228 lines
21 KiB
XML
228 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.71.e60921" ID-GBIF-Dataset="542339bb-d944-41a5-b45b-664a07c655b6" ID-GBIF-Taxon="191980069" ID-Pensoft-Pub="2625-8498-71-75" ID-Pensoft-UUID="C87DE9A5FDFA5B5FA5603914E35A1446" ID-ZooBank="1C61F5FE1FA44083B91313FD42157FCA" ModsDocID="2625-8498-71-75" checkinTime="1641857443470" checkinUser="pensoft" docAuthor="Quah, Evan S. H., Grismer, L. Lee, Wood Jr., Perry L., Lim, Kelvin K. P., Imbun, Paul Y. & Anuar, M. S. Shahrul" docDate="2021" docId="3AF536568CCF55129D1A09586C0F23D3" docLanguage="en" docName="VertZool 71: 75-99" docOrigin="Vertebrate Zoology 71" docPubDate="2021-03-11" docSource="http://dx.doi.org/10.3897/vz.71.e60921" docTitle="Abavorana nazgul Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin 2017" docType="treatment" docVersion="4" id="C87DE9A5FDFA5B5FA5603914E35A1446" lastPageNumber="75" masterDocId="C87DE9A5FDFA5B5FA5603914E35A1446" masterDocTitle="An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo" masterLastPageNumber="99" masterPageNumber="75" pageId="0" pageNumber="75" updateTime="1641863460135" updateUser="ExternalLinkService">
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<mods:title>An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo</mods:title>
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<mods:namePart>Quah, Evan S. H.</mods:namePart>
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<mods:affiliation>Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia, https: // orcid. org / 0000 - 0002 - 5357 - 1953 & Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698 & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
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<mods:namePart>Grismer, L. Lee</mods:namePart>
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<mods:affiliation>Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wood Jr., Perry L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3767-5274</mods:nameIdentifier>
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<mods:affiliation>Department of Biological Sciences & Museum of Natural History, Auburn University, Auburn, Alabama 36849, USA, https: // orcid. org / 0000 - 0003 - 3767 - 5274</mods:affiliation>
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<mods:namePart>Lim, Kelvin K. P.</mods:namePart>
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<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
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<mods:namePart>Imbun, Paul Y.</mods:namePart>
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<mods:affiliation>Museum of Zoology, Research & Education Division, Sabah Parks, 10626 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Anuar, M. S. Shahrul</mods:namePart>
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<mods:affiliation>School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia, https: // orcid. org / 0000 - 0003 - 0648 - 7318 & Center for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Malaysia</mods:affiliation>
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<mods:title>Vertebrate Zoology</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-03-11</mods:number>
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<mods:number>71</mods:number>
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<mods:start>75</mods:start>
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<mods:url>http://dx.doi.org/10.3897/vz.71.e60921</mods:url>
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<mods:classification>journal article</mods:classification>
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<subSection pageId="0" pageNumber="75" type="taxonomy of abavorana">
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<treatment ID-GBIF-Taxon="191980069" LSID="urn:lsid:plazi:treatment:3AF536568CCF55129D1A09586C0F23D3" httpUri="http://treatment.plazi.org/id/3AF536568CCF55129D1A09586C0F23D3" lastPageNumber="75" pageId="0" pageNumber="75">
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<subSubSection pageId="0" pageNumber="75" type="nomenclature">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName LSID="3AF53656-8CCF-5512-9D1A-09586C0F23D3" authority="Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin 2017" authorityName="Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin" authorityYear="2017" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana nazgul" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="nazgul">Abavorana nazgul Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin 2017</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Peninsular Malaysian, Thai and Sumatran Abavorana luctuosa, and A. nazgul; A: A. luctuosa from Sungai Durian Forest Reserve, Kelantan, Peninsular Malaysia. B: A. luctuosa from Janda Baik, Pahang, Peninsular Malaysia. C: A. luctuosa from Semenyih, Selangor, Peninsular Malaysia. D: A. luctuosa from Fraser's Hill, Pahang, Peninsular Malaysia. E: A. luctuosa from Hala-Bala Wildlife Sanctuary, Thailand. F: A. luctuosa from Sumatra, Indonesia. G: A. nazgul (USMHC 2572) from Gunung Jerai, Kedah, Peninsular Malaysia. H. A. cf. nazgul from Thale Ban National Park, Satun Province, Thailand. (Photographs by Zaharil Dzulkafly [A], Steven Wong [B], Kurt H. P. Guek [C], Evan Quah [D & G], Ian Dugdale [E], Hellen Kurniati [F] & Bunyarit Dechochai [H])." figureDoi="10.3897/vz.71.e60921.figure8" httpUri="https://binary.pensoft.net/fig/518981" pageId="0" pageNumber="75">Gunung Jerai Black Stream-Frog Figs 8G-H</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Colour pattern comparison among species of the genus Abavorana; Dorsum (A, C & E) and venter (B, D & F) in life of A. luctuosa (USMHC 2576) (A & B) from Penang Hill, Penang Island, Malaysia, A. decorata from Maliau Basin, Sabah, East Malaysia, Borneo (C & D) and A. nazgul (USMHC 2233) (E & F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E & F] and Stefan T. Hertwig [C & D])." figureDoi="10.3897/vz.71.e60921.figure9" httpUri="https://binary.pensoft.net/fig/518982" pageId="0" pageNumber="75">, 9E-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="reference_group">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">Rana luctuosa</taxonomicName>
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Sukumaran, J. 2005: 38
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="holotype">
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<paragraph pageId="0" pageNumber="75">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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An adult male (USMHC 2231) collected on 9 April 2016 from Gunung Jerai, Kedah, Peninsular Malaysia (
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<geoCoordinate degrees="5" direction="north" minutes="47.719" orientation="latitude" precision="1" value="5.7953167">5°47.719'N</geoCoordinate>
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;
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<geoCoordinate degrees="100" direction="east" minutes="26.222" orientation="longitude" precision="1" value="100.437035">100°26.222'E</geoCoordinate>
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, 948 m in elevation) by Evan S.H. Quah and David Chan.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
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<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin" authorityYear="2017" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana nazgul" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="nazgul">
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<emphasis italics="true" pageId="0" pageNumber="75">Abavorana nazgul</emphasis>
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</taxonomicName>
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can be differentiated from its congeners by the following combination of characters: adult males 41.51-44.1 mm SVL and single adult female 52.51 mm; adult males with SW 7.98-8.70 mm, single adult female SW 9.59 mm; adult males with IOD 3.94-4.48 mm, single adult female IOD 4.94 mm; adult males with TD 3.52-4.01 mm, single adult female TD 4.10 mm; adult males with FL 22.20-23.22 mm, single adult female FL 27.80 mm; nuptial pads absent in males; humeral glands in males small (2.4-2.98 mm) (Table
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<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Summary statistics and ventral colour pattern among members of the genus Abavorana. Summary statistics and ventral colour pattern for Abavorana decorata, A. nazgul, and A. luctuosa populations from the Malay Peninsula and Borneo, SD = standard deviation, N = sample size, / = no data. Humeral gland only present in males and all measurements in mm." httpUri="http://table.plazi.org/id/B91F734E592A25063FCA3E8C57D388D3" pageId="0" pageNumber="75" tableUuid="B91F734E592A25063FCA3E8C57D388D3">5</tableCitation>
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); dorsolateral stripe continuous, orange to yellow in colour; dorsum between dorsolateral stripes black, with or without faint orange or yellow speckles; flanks black, colouration unstratified; distinct cream-colored spots on flanks, dorsal surfaces of limbs, and upper lip; venter grey-brown with prominent light spots on throat and belly, smaller spots on underside of thigh (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Colour pattern comparison among species of the genus Abavorana; Dorsum (A, C & E) and venter (B, D & F) in life of A. luctuosa (USMHC 2576) (A & B) from Penang Hill, Penang Island, Malaysia, A. decorata from Maliau Basin, Sabah, East Malaysia, Borneo (C & D) and A. nazgul (USMHC 2233) (E & F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E & F] and Stefan T. Hertwig [C & D])." figureDoi="10.3897/vz.71.e60921.figure9" httpUri="https://binary.pensoft.net/fig/518982" pageId="0" pageNumber="75">9E-F</figureCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4320.2.4" author="Quah, ESH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="272 - 288" refId="B75" refString="Quah, ESH, Shahrul, AMS, Grismer, LL, Wood, JrPL, Siti, AMN, Muin, MA, 2017. A new species of frog of the genus Abavorana Oliver, Prendini, Kraus & Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 4320: 272 - 288, DOI: https://doi.org/10.11646/zootaxa.4320.2.4" title="A new species of frog of the genus Abavorana Oliver, Prendini, Kraus & Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia." url="https://doi.org/10.11646/zootaxa.4320.2.4" volume="4320" year="2017">Quah et al. 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="distribution">
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<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="Quah, Shahrul Anuar, Grismer, Wood, Siti Azizah & Muin" authorityYear="2017" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana nazgul" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="nazgul">
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<emphasis italics="true" pageId="0" pageNumber="75">Abavorana nazgul</emphasis>
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</taxonomicName>
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is thus far confirmed from the upper elevations of Gunung Jerai, Kedah, Peninsular Malaysia (
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4320.2.4" author="Quah, ESH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="272 - 288" refId="B75" refString="Quah, ESH, Shahrul, AMS, Grismer, LL, Wood, JrPL, Siti, AMN, Muin, MA, 2017. A new species of frog of the genus Abavorana Oliver, Prendini, Kraus & Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 4320: 272 - 288, DOI: https://doi.org/10.11646/zootaxa.4320.2.4" title="A new species of frog of the genus Abavorana Oliver, Prendini, Kraus & Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia." url="https://doi.org/10.11646/zootaxa.4320.2.4" volume="4320" year="2017">Quah et al. 2017</bibRefCitation>
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). Specimens resembling this species have been photographed at Thale Ban National Park, Satun Province in southern Thailand at approximately 500-600m in elevation around wild boar wallows but are only tentatively identified as
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<taxonomicName genus="A." lsidName="A. nazgul" pageId="0" pageNumber="75" rank="species" species="nazgul">A. cf. nazgul</taxonomicName>
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pending positive evidence (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Peninsular Malaysian, Thai and Sumatran Abavorana luctuosa, and A. nazgul; A: A. luctuosa from Sungai Durian Forest Reserve, Kelantan, Peninsular Malaysia. B: A. luctuosa from Janda Baik, Pahang, Peninsular Malaysia. C: A. luctuosa from Semenyih, Selangor, Peninsular Malaysia. D: A. luctuosa from Fraser's Hill, Pahang, Peninsular Malaysia. E: A. luctuosa from Hala-Bala Wildlife Sanctuary, Thailand. F: A. luctuosa from Sumatra, Indonesia. G: A. nazgul (USMHC 2572) from Gunung Jerai, Kedah, Peninsular Malaysia. H. A. cf. nazgul from Thale Ban National Park, Satun Province, Thailand. (Photographs by Zaharil Dzulkafly [A], Steven Wong [B], Kurt H. P. Guek [C], Evan Quah [D & G], Ian Dugdale [E], Hellen Kurniati [F] & Bunyarit Dechochai [H])." figureDoi="10.3897/vz.71.e60921.figure8" httpUri="https://binary.pensoft.net/fig/518981" pageId="0" pageNumber="75">8H</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/vz.71.e60921.figure8" httpUri="https://binary.pensoft.net/fig/518981" pageId="0" pageNumber="75" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 8.</emphasis>
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Peninsular Malaysian, Thai and Sumatran
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<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">Abavorana luctuosa</emphasis>
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</taxonomicName>
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, and
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<taxonomicName genus="A." lsidName="A. nazgul" pageId="0" pageNumber="75" rank="species" species="nazgul">
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<emphasis italics="true" pageId="0" pageNumber="75">A. nazgul</emphasis>
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</taxonomicName>
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; A:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from Sungai Durian Forest Reserve, Kelantan, Peninsular Malaysia. B:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from Janda Baik, Pahang, Peninsular Malaysia. C:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from Semenyih, Selangor, Peninsular Malaysia. D:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from
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<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
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Hill, Pahang, Peninsular Malaysia. E:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from Hala-Bala Wildlife Sanctuary, Thailand. F:
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
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<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
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</taxonomicName>
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from Sumatra, Indonesia. G:
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<taxonomicName genus="A." lsidName="A. nazgul" pageId="0" pageNumber="75" rank="species" species="nazgul">
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<emphasis italics="true" pageId="0" pageNumber="75">A. nazgul</emphasis>
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</taxonomicName>
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(USMHC 2572) from Gunung Jerai, Kedah, Peninsular Malaysia. H.
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<taxonomicName genus="A." lsidName="A. nazgul" pageId="0" pageNumber="75" rank="species" species="nazgul">A. cf. nazgul</taxonomicName>
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from Thale Ban National Park, Satun Province, Thailand. (Photographs by Zaharil Dzulkafly [A], Steven Wong [B], Kurt H.P. Guek [C], Evan Quah [D & G], Ian Dugdale [E], Hellen Kurniati [F] & Bunyarit Dechochai [H]).
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</paragraph>
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</caption>
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<caption doi="10.3897/vz.71.e60921.figure9" httpUri="https://binary.pensoft.net/fig/518982" pageId="0" pageNumber="75" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 9.</emphasis>
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Colour pattern comparison among species of the genus
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<taxonomicName authorityName="Oliver, Prendini, Kraus & Raxworthy" authorityYear="2015" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="75">Abavorana</emphasis>
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</taxonomicName>
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; Dorsum (A,C & E) and venter (B,D & F) in life of
|
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<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
|
||
</taxonomicName>
|
||
(USMHC 2576) (A & B) from Penang Hill, Penang Island, Malaysia,
|
||
<taxonomicName genus="A." lsidName="A. decorata" pageId="0" pageNumber="75" rank="species" species="decorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">A. decorata</emphasis>
|
||
</taxonomicName>
|
||
from Maliau Basin, Sabah, East Malaysia, Borneo (C & D) and
|
||
<taxonomicName genus="A." lsidName="A. nazgul" pageId="0" pageNumber="75" rank="species" species="nazgul">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">A. nazgul</emphasis>
|
||
</taxonomicName>
|
||
(USMHC 2233) (E & F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E & F] and Stefan T. Hertwig [C & D]).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/vz.71.e60921.figure10" httpUri="https://binary.pensoft.net/fig/518983" pageId="0" pageNumber="75" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="75">
|
||
<emphasis bold="true" pageId="0" pageNumber="75">Figure 10.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana decorata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
|
||
</taxonomicName>
|
||
in Borneo; Known distribution of
|
||
<taxonomicName authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana decorata</emphasis>
|
||
</taxonomicName>
|
||
(red) and
|
||
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
|
||
</taxonomicName>
|
||
(blue) in Borneo. Star = type locality.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |