="1668169715782" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smith, Sarah M.</mods:namePart>
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<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
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<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beaver, Roger A.</mods:namePart>
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<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cognato, Anthony I.</mods:namePart>
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<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>983</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>442</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.983.52630</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
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<mods:identifier type="ZooBank">7DED4CE2934C4539945F758930C927F9</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C890C7FD4B2D57A8B1A062305ED42D53</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168754724" LSID="urn:lsid:plazi:treatment:665ABE3EB56B5A68ADCA3FE194EFCA72" httpUri="http://treatment.plazi.org/id/665ABE3EB56B5A68ADCA3FE194EFCA72" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="(Eichhoff, 1878)" baseAuthorityName="Eichhoff" baseAuthorityYear="1878" class="Insecta" family="Curculionidae" genus="Debus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Debus emarginatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginatus">Debus emarginatus (Eichhoff, 1878)</taxonomicName>
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<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Dorsal, lateral and declivital view of Debus emarginatus, 3.3 - 3.6 mm (A, B, I), D. pumilus, 2.5 - 3.2 mm (C, D, J), D. quadrispinus, 2.2 - 2.5 mm (E, F, K), and D. shoreae, 3.0 - 3.8 mm (G, H, L)." figureDoi="10.3897/zookeys.983.52630.figure48" httpUri="https://binary.pensoft.net/fig/470848" pageId="0" pageNumber="1">Fig. 48A, B, I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus emarginatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginatus">Xyleborus emarginatus</taxonomicName>
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Eichhoff, 1878a: 392.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Eichhoff" baseAuthorityYear="1878" class="Insecta" family="Curculionidae" genus="Debus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Debus emarginatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginatus">Debus emarginatus</taxonomicName>
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(Eichhoff):
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2717.1.1" author="Hulcr, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 33" refId="B202" refString="Hulcr, J, Cognato, AI, 2010a. New genera of Palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae) based on congruence between morphological and molecular characters. Zootaxa 2717: 1 - 33, DOI: https://doi.org/10.11646/zootaxa.2717.1.1" title="New genera of Palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae) based on congruence between morphological and molecular characters." url="https://doi.org/10.11646/zootaxa.2717.1.1" volume="2717" year="2010 a">Hulcr and Cognato 2010a</bibRefCitation>
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: 14.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Blandford" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus exesus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="exesus">Xyleborus exesus</taxonomicName>
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Blandford, 1894b: 119. Synonymy:
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.56.520" author="Hulcr, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="105 - 119" refId="B200" refString="Hulcr, J, 2010. Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae). ZooKeys 56: 105 - 119, DOI: https://doi.org/10.3897/zookeys.56.520" title="Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.3897/zookeys.56.520" volume="56" year="2010">Hulcr 2010</bibRefCitation>
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: 111.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Curculionidae" genus="Ips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ips cinchonae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cinchonae">Ips cinchonae</taxonomicName>
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Veen, 1897: 135. Synonymy:
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<bibRefCitation author="Kalshoven, LGE" journalOrPublisher="Entomologische Berichten, Amsterdam" pageId="0" pageNumber="1" pagination="93 - 97" refId="B219" refString="Kalshoven, LGE, 1959a. New cases of synonymy in Indomalayan scolytids. Entomologische Berichten, Amsterdam 19: 93 - 97" title="New cases of synonymy in Indomalayan scolytids." volume="19" year="1959 a">Kalshoven 1959a</bibRefCitation>
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: 96.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Hagedorn" authorityYear="1910" class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus cordatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cordatus">Xyleborus cordatus</taxonomicName>
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Hagedorn, 1910a: 12. Synonymy:
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<bibRefCitation author="Schedl, KE" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="1 - 49" refId="B371" refString="Schedl, KE, 1942c. Neue Scolytidae aus Java. 76. Beitrag zur Morphologie und Systematik der Scolytoidea. Tijdschrift voor Entomologie 85: 1 - 49" title="Neue Scolytidae aus Java. 76. Beitrag zur Morphologie und Systematik der Scolytoidea." volume="85" year="1942 c">Schedl 1942c</bibRefCitation>
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: 6.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus palmeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="palmeri">Xyleborus palmeri</taxonomicName>
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Hopkins, 1915a: 54. Synonymy:
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.56.520" author="Hulcr, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="105 - 119" refId="B200" refString="Hulcr, J, 2010. Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae). ZooKeys 56: 105 - 119, DOI: https://doi.org/10.3897/zookeys.56.520" title="Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.3897/zookeys.56.520" volume="56" year="2010">Hulcr 2010</bibRefCitation>
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: 111.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus terminaliae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terminaliae">Xyleborus terminaliae</taxonomicName>
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Hopkins, 1915a: 54. Synonymy:
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.56.520" author="Hulcr, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="105 - 119" refId="B200" refString="Hulcr, J, 2010. Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae). ZooKeys 56: 105 - 119, DOI: https://doi.org/10.3897/zookeys.56.520" title="Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.3897/zookeys.56.520" volume="56" year="2010">Hulcr 2010</bibRefCitation>
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: 110.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus emarginatus subsp. semicircularis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="emarginatus" subSpecies="semicircularis">Xyleborus emarginatus semicircularis</taxonomicName>
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Schedl, 1973: 92. Synonymy:
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.22642" author="Wood, SL" journalOrPublisher="The Great Basin Naturalist" pageId="0" pageNumber="1" pagination="167 - 185" refId="B487" refString="Wood, SL, 1989. Nomenclatural changes and new species of Scolytidae (Coleoptera), part IV. The Great Basin Naturalist 49: 167 - 185, DOI: https://doi.org/10.5962/bhl.part.22642" title="Nomenclatural changes and new species of Scolytidae (Coleoptera), part IV." url="https://doi.org/10.5962/bhl.part.22642" volume="49" year="1989">Wood 1989</bibRefCitation>
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: 176.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus emarginatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Xyleborus emarginatus</emphasis>
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</taxonomicName>
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(MIZ).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
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<taxonomicName authorityName="Blandford" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus exesus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="exesus">
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<emphasis italics="true" pageId="0" pageNumber="1">Xyleborus exesus</emphasis>
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</taxonomicName>
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(NHMUK, 2).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus palmeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="palmeri">
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<emphasis italics="true" pageId="0" pageNumber="1">Xyleborus palmeri</emphasis>
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</taxonomicName>
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(NMNH).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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<taxonomicName class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus terminaliae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terminaliae">
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<emphasis italics="true" pageId="0" pageNumber="1">Xyleborus terminaliae</emphasis>
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</taxonomicName>
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(NMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="new records">
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<paragraph pageId="0" pageNumber="1">New records.</paragraph>
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<paragraph pageId="0" pageNumber="1">China: N Guangxi reg., Miaoershan, S slope, 1300-2000 m, 25-28.vi.1997, Bolm (RABC, 1).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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3.3-3.6 mm long (mean = 3.48 mm; n = 4); 2.83-3.0
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide. This species is distinguished by the posterolateral extensions of elytra short, less than the width of apical emargination, and declivity shallowly excavated; declivity clearly, confusedly punctate.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is very similar to
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<taxonomicName lsidName="D. shoreae" pageId="0" pageNumber="1" rank="species" species="shoreae">
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<emphasis italics="true" pageId="0" pageNumber="1">D. shoreae</emphasis>
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</taxonomicName>
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and is distinguished by the punctation of the declivity.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="similar species">
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<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Smith & Beaver & Cognato" authorityYear="2020" baseAuthorityName="Motschulsky" baseAuthorityYear="1863" class="Insecta" family="Curculionidae" genus="Debus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Debus quadrispinus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quadrispinus">
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<emphasis italics="true" pageId="0" pageNumber="1">Debus quadrispinus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="D. shoreae" pageId="0" pageNumber="1" rank="species" species="shoreae">
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<emphasis italics="true" pageId="0" pageNumber="1">D. shoreae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">From India and southern China through southeast Asia, the Philippines and Indonesia to New Guinea and the Solomon Islands in the East, northwards to Japan. Recorded in the study region from China (Fujian, Guangxi*, Guizhou, Hubei, Hunan, Shaanxi, Shanxi, Sichuan, Xizang, Yunnan), India (Nicobar Is), Laos, Taiwan, Thailand, Vietnam.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="host">
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<paragraph pageId="0" pageNumber="1">Host plants.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Strongly polyphagous (e.g.,
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<bibRefCitation author="Browne, FG" journalOrPublisher="Malayan Forest Records" pageId="0" pageNumber="1" pagination="1 - 255" refId="B70" refString="Browne, FG, 1961b. The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records 22: 1 - 255" title="The biology of Malayan Scolytidae and Platypodidae." volume="22" year="1961 b">Browne 1961b</bibRefCitation>
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;
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<bibRefCitation author="Ohno, S" journalOrPublisher="Research Bulletin of the Plant Protection Service, Japan" pageId="0" pageNumber="1" pagination="83 - 94" refId="B298" refString="Ohno, S, 1990. The Scolytidae and Platypodidae (Coleoptera) from Borneo found in logs at Nagoya port 1. Research Bulletin of the Plant Protection Service, Japan 26: 83 - 94" title="The Scolytidae and Platypodidae (Coleoptera) from Borneo found in logs at Nagoya port 1." volume="26" year="1990">Ohno 1990</bibRefCitation>
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;
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<bibRefCitation author="Wood, SL" journalOrPublisher="The Great Basin Naturalist Memoirs" pageId="0" pageNumber="1" pagination="1 - 1553" refId="B490" refString="Wood, SL, Bright, DE, 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic index. The Great Basin Naturalist Memoirs 13: 1 - 1553" title="A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic index." volume="13" year="1992">Wood and Bright 1992</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Browne, FG" journalOrPublisher="Malayan Forest Records" pageId="0" pageNumber="1" pagination="1 - 255" refId="B70" refString="Browne, FG, 1961b. The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records 22: 1 - 255" title="The biology of Malayan Scolytidae and Platypodidae." volume="22" year="1961 b">Browne (1961b)</bibRefCitation>
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provides further information on the habits of the species.
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</paragraph>
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<caption doi="10.3897/zookeys.983.52630.figure48" httpUri="https://binary.pensoft.net/fig/470848" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
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<paragraph pageId="0" pageNumber="1">
|
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
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Dorsal, lateral and declivital view of
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<taxonomicName baseAuthorityName="Eichhoff" baseAuthorityYear="1878" class="Insecta" family="Curculionidae" genus="Debus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Debus emarginatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Debus emarginatus</emphasis>
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</taxonomicName>
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, 3.3-3.6 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">A, B, I</emphasis>
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),
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<taxonomicName lsidName="D. pumilus" pageId="0" pageNumber="1" rank="species" species="pumilus">
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<emphasis italics="true" pageId="0" pageNumber="1">D. pumilus</emphasis>
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</taxonomicName>
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, 2.5-3.2 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">C, D, J</emphasis>
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),
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<taxonomicName lsidName="D. quadrispinus" pageId="0" pageNumber="1" rank="species" species="quadrispinus">
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<emphasis italics="true" pageId="0" pageNumber="1">D. quadrispinus</emphasis>
|
||
</taxonomicName>
|
||
, 2.2-2.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E, F, K</emphasis>
|
||
), and
|
||
<taxonomicName lsidName="D. shoreae" pageId="0" pageNumber="1" rank="species" species="shoreae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">D. shoreae</emphasis>
|
||
</taxonomicName>
|
||
, 3.0-3.8 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">G, H, L</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |