treatments-xml/data/D1/77/87/D17787854F5B1F2C0DD76FAD1563FBB9.xml
2024-06-21 12:52:41 +02:00

174 lines
20 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0E2D9E84925EDDF8DB340195E85CF8D5" ID-CLB-Dataset="32713" ID-DOI="10.11646/zootaxa.4287.1.1" ID-GBIF-Dataset="91e8f5fd-1e15-4c4f-872f-3d4c50d0c8a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="829051" ID-ZooBank="13C4F702-EF00-4F04-B38E-3F0AA6CAF718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110702684" checkinUser="plazi" docAuthor="ROBERTO CALDARA, CHARLES W. OBRIEN &amp; MASSIMO MEREGALLI" docDate="2017" docId="D17787854F5B1F2C0DD76FAD1563FBB9" docLanguage="en" docName="zootaxa.4287.1.1.pdf.imf" docOrigin="Zootaxa 4287 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parabagous Schilsky 1907" docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="2D4EFFFD4F7D1F0A0D406F35171FFFD9" masterDocTitle="A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults" masterLastPageNumber="63" masterPageNumber="1" pageNumber="39" updateTime="1698465391549" updateUser="ExternalLinkService">
<mods:mods id="43D0C433A11D1415BCD34BD90A8B4D99" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F6B1C9AA34CE512B14E530C9E5CD58D0">
<mods:title id="5722A3CD423542EBADBE6238AB56E392">A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults</mods:title>
</mods:titleInfo>
<mods:name id="F8A3114E45CCA59E2D4649A27EE1CF5E" type="personal">
<mods:role id="C9249B06E3BB7FD143516E6D885CC149">
<mods:roleTerm id="5D02F869CB2CF5FF7EA2A9FB4ED5D89D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1C1882ECD4A4E6EE7B8901DCF10309A">ROBERTO CALDARA</mods:namePart>
</mods:name>
<mods:name id="959BC4761E6CE68190ADDDE8EA556A68" type="personal">
<mods:role id="F67178F1462CEC2D21C05406FC70615D">
<mods:roleTerm id="F6BF08D5596BA7540A4F1CF1A1EC7922">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB8C41B1B789003D6664C2CDFA03AA66">CHARLES W. OBRIEN</mods:namePart>
</mods:name>
<mods:name id="92830F5DAA4BB968D921C3B9F9D2901F" type="personal">
<mods:role id="03B0E49AF63B8464949006CAB8BD3C91">
<mods:roleTerm id="9E85D32A9F77B28887C1CF21F8248A96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26251D65E542C39C342F4DB88AC7F4E2">MASSIMO MEREGALLI</mods:namePart>
</mods:name>
<mods:typeOfResource id="0D29829CBCC77DCEFBD89DDAAA7DBA33">text</mods:typeOfResource>
<mods:relatedItem id="053FA6D0E1DF8B96D17CCB6BBD273ABC" type="host">
<mods:titleInfo id="436CBD86EAD1D55E14F47D6D83F00830">
<mods:title id="CDCF0EC9AA53EDFF0D97026C3252172E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="142904969153F282E519C7B4746CDE10">
<mods:date id="488E2E50B03D2EA43D5CB78EFE9DF24B">2017</mods:date>
<mods:detail id="69F30EF06DCDB1137C1E95CDDA7FE2E1" type="pubDate">
<mods:number id="B6969A0C07A6BB66AF81357C4798A39E">2017-07-05</mods:number>
</mods:detail>
<mods:detail id="4863FAF97737A33A7ECD4D78B081DA1A" type="volume">
<mods:number id="F1D4A09063A74B04F2F6C532E743C2B4">4287</mods:number>
</mods:detail>
<mods:detail id="0DF68F605FD9D27E02C18DF69621310D" type="issue">
<mods:number id="DA88AB5CD1B67FB7B68A8070943164B0">1</mods:number>
</mods:detail>
<mods:extent id="1F69014D86522B326811414C975507E7" unit="page">
<mods:start id="5AC2178A249EAAA21AD2E31F8F8BB3FA">1</mods:start>
<mods:end id="659DF4D066B8CF2629BAD708D1ADC34A">63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="88E6DB33400E8D178F58C0AB19F70F05">journal article</mods:classification>
<mods:identifier id="46F7AC7DDCDFCCC2DDCBED7359107377" type="CLB-Dataset">32713</mods:identifier>
<mods:identifier id="B2338C86B5D834B3940B824E30609B1F" type="DOI">10.11646/zootaxa.4287.1.1</mods:identifier>
<mods:identifier id="05509D40C8CAF75DDE5AEE37FCDA9E12" type="GBIF-Dataset">91e8f5fd-1e15-4c4f-872f-3d4c50d0c8a7</mods:identifier>
<mods:identifier id="56519F9865CC4F45F7F0D35451A9785F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A95EDB62E634DDFD5438637ACE553770" type="Zenodo-Dep">829051</mods:identifier>
<mods:identifier id="5CDFEFE2113D93956E2120504E4003A0" type="ZooBank">13C4F702-EF00-4F04-B38E-3F0AA6CAF718</mods:identifier>
</mods:mods>
<treatment id="D17787854F5B1F2C0DD76FAD1563FBB9" ID-DOI="http://doi.org/10.5281/zenodo.4386655" ID-GBIF-Taxon="132549332" ID-Zenodo-Dep="4386655" LSID="urn:lsid:plazi:treatment:D17787854F5B1F2C0DD76FAD1563FBB9" httpUri="http://treatment.plazi.org/id/D17787854F5B1F2C0DD76FAD1563FBB9" lastPageNumber="39" pageId="38" pageNumber="39">
<subSubSection id="11C465184F5B1F2C0DD76FAD145EFF6B" box="[151,833,151,178]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="596136934F5B1F2C0DD76FAD145EFF6B" blockId="38.[151,833,151,178]" box="[151,833,151,178]" pageId="38" pageNumber="39">
<heading id="022981FF4F5B1F2C0DD76FAD145EFF6B" bold="true" box="[151,833,151,178]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="6BAAEA814F5B1F2C0DD76FAD145EFF6B" bold="true" box="[151,833,151,178]" pageId="38" pageNumber="39">
Subgenus
<taxonomicName id="9EDE4D104F5B1F2C0C576FA21540FF6B" ID-CoL="8HNSP" authority="Schilsky, 1907" authorityName="Schilsky" authorityYear="1907" box="[279,607,151,178]" class="Insecta" family="Curculionidae" genus="Parabagous" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5B1F2C0C576FA216B7FF68" bold="true" box="[279,424,151,177]" italics="true" pageId="38" pageNumber="39">Parabagous</emphasis>
<bibRefCitation id="3D4F4B624F5B1F2C0CEF6FAD1540FF6B" author="Schilsky" box="[431,607,152,178]" pageId="38" pageNumber="39" refString="Schilsky, J. (1907) Die Kafer Europa's. Nach der Natur beschrieben von Dr. H. C. Kuster und Dr. G. Kraatz. Heft 44. Bauer und Raspe (E. Kuster), Nurnberg,. iv + A-II + 100 nr. + [2 Nachtrage und Berichtigungen] pp." type="book chapter" year="1907">Schilsky, 1907</bibRefCitation>
</taxonomicName>
resurrected name
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5B1F2C0DD76FD514B3FEB6" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="596136934F5B1F2C0DD76FD516B1FEEB" blockId="38.[151,1436,222,367]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="79CE11BD4F5B1F2C0DD76FD516B1FEEB" pageId="38" pageNumber="39">
<taxonomicName id="9EDE4D104F5B1F2C0DD76FD517FCFF2C" ID-CoL="8HCYW" box="[151,227,224,245]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5B1F2C0DD76FD517FCFF2C" box="[151,227,224,245]" italics="true" pageId="38" pageNumber="39">Bagous</emphasis>
</taxonomicName>
subgen.
<taxonomicName id="9EDE4D104F5B1F2C0C006FEB1542FF2D" ID-CoL="8HNSP" authority="Schilsky, 1907" authorityName="Schilsky" authorityYear="1907" box="[320,605,222,244]" class="Insecta" family="Curculionidae" genus="Parabagous" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5B1F2C0C006FEB16A3FF2D" box="[320,444,222,244]" italics="true" pageId="38" pageNumber="39">Parabagous</emphasis>
<bibRefCitation id="3D4F4B624F5B1F2C0C846FEB1542FF2D" author="Schilsky" box="[452,605,222,244]" pageId="38" pageNumber="39" refString="Schilsky, J. (1907) Die Kafer Europa's. Nach der Natur beschrieben von Dr. H. C. Kuster und Dr. G. Kraatz. Heft 44. Bauer und Raspe (E. Kuster), Nurnberg,. iv + A-II + 100 nr. + [2 Nachtrage und Berichtigungen] pp." type="book chapter" year="1907">Schilsky, 1907</bibRefCitation>
</taxonomicName>
: N (type species:
<taxonomicName id="9EDE4D104F5B1F2C0E566FD5136EFF2D" ID-CoL="KDNS" authority="Tournier, 1874" authorityName="Tournier" authorityYear="1874" box="[790,1137,222,245]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="6BAAEA814F5B1F2C0E566FD514D0FF2D" box="[790,975,222,245]" italics="true" pageId="38" pageNumber="39">Bagous chevrolati</emphasis>
<bibRefCitation id="3D4F4B624F5B1F2C0E966FEB136EFF2D" author="Tournier" box="[982,1137,222,244]" pageId="38" pageNumber="39" refString="Tournier, H. (1874) Materiaux pour la monographie de la tribu des Erirrhinides de la famille des Curculionides (Coleopteres). Annales de la Societe Entomologique de Belgique, 17, 63 - 116." type="journal article" year="1874">Tournier, 1874</bibRefCitation>
</taxonomicName>
).
<treatmentCitation id="D87F10824F5B1F2C09C36FEB122DFF2D" author="Klima" box="[1155,1330,222,244]" page="111" pageId="38" pageNumber="39" year="1934">
<bibRefCitation id="3D4F4B624F5B1F2C09C36FEB121FFF2D" author="Klima" box="[1155,1280,222,244]" pageId="38" pageNumber="39" refString="Klima, A. (1934) Pars 140. Curculionidae: Erirrhininae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, s' - Gravenhage, 167 pp." type="book" year="1934">Klima 1934</bibRefCitation>
: 111
</treatmentCitation>
.
<treatmentCitation id="D87F10824F5B1F2C087D6FEB162DFECA" author="Hustache" page="206" pageId="38" pageNumber="39" year="1930">
<bibRefCitation id="3D4F4B624F5B1F2C087D6FEB17E2FECA" author="Hustache" pageId="38" pageNumber="39" refString="Hustache, A. (1930) Curculionidae Gallo-Rhenans. Annales de la Societe Entomologique de France, 49, 81 - 272." type="journal article" year="1930">Hustache 1930</bibRefCitation>
: 206
</treatmentCitation>
.
<treatmentCitation id="D87F10824F5B1F2C0C7E6FC81506FECA" author="Hoffmann" box="[318,537,253,275]" page="716" pageId="38" pageNumber="39" year="1954">
<bibRefCitation id="3D4F4B624F5B1F2C0C7E6FC816FBFECA" author="Hoffmann" box="[318,484,253,275]" pageId="38" pageNumber="39" refString="Hoffmann, A. (1954) Faune de France 59 Coleopteres Curculionides (Deuxieme partie). Le Chevalier, Paris, pp. 487 - 1208." type="journal article" year="1954">Hoffmann 1954</bibRefCitation>
: 716
</treatmentCitation>
, 722.
<treatmentCitation id="D87F10824F5B1F2C0F1A6FC8140CFECA" author="Gonzalez" box="[602,787,253,275]" page="5" pageId="38" pageNumber="39" year="1971">
<bibRefCitation id="3D4F4B624F5B1F2C0F1A6FC815E6FECA" author="Gonzalez" box="[602,761,253,275]" pageId="38" pageNumber="39" refString="Gonzalez, M. (1971) Sobre el subgenero Parabagous Schilsky. Publicaciones del Instituto de Biologia Aplicada, 50, 5 - 16." type="journal article" year="1971">González 1971</bibRefCitation>
: 5
</treatmentCitation>
.
<treatmentCitation id="D87F10824F5B1F2C0E606FC81349FECA" author="Caldara" box="[800,1110,253,275]" page="137" pageId="38" pageNumber="39" year="1998">
<bibRefCitation id="3D4F4B624F5B1F2C0E606FC8133EFECA" author="Caldara" box="[800,1057,253,275]" pageId="38" pageNumber="39" refString="Caldara, R. &amp; O'Brien, C. W. (1998) Systematics and evolution of weevils of the genus Bagous. VI. Taxonomic treatment of the species of the western Palaearctic Region (Coleoptera Curculionidae). Memorie della Societa Entomologica Italiana, 76 [1997], 131 - 347." type="journal article" year="1998">Caldara &amp; O'Brien 1998</bibRefCitation>
: 137
</treatmentCitation>
.
<bibRefCitation id="3D4F4B624F5B1F2C09206FC81286FECA" author="Alonso-Zarazaga" box="[1120,1433,253,275]" pageId="38" pageNumber="39" refString="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C. (1999) A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) (Excepting Scolytidae and Platypodidae). Entomopraxis S. C. P. Edition, Barcelona, 315 pp." type="book" year="1999">Alonso-Zarazaga &amp; Lyal 1999</bibRefCitation>
. 88.
<treatmentCitation id="D87F10824F5B1F2C0DAD6E2916B4FEEB" author="Caldara" box="[237,427,284,306]" page="172" pageId="38" pageNumber="39" year="2013">
<bibRefCitation id="3D4F4B624F5B1F2C0DAD6E291667FEEB" author="Caldara" box="[237,376,284,306]" pageId="38" pageNumber="39" refString="Caldara, R. (2013) Curculionidae: Bagoinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 8. Leiden, Brill, pp. 56 - 58, 172 - 176" type="journal article" year="2013">Caldara 2013</bibRefCitation>
: 172
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="596136934F5B1F2C0DD76E0F14B3FEB6" blockId="38.[151,1436,222,367]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="79CE11BD4F5B1F2C0DD76E0F14B3FEB6" pageId="38" pageNumber="39">
<taxonomicName id="9EDE4D104F5B1F2C0DD76E0F1516FE89" authority="Hoffmann, 1962: 120" authorityName="Hoffmann" authorityPageNumber="120" authorityYear="1962" box="[151,521,314,336]" class="Insecta" family="Curculionidae" genus="Fontenelleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5B1F2C0DD76E0F1605FE89" box="[151,282,314,336]" italics="true" pageId="38" pageNumber="39">Fontenelleus</emphasis>
<treatmentCitation id="D87F10824F5B1F2C0C656E0F1516FE89" author="Hoffmann" box="[293,521,314,336]" page="120" pageId="38" pageNumber="39" year="1962">
<bibRefCitation id="3D4F4B624F5B1F2C0C656E0F16CBFE89" author="Hoffmann" box="[293,468,314,336]" pageId="38" pageNumber="39" refString="Hoffmann, A. (1962) Curculionides inedits et observations diverses concernant quelques especes de cette familie appartenant a la faune africaine. Bulletin de la Societe Entomologique de France 67, 120 - 128." type="journal article" year="1962">Hoffmann, 1962</bibRefCitation>
: 120
</treatmentCitation>
</taxonomicName>
(type species:
<taxonomicName id="9EDE4D104F5B1F2C0FF06E0E1359FE89" ID-CoL="KDPN" authority="Peyerimhoff, 1929" authorityName="Peyerimhoff" authorityYear="1929" box="[688,1094,314,336]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="cyperorum">
<emphasis id="6BAAEA814F5B1F2C0FF06E0E146BFE89" box="[688,884,315,336]" italics="true" pageId="38" pageNumber="39">Bagous cyperorum</emphasis>
Peyerimhoff, 1929
</taxonomicName>
).
<treatmentCitation id="D87F10824F5B1F2C091D6E0F1283FE89" author="Caldara" box="[1117,1436,314,336]" page="137" pageId="38" pageNumber="39" year="1998">
<bibRefCitation id="3D4F4B624F5B1F2C091D6E0F1278FE89" author="Caldara" box="[1117,1383,314,336]" pageId="38" pageNumber="39" refString="Caldara, R. &amp; O'Brien, C. W. (1998) Systematics and evolution of weevils of the genus Bagous. VI. Taxonomic treatment of the species of the western Palaearctic Region (Coleoptera Curculionidae). Memorie della Societa Entomologica Italiana, 76 [1997], 131 - 347." type="journal article" year="1998">Caldara &amp; O'Brien 1998</bibRefCitation>
: 137
</treatmentCitation>
(
<taxonomicName id="9EDE4D104F5B1F2C0D8F6E6C1659FEB6" box="[207,326,345,367]" class="Insecta" family="Curculionidae" genus="Fontenellus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5B1F2C0D8F6E6C1659FEB6" box="[207,326,345,367]" italics="true" pageId="38" pageNumber="39">Fontenellus</emphasis>
</taxonomicName>
err.).
<bibRefCitation id="3D4F4B624F5B1F2C0CC36E6C15A4FEB6" author="Alonso-Zarazaga" box="[387,699,345,367]" pageId="38" pageNumber="39" refString="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C. (1999) A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) (Excepting Scolytidae and Platypodidae). Entomopraxis S. C. P. Edition, Barcelona, 315 pp." type="book" year="1999">Alonso-Zarazaga &amp; Lyal 1999</bibRefCitation>
. 88.
<treatmentCitation id="D87F10824F5B1F2C0FAA6E6C14B7FEB6" author="Caldara" box="[746,936,345,367]" page="172" pageId="38" pageNumber="39" year="2013">
<bibRefCitation id="3D4F4B624F5B1F2C0FAA6E6C146AFEB6" author="Caldara" box="[746,885,345,367]" pageId="38" pageNumber="39" refString="Caldara, R. (2013) Curculionidae: Bagoinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 8. Leiden, Brill, pp. 56 - 58, 172 - 176" type="journal article" year="2013">Caldara 2013</bibRefCitation>
: 172
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5B1F2C0DD76EA91459FCC4" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="596136934F5B1F2C0DD76EA91459FCC4" blockId="38.[151,1437,412,1120]" pageId="38" pageNumber="39">
<emphasis id="6BAAEA814F5B1F2C0DD76EA9160EFE6C" bold="true" box="[151,273,412,437]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
Antenna with segment 7 of funicle nearly as wide as segment 6 (char. 21.0) and distinctly separated from club (except for
<taxonomicName id="9EDE4D104F5B1F2C0CD16EF416EEFE01" box="[401,497,447,472]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="belloi">
<emphasis id="6BAAEA814F5B1F2C0CD16EF416EEFE01" box="[401,497,447,472]" italics="true" pageId="38" pageNumber="39">B. belloi</emphasis>
</taxonomicName>
with 6 funicular segments). Forehead neither sulcate nor foveate (char. 24.0), with tuberculate elevation along margin of eye (excluding some species). Vestiture of odd-numbered elytral intervals generally tesselate, with alternate rectangles of dark and pale scales; setae of odd-numbered elytral intervals surrounded usually by rosette of scales of same color. Elytra lacking declivital callosities (char. 36.0). Metasternum distinctly shorter than ventrite 1. Tibiae stout, with inner margin weakly bisinuate, lacking denticles and with several moderately short bristles; outer margin of tibiae, especially on hind tibiae, straight or nearly so toward apex (in lateral view) (char. 48.0). Tarsi short, with tarsomere 3 subcordate. Basal sclerite complex with two pairs of subclavate sclerites variously articulated (char. 90.1). Sclerotization of ventral surface of penis body broadly membranous. Apodemes of spiculum ventrale very broadly and arcuately divergent from base (U-shaped) (char. 114.1). Bursa with digitate sclerotized structure (char. 116.1).
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5B1F2C0D876C121563FBB9" pageId="38" pageNumber="39" type="discussion">
<paragraph id="596136934F5B1F2C0D876C121498FBE4" blockId="38.[151,1437,412,1120]" pageId="38" pageNumber="39">
<emphasis id="6BAAEA814F5B1F2C0D876C121624FC99" bold="true" box="[199,315,807,832]" pageId="38" pageNumber="39">Remarks.</emphasis>
This subgenus was fully supported in BI and strongly supported in ML and MP (96% bs and sr). The species belonging to this subgenus share four synapomorphies: shape of funicular segment 7 (char. 21.0), features of the sclerite complex at the base of the endophallus of the penis (char. 90.1), shape of the spiculum ventrale (char. 114.1) and the sclerite complex of the bursa (char. 116.1). Some strongly modified external features (
<emphasis id="6BAAEA814F5B1F2C0DE06C8C17DCFC09" box="[160,195,953,976]" italics="true" pageId="38" pageNumber="39">i.e.</emphasis>
, short and robust legs, scale vestiture, long elytral setae) might be related to the biology of these species, which is generally poorly known. These species appear to be more strictly associated with the soil than aquatic habitats. Specimens with known data were collected either in desert or forested habitats, from lowland to high altitude, usually by sifting soil and debris far from permanent aquatic pools.
</paragraph>
<paragraph id="596136934F5B1F2C0D876B7D1563FBB9" blockId="38.[151,1437,412,1120]" box="[199,636,1095,1120]" pageId="38" pageNumber="39">We refer three groups to this subgenus.</paragraph>
</subSubSection>
</treatment>
</document>