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<document id="1216469F42B27637591FCBCCD3B28DA0" ID-DOI="10.11865/zs.2024101" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941018" ID-ZooBank="CA760FC0-4A28-4FEC-8D5F-766024ACF490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576729003" checkinUser="felipe" docAuthor="Lin, Yejie, Li, Shuqiang, Mo, Haolin &amp; Wang, Xihao" docDate="2024" docId="81114138FF8CFFEA7D9FFD89C7F6FA40" docLanguage="en" docName="ZoolSyst.49.1.4-98.pdf" docOrigin="Zoological Systematics 49 (1)" docSource="https://www.mendeley.com/catalogue/be4c04f9-10f9-351c-8fd7-9c0289bee230/" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Langlibaitiao Lin &amp; Li 2024, gen. nov." docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="7D283940FF84FFE27D17FFD2C25AFFC3" masterDocTitle="Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam" masterLastPageNumber="98" masterPageNumber="4" pageNumber="12" updateTime="1712577654997" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A5F946AA503AC6EAB4F56B5B923094B2">Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam</mods:title>
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<mods:namePart id="C32D04196CC8CCF8A8907EB65ADEAD54">Lin, Yejie</mods:namePart>
<mods:affiliation id="4582AC74D6F0AD1CCA5763A5DF3E9E50">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="72ABFA9D270197EF6B306BDC5EDFBAB9">Li, Shuqiang</mods:namePart>
<mods:affiliation id="99879F3E96B8E60FFD0E74DB42F7A292">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="852EF9D696D3F2236355F1FEF767F71B">Mo, Haolin</mods:namePart>
<mods:affiliation id="096CE81D0DB73EA70C2449F88168752D">Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, China</mods:affiliation>
</mods:name>
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<mods:namePart id="00FB0D8AA182DE5557FCE63ACF50434E">Wang, Xihao</mods:namePart>
<mods:affiliation id="7D13359BF92B71DC9CD3FE658F110A67">Jinan Licheng No. 2 High School, Jinan 250109, China Corresponding author, E-mail: lisq @ ioz. ac. cn</mods:affiliation>
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<mods:title id="FEF5E1E33CB03DFB51DC4A8B936F1621">Zoological Systematics</mods:title>
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<mods:date id="4A649BDA76054C328092BA5A435C7CDA">2024</mods:date>
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<mods:number id="9C98E06F2058E41E1C96357F72109140">49</mods:number>
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<mods:classification id="C0EFC1C4404D6212906AA6B49F3D7C1A">journal article</mods:classification>
<mods:identifier id="A4EDD855E1E3A64FBF39C30B96E91700" type="DOI">10.11865/zs.2024101</mods:identifier>
<mods:identifier id="63BDE8960249C856949456C377358336" type="ISSN">2095-6827</mods:identifier>
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<treatment id="81114138FF8CFFEA7D9FFD89C7F6FA40" ID-DOI="http://doi.org/10.5281/zenodo.10941084" ID-Zenodo-Dep="10941084" LSID="urn:lsid:plazi:treatment:81114138FF8CFFEA7D9FFD89C7F6FA40" httpUri="http://treatment.plazi.org/id/81114138FF8CFFEA7D9FFD89C7F6FA40" lastPageNumber="12" pageId="8" pageNumber="12">
<subSubSection id="41A2A3A5FF8CFFEA7D9FFD89C3A0FDB7" box="[136,506,603,628]" pageId="8" pageNumber="12" type="nomenclature">
<paragraph id="0907F02EFF8CFFEA7D9FFD89C3A0FDB7" blockId="8.[136,506,603,628]" box="[136,506,603,628]" pageId="8" pageNumber="12">
<heading id="524F4742FF8CFFEA7D9FFD89C3A0FDB7" box="[136,506,603,628]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="12" reason="2">
<emphasis id="3BCC2C3CFF8CFFEA7D9FFD89C3A0FDB7" bold="true" box="[136,506,603,628]" pageId="8" pageNumber="12">
<taxonomicName id="CEB88BADFF8CFFEA7D9FFD89C3D0FDB0" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2024" box="[136,394,603,627]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7D9FFD89C346FDB0" bold="true" box="[136,284,603,627]" italics="true" pageId="8" pageNumber="12">Langlibaitiao</emphasis>
Lin &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="20FF9147FF8CFFEA7C82FD8EC3A0FDB7" box="[405,506,604,628]" pageId="8" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="41A2A3A5FF8CFFEA7DAAFD46C7F6FA40" pageId="8" pageNumber="12" type="materials_examined">
<paragraph id="0907F02EFF8CFFEA7DAAFD46C10BFD68" blockId="8.[136,1452,659,1411]" box="[189,849,659,684]" pageId="8" pageNumber="12">
<materialsCitation id="B9D0FA73FF8CFFEA7DAAFD46C117FD68" box="[189,845,659,684]" country="China" location="Guizhou" pageId="8" pageNumber="12" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="D6034E8CFF8CFFEA7DAAFD46C2A8FD6F" box="[189,242,660,684]" pageId="8" pageNumber="12">Type</typeStatus>
species.
<taxonomicName id="CEB88BADFF8CFFEA7C44FD41C005FD68" authority="Li, 2017" authorityName="Li" authorityYear="2017" box="[339,607,659,683]" class="Arachnida" family="Dictynidae" genus="Lathys" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="inaffecta">
<emphasis id="3BCC2C3CFF8CFFEA7C44FD41C3A4FD68" box="[339,510,659,683]" italics="true" pageId="8" pageNumber="12">Lathys inaffecta</emphasis>
<bibRefCitation id="6D298DDFFF8CFFEA7F12FD41C005FD68" author="Li" box="[517,607,659,683]" pageId="8" pageNumber="12" pagination="125 - 228" refId="ref30297" refString="Li, S., Wang, X. 2017. New cave-dwelling spiders of the family Dictynidae (Arachnida, Araneae) from Guangxi and Guizhou, China. Zoological Systematics, 42 (2): 125 - 228. doi: 10.11865 / zs. 201711" type="journal article" year="2017">Li, 2017</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="71AFB0BEFF8CFFEA7FB6FD41C0BEFD68" box="[673,740,659,683]" name="China" pageId="8" pageNumber="12">China</collectingCountry>
,
<collectingRegion id="CB7C3ECCFF8CFFEA7FF9FD41C117FD68" box="[750,845,659,683]" country="China" name="Guizhou" pageId="8" pageNumber="12">Guizhou</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFD64C098FD33" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">
Etymology. The genus is named after
<taxonomicName id="CEB88BADFF8CFFEA7F73FD64C0AFFD0D" authorityName="Lin &amp; Li" authorityYear="2024" box="[612,757,694,718]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="3BCC2C3CFF8CFFEA7F73FD64C0AFFD0D" box="[612,757,694,718]" italics="true" pageId="8" pageNumber="12">Langlibaitiao</emphasis>
</taxonomicName>
, nickname for one of the 108 outlaws in the classical Chinese literature
<emphasis id="3BCC2C3CFF8CFFEA7DF8FD0AC38DFD33" box="[239,471,728,753]" italics="true" pageId="8" pageNumber="12">Outlaws of the Marsh</emphasis>
; masculine in gender.
</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFD29C6AEFCB8" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">
Diagnosis.
<taxonomicName id="CEB88BADFF8CFFEA7C21FD29C06BFCD0" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2024" box="[310,561,763,787]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7C21FD29C39DFCD0" box="[310,455,763,787]" italics="true" pageId="8" pageNumber="12">Langlibaitiao</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="3BCC2C3CFF8CFFEA7F2BFD29C0FBFCD0" bold="true" box="[572,673,763,787]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7F2BFD29C0FBFCD0" box="[572,673,763,787]" pageId="8" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="CEB88BADFF8CFFEA7E39FD29C65FFCD0" authority="Simon, 1885" authorityName="Simon" authorityYear="1885" box="[814,1029,763,787]" class="Arachnida" family="Dictynidae" genus="Lathys" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="3BCC2C3CFF8CFFEA7E39FD29C12FFCD0" box="[814,885,763,787]" italics="true" pageId="8" pageNumber="12">Lathys</emphasis>
Simon, 1885
</taxonomicName>
(see
<bibRefCitation id="6D298DDFFF8CFFEA7928FD29C740FCD0" author="Marusik" box="[1087,1306,763,787]" pageId="8" pageNumber="12" pagination="21 - 33" refId="ref30805" refString="Marusik, Y. M., Kovblyuk, M. M., Nadolny, A. A. 2009. A survey of Lathys Simon, 1884, from Crimea with resurrection of Scotolathys Simon, 1884 (Aranei: Dictynidae). Arthropoda Selecta, 18: 21 - 33." type="journal article" year="2009">
Marusik
<emphasis id="3BCC2C3CFF8CFFEA7988FD2EC68FFCD0" box="[1183,1237,763,787]" italics="true" pageId="8" pageNumber="12">et al.</emphasis>
, 2009
</bibRefCitation>
, generotype), but the male can be distinguished by the absence of a patellar apophysis, with an obvious spine, tibia with 3 obvious spines directed dorsally, ventral tibial apophysis absent, retrolateral tibial apophysis tip with a strong spine and base of conductor with a semicircular process. Females can be distinguished by the simple, slightly curved copulatory ducts.
</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFC54C148FBC5" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">Description. Male. Total length 2.00–3.01. Cephalic part strongly elevated, with several setae. Furrows indistinct, fovea reduced, longitudinally. Six eyes, PER procurved in dorsal view, ALE absent. All eyes encircled with black. Eyes subequal in size. Chelicerae with distinct lateral condyles. Leg formula 1423. Opisthosoma oval, with black anchor-shaped pattern, venter without distinct pattern. Cribellum narrow, undivided.</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFBC2C70AFBAD" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">Palp. Femur almost 3 times longer than patella. Patella with dorsal spine. Tibia with 3 spines retrolaterally. Tip of retrolateral tibial apophysis with strong spine retrolaterally. Cymbium pyriform. Conductor with anterior and posterior arms, base of conductor in the centre of the bulb. Embolus filiform, originating at approximately 10:00-o’clock-position.</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFBAAC11BFB53" blockId="8.[136,1452,659,1411]" box="[189,833,1144,1168]" pageId="8" pageNumber="12">Female. Total length 2.0–2.5. Somatic characters as in male.</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFB49C0BEFB15" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">Genitalia. Copulatory openings small, round and sclerotized. Copulatory ducts short, slightly curved, touching at midline, forming an &quot;X&quot;. Spermathecae, oval, separated.</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFB32C090FB3B" blockId="8.[136,1452,659,1411]" box="[189,714,1248,1272]" pageId="8" pageNumber="12">
Distribution.
<collectingCountry id="71AFB0BEFF8CFFEA7C5BFB32C3D6FB3B" box="[332,396,1248,1272]" name="China" pageId="8" pageNumber="12">China</collectingCountry>
(
<collectingRegion id="CB7C3ECCFF8CFFEA7C8EFB32C3A3FB3B" box="[409,505,1248,1272]" country="China" name="Guizhou" pageId="8" pageNumber="12">Guizhou</collectingRegion>
,
<collectingRegion id="CB7C3ECCFF8CFFEA7F13FB32C039FB3B" box="[516,611,1248,1272]" country="China" name="Guangxi" pageId="8" pageNumber="12">Guangxi</collectingRegion>
,
<collectingRegion id="CB7C3ECCFF8CFFEA7F7AFB32C0E5FB3B" box="[621,703,1248,1272]" country="China" name="Hainan" pageId="8" pageNumber="12">Hainan</collectingRegion>
).
</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFAD6C65AFAA3" blockId="8.[136,1452,659,1411]" pageId="8" pageNumber="12">
Notes. Because of the tibia with 3 spines retrolaterally and the X-shaped copulatory ducts, we transfer the species
<taxonomicName id="CEB88BADFF8CFFEA7873FAD1C00AFAFD" authority="Zhang, Yang &amp; Zhang, 2009" authorityName="Zhang, Yang &amp; Zhang" authorityYear="2009" class="Arachnida" family="Dictynidae" genus="Lathys" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="chishuiensis">
<emphasis id="3BCC2C3CFF8CFFEA7873FAD1C351FAFD" italics="true" pageId="8" pageNumber="12">Lathys chishuiensis</emphasis>
Zhang, Yang &amp;
<bibRefCitation id="6D298DDFFF8CFFEA7CD0FAF4C00AFAFD" author="Zhang" box="[455,592,1318,1342]" pageId="8" pageNumber="12" pagination="199 - 202" refId="ref31988" refString="Zhang, Z., Yang, Z., Zhang, Y. 2009. A new species of the genus Lathys (Araneae, Dictynidae) from China. Acta Zootaxonomica Sinica, 34: 199 - 202." type="journal article" year="2009">Zhang, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="CEB88BADFF8CFFEA7F9DFAF4C162FAFD" authorityName="Li" authorityYear="2017" box="[650,824,1318,1342]" class="Arachnida" family="Dictynidae" genus="Lathys" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="inaffecta">
<emphasis id="3BCC2C3CFF8CFFEA7F9DFAF4C162FAFD" box="[650,824,1318,1342]" italics="true" pageId="8" pageNumber="12">Lathys inaffecta</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="CEB88BADFF8CFFEA7EC4FAF4C63EFAFD" authorityName="Lin &amp; Li" authorityYear="2024" box="[979,1124,1318,1342]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="3BCC2C3CFF8CFFEA7EC4FAF4C63EFAFD" box="[979,1124,1318,1342]" italics="true" pageId="8" pageNumber="12">Langlibaitiao</emphasis>
</taxonomicName>
as
<taxonomicName id="CEB88BADFF8CFFEA7999FAF4C390FAA3" authority="(Zhang, Yang &amp; Zhang, 2009)" authorityName="Lin &amp; Li &amp; Mo &amp; Wang" authorityYear="2024" baseAuthorityName="Zhang, Yang &amp; Zhang" baseAuthorityYear="2009" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="chishuiensis" status="comb. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7999FAF4C7F6FAFD" box="[1166,1452,1318,1342]" italics="true" pageId="8" pageNumber="12">Langlibaitiao chishuiensis</emphasis>
(Zhang, Yang &amp;
<bibRefCitation id="6D298DDFFF8CFFEA7C2CFA9AC399FAA3" author="Zhang" box="[315,451,1352,1376]" pageId="8" pageNumber="12" pagination="199 - 202" refId="ref31988" refString="Zhang, Z., Yang, Z., Zhang, Y. 2009. A new species of the genus Lathys (Araneae, Dictynidae) from China. Acta Zootaxonomica Sinica, 34: 199 - 202." type="journal article" year="2009">Zhang, 2009</bibRefCitation>
)
</taxonomicName>
<emphasis id="3BCC2C3CFF8CFFEA7CC6FA9AC016FAA3" bold="true" box="[465,588,1352,1376]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7CC6FA9AC016FAA3" box="[465,588,1352,1376]" pageId="8" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CEB88BADFF8CFFEA7F68FA9AC125FAA3" authorityName="Lin &amp; Li &amp; Mo &amp; Wang" authorityYear="2024" baseAuthorityName="Li" baseAuthorityYear="2017" box="[639,895,1352,1376]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="inaffectus" status="comb. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7F68FA9AC125FAA3" box="[639,895,1352,1376]" italics="true" pageId="8" pageNumber="12">Langlibaitiao inaffectus</emphasis>
</taxonomicName>
<emphasis id="3BCC2C3CFF8CFFEA7E92FA9AC65AFAA3" bold="true" box="[901,1024,1352,1376]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7E92FA9AC65AFAA3" box="[901,1024,1352,1376]" pageId="8" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="0907F02EFF8CFFEA7DAAFAB9C7F6FA40" blockId="8.[136,1452,659,1411]" box="[189,1452,1387,1411]" pageId="8" pageNumber="12">
Species included.
<taxonomicName id="CEB88BADFF8CFFEA7C6AFAB9C0CDFA40" authorityName="Lin &amp; Li &amp; Mo &amp; Wang" authorityYear="2024" baseAuthorityName="Zhang, Yang &amp; Zhang" baseAuthorityYear="2009" box="[381,663,1387,1411]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="chishuiensis" status="comb. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7C6AFAB9C0CDFA40" box="[381,663,1387,1411]" italics="true" pageId="8" pageNumber="12">Langlibaitiao chishuiensis</emphasis>
</taxonomicName>
<emphasis id="3BCC2C3CFF8CFFEA7F8AFAB9C14DFA40" bold="true" box="[669,791,1387,1411]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7F8AFAB9C14DFA40" box="[669,791,1387,1411]" pageId="8" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="CEB88BADFF8CFFEA7E33FABEC1FDFA40" authorityName="Lin &amp; Li &amp; Mo &amp; Wang" authorityYear="2024" baseAuthorityName="Li" baseAuthorityYear="2017" box="[804,935,1387,1411]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="inaffectus" status="comb. nov.">
<emphasis id="3BCC2C3CFF8CFFEA7E33FABEC1FDFA40" box="[804,935,1387,1411]" italics="true" pageId="8" pageNumber="12">L. inaffectus</emphasis>
</taxonomicName>
<emphasis id="3BCC2C3CFF8CFFEA7EBAFAB9C67DFA40" bold="true" box="[941,1063,1387,1411]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7EBAFAB9C67DFA40" box="[941,1063,1387,1411]" pageId="8" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CEB88BADFF8CFFEA794EFABEC716FA40" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2024" box="[1113,1356,1387,1411]" class="Arachnida" family="Dictynidae" genus="Langlibaitiao" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="zhangshun" status="sp. nov.">
<emphasis id="3BCC2C3CFF8CFFEA794EFABEC6BCFA40" box="[1113,1254,1387,1411]" italics="true" pageId="8" pageNumber="12">L. zhangshun</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="3BCC2C3CFF8CFFEA7840FAB9C7F6FA40" bold="true" box="[1367,1452,1387,1411]" pageId="8" pageNumber="12">
<taxonomicNameLabel id="20FF9147FF8CFFEA7840FAB9C7F6FA40" box="[1367,1452,1387,1411]" pageId="8" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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