351 lines
47 KiB
XML
351 lines
47 KiB
XML
<document id="15352B46121BDE027963AA5BFC550F61" ID-CLB-Dataset="24900" ID-DOI="10.1093/isd/ixz015" ID-GBIF-Dataset="56172c62-3e7c-4ab7-83d4-1dd9f5c57eb5" ID-Zenodo-Dep="3542130" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573763013583" checkinUser="plazi" docAuthor="Hita-Garcia, Francisco, Lieberman, Ziv, Audisio, Tracy L., Liu, Cong & Economo, Evan P." docDate="2019" docId="03D9AC4AE574FF9FFCC4F95EBF9C0508" docLanguage="en" docName="insectSystematicsDiversity.ixz015.pdf" docOrigin="Insect Systematics and Diversity 5" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Discothyrea chimera Hita-Garcia & Lieberman & Audisio & Liu & Economo 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="37" masterDocId="FFE0D432E555FFBBFFF6FF95BB120352" masterDocTitle="Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy" masterLastPageNumber="84" masterPageNumber="1" pageNumber="34" updateTime="1698765580234" updateUser="ExternalLinkService">
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<mods:title id="72D1AE476F7E3747EAC0FF19CA0375DD">Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy</mods:title>
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<mods:namePart id="4DDAA3FEB8E0CBB3B261290AEEB1F4E7">Audisio, Tracy L.</mods:namePart>
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<mods:namePart id="067CB2042A04EB7BC39DA94B22B51A95">Liu, Cong</mods:namePart>
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<mods:title id="E977F30FAA65B243F7033190E913D2F9">Insect Systematics and Diversity</mods:title>
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<taxonomicName id="4C7066DFE574FF9AFCC4F95EBE2405B0" authority="Hita Garcia & Lieberman" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[818,1334,1738,1762]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="chimera" status="sp. nov.">
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<emphasis id="B904C14EE574FF9AFCC4F95EBF0805B0" box="[818,1050,1739,1762]" italics="true" pageId="33" pageNumber="34">Discothyrea chimera</emphasis>
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Hita Garcia & Lieberman
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(
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<figureCitation id="134B01D9E574FF9AFCCCF978B8830456" box="[826,913,1773,1796]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="33" pageNumber="34">Figs. 4E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFC69F978B8A90456" box="[927,955,1773,1796]" captionStart="Fig" captionStartId="10.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@10.[154,1434,147,1419]" captionTargetPageId="10" captionText="Fig. 6. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542144" httpUri="https://zenodo.org/record/3542144/files/figure.png" pageId="33" pageNumber="34">6E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFC3EF978B8F70456" box="[968,997,1773,1796]" captionStart="Fig" captionStartId="11.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@11.[152,1435,147,1422]" captionTargetPageId="11" captionText="Fig. 7. Diagnostic plate showing still images from surface volume renderings of the head in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542148" httpUri="https://zenodo.org/record/3542148/files/figure.png" pageId="33" pageNumber="34">7E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFC04F978BF1C0456" box="[1010,1038,1773,1796]" captionStart="Fig" captionStartId="12.[146,172,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@12.[152,1435,147,1422]" captionTargetPageId="12" captionText="Fig. 8. Diagnostic plate showing still images from surface volume renderings of the right mandible in dorsal view (the remainder of the body virtually removed).All Afrotropical species are shown,species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542150" httpUri="https://zenodo.org/record/3542150/files/figure.png" pageId="33" pageNumber="34">8E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFBEDF978BF250456" box="[1051,1079,1773,1796]" captionStart="Fig" captionStartId="13.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@13.[152,1435,147,1422]" captionTargetPageId="13" captionText="Fig. 9. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542152" httpUri="https://zenodo.org/record/3542152/files/figure.png" pageId="33" pageNumber="34">9E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFBB2F978BF7E0456" box="[1092,1132,1773,1796]" captionStart="Fig" captionStartId="14.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@14.[152,1435,147,1422]" captionTargetPageId="14" captionText="Fig. 10. Diagnostic plate showing still images from surface volume renderings of the body in dorsal view with focus on mesosoma.All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542154" httpUri="https://zenodo.org/record/3542154/files/figure.png" pageId="33" pageNumber="34">10E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFB8CF978BF8F0456" box="[1146,1181,1773,1796]" captionStart="Fig" captionStartId="15.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@15.[154,1434,147,1419]" captionTargetPageId="15" captionText="Fig. 11. Diagnostic plate showing still images from surface volume renderings of the posterior propodeum in posterior view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542156" httpUri="https://zenodo.org/record/3542156/files/figure.png" pageId="33" pageNumber="34">11E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFB5BF978BFC40456" box="[1197,1238,1773,1796]" captionStart="Fig" captionStartId="16.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@16.[154,1434,147,1419]" captionTargetPageId="16" captionText="Fig. 12. Diagnostic plate showing still images from surface volume renderings of the petiole and gaster in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542158" httpUri="https://zenodo.org/record/3542158/files/figure.png" pageId="33" pageNumber="34">12E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFB12F978BE190456" box="[1252,1291,1773,1796]" captionStart="Fig" captionStartId="18.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1419]" captionTargetId="figure@18.[154,1434,147,1419]" captionTargetPageId="18" captionText="Fig. 14. Diagnostic plate showing differences in body pilosity. (A) D. mixta (CASENT0235473),(B) D. oculata (CASENT0235467),(C) D. aisnetu (CASENT0235475), (D) D.athene (CASENT0235476),(E) D.chimera (CASENT0235471),(F) D.damato (CASENT0247362),(G) D.dryad (CASENT0247371),(H) D.gaia (CASENT0247040), (I) D. gryphon (CASENT0247367), (J) D. hawkesi (CASENT0235470), (K) D. kalypso (CASENT0235468), (L) D. maia (CASENT0790541), (M) D. michelae (CASENT0235469), (N) D. patrizii (CASENT0235472), (O) D. penthos (CASENT0247383), (P) D. poweri (SAM-ENT-0011509), (Q) D. schulzei (CASENT0247370), (R) D. traegaordhi (CASENT0790122), (S) D. venus (CASENT0247017), (T) D. wakanda (CASENT0790326).With the exception of F, L, R,T, all other images are from https://www.antweb.org—photographers Michele Esposito and Will Ericson." figureDoi="http://doi.org/10.5281/zenodo.3542162" httpUri="https://zenodo.org/record/3542162/files/figure.png" pageId="33" pageNumber="34">14E</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFAECF978BE220456" box="[1306,1328,1773,1796]" captionStart="Fig" captionStartId="34.[146,173,1061,1078]" captionTargetBox="[147,768,147,1026]" captionTargetId="figure@34.[147,768,147,1026]" captionTargetPageId="34" captionText="Fig. 27. Stacked digital color images of D. chimera sp. n. holotype (CASENT0235471- from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.3542188" httpUri="https://zenodo.org/record/3542188/files/figure.png" pageId="33" pageNumber="34">27</figureCitation>
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,
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<figureCitation id="134B01D9E574FF9AFACBF978BE4A0456" box="[1341,1368,1773,1796]" captionStart="Fig" captionStartId="35.[146,173,1782,1799]" captionTargetBox="[202,1386,147,1747]" captionTargetId="figure@35.[202,1386,147,1747]" captionTargetPageId="35" captionText="Fig. 28. Still images from shaded surface display volume renderings of D. chimera sp. n. holotype (CASENT0235471) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3542190" httpUri="https://zenodo.org/record/3542190/files/figure.png" pageId="33" pageNumber="34">28</figureCitation>
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; Supp Video S5 [online only])
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<emphasis id="B904C14EE574FF9AFCC4F8A5B8A60411" bold="true" box="[818,948,1840,1859]" pageId="33" pageNumber="34">Type Material</emphasis>
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</paragraph>
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<materialsCitation id="3B181701E574FF9AFCC4F8D8BEB004C9" ID-GBIF-Occurrence="2451560188" country="Tanzania" elevation="1989" latitude="-6.3753" location="Mamiwa-Kisara Forest Reserve" longLatPrecision="5" longitude="36.93711" municipality="Kilosa" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Morogoro" typeStatus="holotype">
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<typeStatus id="54CBA3FEE574FF9AFCC4F8D8B8B90432" box="[818,939,1869,1888]" pageId="33" pageNumber="34" type="holotype">HOLOTYPE</typeStatus>
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, pinned worker,
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<collectingCountry id="F3675DCCE574FF9AFB94F8DBBFCB0433" box="[1122,1241,1870,1889]" name="Tanzania" pageId="33" pageNumber="34">TANZANIA</collectingCountry>
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,
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<collectingRegion id="49B4D3BEE574FF9AFB1AF8DBBE8C0432" box="[1260,1438,1869,1889]" country="Tanzania" name="Morogoro" pageId="33" pageNumber="34">Morogoro Region</collectingRegion>
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,
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<collectingMunicipality id="6BAB8726E574FF9AFCC4F8FEB87C042C" box="[818,878,1899,1918]" pageId="33" pageNumber="34">Kilosa</collectingMunicipality>
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,
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<location id="8EAF4B87E574FF9AFC8CF8FEBFB1042C" LSID="urn:lsid:plazi:treatment:03D9AC4AE574FF9FFCC4F95EBF9C0508:8EAF4B87E574FF9AFC8CF8FEBFB1042C" box="[890,1187,1899,1918]" country="Tanzania" latitude="-6.3753" longLatPrecision="5" longitude="36.93711" municipality="Kilosa" name="Mamiwa-Kisara Forest Reserve" pageId="33" pageNumber="34" stateProvince="Morogoro">Mamiwa-Kisara Forest Reserve</location>
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,
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<quantity id="4C88B0B9E574FF9AFA84F8FEB85704CE" metricMagnitude="3" metricUnit="m" metricValue="1.989" pageId="33" pageNumber="34" unit="m" value="1989.0">
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<elevation id="005DFA6FE574FF9AFA84F8FEB85704CE" metricMagnitude="3" metricUnit="m" metricValue="1.989" pageId="33" pageNumber="34" unit="m" value="1989.0">1989 m</elevation>
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, primary forest, leaf litter, Winkler, collection code CEPF-TZ-1.2,
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|
||
<emphasis id="B904C14EE577FF99FF64FBB0BBC60764" bold="true" box="[146,212,1061,1078]" pageId="34" pageNumber="35">Fig. 27.</emphasis>
|
||
Stacked digital color images of
|
||
<taxonomicName id="4C7066DFE577FF99FDFEFBB0B97B0764" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[520,617,1061,1078]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="chimera" status="sp. nov.">
|
||
<emphasis id="B904C14EE577FF99FDFEFBB0B97B0764" box="[520,617,1061,1078]" italics="true" pageId="34" pageNumber="35">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E577FF99FD81FBB0B9BF0764" box="[631,685,1061,1078]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
||
holotype (CASENT0235471- from https://www.antweb.org, photographer Will Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF1D5CE577FF99FF64FB1ABA5B07ED" blockId="34.[146,771,1167,1215]" pageId="34" pageNumber="35">
|
||
<date id="FFCE3B9CE577FF99FF64FB1ABA2007F0" box="[146,306,1167,1186]" pageId="34" pageNumber="35" value="2005-08-16" valueMax="2005-08-21" valueMin="2005-08-16">16.–21.VIII.2005</date>
|
||
(
|
||
<emphasis id="B904C14EE577FF99FEB5FB05B9B707F0" box="[323,677,1167,1186]" italics="true" pageId="34" pageNumber="35">P. Hawkes, J. Makwati & R. Mtana</emphasis>
|
||
) (
|
||
<collectionCode id="ED618599E577FF99FD4BFB1AB9ED07F0" box="[701,767,1167,1186]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="34" pageNumber="35" type="Museum">SAMC</collectionCode>
|
||
: CASENT0235471).
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FF64FB7FBA440557" blockId="34.[146,770,1258,1541]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FF64FB7FBBEA07AF" box="[146,248,1258,1277]" italics="true" pageId="34" pageNumber="35">Cybertype.</emphasis>
|
||
Volumetric raw data (in DICOM format), 3D rotation video, still images of surface volume rendering, and 3D surface (in PLY format) of the physical
|
||
<typeStatus id="54CBA3FEE577FF99FE55FAB1BAE60665" box="[419,500,1316,1335]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
(CASENT0235471) in addition to stacked digital color images illustrating head in full-face view, profile and dorsal views of the body. The data are deposited at Dryad (Hita Garcia et al. 2019, http://doi.org/10.5061/ dryad.3qm4183) and can be freely accessed as virtual representation of the type. In addition to the cybertype data at Dryad, we also provide a freely accessible 3D surface model of the
|
||
<typeStatus id="54CBA3FEE577FF99FD6EFA41B9FB06B5" box="[664,745,1492,1511]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
at Sketchfab (Model 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E577FF99FF64F9BAB90B047F" pageId="34" pageNumber="35" type="diagnosis">
|
||
<paragraph id="8BCF1D5CE577FF99FF64F9BABBFE0510" blockId="34.[146,770,1583,1837]" box="[146,236,1583,1602]" pageId="34" pageNumber="35">
|
||
<heading id="D087AA30E577FF99FF64F9BABBFE0510" bold="true" box="[146,236,1583,1602]" fontSize="8" level="3" pageId="34" pageNumber="35" reason="6">
|
||
<emphasis id="B904C14EE577FF99FF64F9BABBFE0510" bold="true" box="[146,236,1583,1602]" pageId="34" pageNumber="35">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FF64F9D9B90B047F" blockId="34.[146,770,1583,1837]" pageId="34" pageNumber="35">
|
||
The following character combination distinguishes
|
||
<taxonomicName id="4C7066DFE577FF99FD9FF9D8B9DD050D" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[617,719,1612,1631]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE577FF99FD9FF9D8B9DD050D" box="[617,719,1612,1631]" italics="true" pageId="34" pageNumber="35">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
from the remainder of the complex: masticatory margin of mandible with a conspicuous, curved subapical tooth; subquadrate head with anterolateral genal angles sharply defined; anterior clypeal margin with long erect setae; propodeum not dentate; mesotibia without apicoventral spur; petiolar node strongly attenuated dorsally and squamiform in profile (LPeI 329); AT4 around 1.2 times longer than AT3; erect setae absent on dorsal surfaces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E577FF9FFF64F8CFB920060B" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="8BCF1D5CE577FF99FF64F8CFB907043F" blockId="34.[146,770,1882,1960]" box="[146,533,1882,1901]" pageId="34" pageNumber="35">
|
||
<heading id="D087AA30E577FF99FF64F8CFB907043F" bold="true" box="[146,533,1882,1901]" fontSize="8" level="3" pageId="34" pageNumber="35" reason="6">
|
||
<emphasis id="B904C14EE577FF99FF64F8CFB907043F" bold="true" box="[146,533,1882,1901]" pageId="34" pageNumber="35">
|
||
Worker Measurements and Indices (
|
||
<emphasis id="B904C14EE577FF99FE2BF8CFBAFB043F" bold="true" box="[477,489,1882,1901]" italics="true" pageId="34" pageNumber="35">n</emphasis>
|
||
= 1)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FF64F8E2B81004D8" blockId="34.[146,770,1882,1960]" box="[146,770,1911,1930]" pageId="34" pageNumber="35">
|
||
<heading id="D087AA30E577FF99FF64F8E2B81004D8" allCaps="true" box="[146,770,1911,1930]" centered="true" fontSize="8" level="4" pageId="34" pageNumber="35" reason="5">
|
||
EL 0.03; HL 0.45; HW 0.35; SL 0.23;
|
||
<collectingCountry id="F3675DCCE577FF99FDF8F8E2B93E04D8" box="[526,556,1911,1930]" name="Philippines" pageId="34" pageNumber="35">PH</collectingCountry>
|
||
0.19; PW 0.31; DML
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FF64F800B81004FA" blockId="34.[146,770,1882,1960]" box="[146,770,1940,1960]" pageId="34" pageNumber="35">0.33; PrH 0.34; WL 0.54; HFL 0.31; PeL 0.07; PeW 0.20; PeH 0.23;</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCC4FF03BEB003FB" blockId="34.[818,1442,149,198]" box="[818,1442,149,169]" pageId="34" pageNumber="35">
|
||
<heading id="D087AA30E577FF99FCC4FF03BEB003FB" allCaps="true" box="[818,1442,149,169]" centered="true" fontSize="8" level="4" pageId="34" pageNumber="35" reason="5">LT3 0.32; LT4 0.39; OI 6; CI 90; SI 60; LMI 52; DMI 57; DMI2 93;</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCC4FF26BF810394" blockId="34.[818,1442,149,198]" box="[818,1171,179,198]" pageId="34" pageNumber="35">ASI 124; HFI 57; DPeI 286; LPeI 329.</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCC4FE9EB8F5024C" blockId="34.[818,1444,267,1958]" box="[818,999,267,286]" pageId="34" pageNumber="35">
|
||
<heading id="D087AA30E577FF99FCC4FE9EB8F5024C" bold="true" box="[818,999,267,286]" fontSize="8" level="3" pageId="34" pageNumber="35" reason="6">
|
||
<emphasis id="B904C14EE577FF99FCC4FE9EB8F5024C" bold="true" box="[818,999,267,286]" pageId="34" pageNumber="35">Worker Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCC4FEBDBEB600F1" blockId="34.[818,1444,267,1958]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCC4FEBDB8710269" box="[818,867,296,315]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
|
||
very broad (CI 90), appearing subquadrate posterad antennal sockets posterior head margin straight; posterodorsal corners of head rounded; in frontal view, sides of head slightly convex; eyes present, relatively long (OI 6), oval, situated slightly less than halfway between anterolateral corner of gena and posterior head margin; eyes just visible in frontal view; anterolateral corner of gena squared, nearly right-angled; frontal lamella bladelike in profile, apex acute, lamella more or less evenly translucent across its disc, without a clearly defined fenestra; medial clypeus broad and sharply transverse, lateral clypeus curving moderately strongly between antennal sockets and anterolateral corners of head, bearing row of long erect setae.
|
||
<emphasis id="B904C14EE577FF99FCC4FDFEB86D012C" box="[818,895,619,638]" italics="true" pageId="34" pageNumber="35">Antenna</emphasis>
|
||
with moderately long scape (SI 60), scape very strongly incrassate, gently bent; pedicel subcylindrical, longer than broad; apparent antennomere count eight, flagellomeres basad apical club highly compressed, taken together only about as long as apical club.
|
||
<emphasis id="B904C14EE577FF99FCC4FD75B8B801A1" box="[818,938,736,755]" italics="true" pageId="34" pageNumber="35">Ventral head</emphasis>
|
||
with short horizontal postoccipital ridge with very short, triangular anteromedian carina; medial region of hypostooma somewhat truncate, arms narrowed; palpal formula not examined.
|
||
<emphasis id="B904C14EE577FF99FCC4FCADB89B0019" box="[818,905,824,843]" italics="true" pageId="34" pageNumber="35">Mandible</emphasis>
|
||
with prominent, curved subapical tooth and small, sharp prebasal denticle; prebasal denticle located nearly at basal angle; ectal face with longitudinal carina running from preapical tooth to basal angle, leaving short depressed prebasal region on masticatory margin.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCA4FC3BBF8907F9" blockId="34.[818,1444,267,1958]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCA4FC3BB8A60092" box="[850,948,942,960]" italics="true" pageId="34" pageNumber="35">Mesosoma</emphasis>
|
||
gently convex in profile, pronotum distinctly higher than propodeum; in dorsal view mesosoma moderately thick (DMI 57; DMI2 93) and somewhat narrowed posteriorly; pronotal humeri moderately rounded; posterior propodeal margin weakly concave; posterodorsal corners of propodeum subangulate, without teeth; declivitous face of propodeum very shallowly concave in profile and oblique posterior view; propodeal spiracle fairly large and round but inconspicuous, directed posteriorly; propodeal lobes poorly developed, very short and bluntly truncated.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCA4FB23B8A90672" blockId="34.[818,1444,267,1958]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCA4FB23B86F079A" box="[850,893,1206,1224]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
|
||
short (HFI 57) and relatively robust; mesotibia without apicoventral spur, with small but distinct seta inserted in apical pit; mesobasitarsus relatively short, about as long as tarsomeres II–IV taken together.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCA4FABEBE1A0531" blockId="34.[818,1444,267,1958]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCA4FABEB8DD066C" box="[850,975,1323,1342]" italics="true" pageId="34" pageNumber="35">Petiolar node</emphasis>
|
||
strongly attenuated dorsally and squamiform in profile, about 3.3 times higher than long (LPeI 329); in profile anterior face of node sloping posterodorsally, apex peaked, posterior face sloping posteroventrally; in dorsal view petiole subrectangular, about 2.9 times broader than long (DPeI 286), sides very slightly convex; in anterior view, petiolar outline sharply pentagonal and strongly, sharply peaked, sides and angles well-defined, in oblique anterior view, anterior face flat; in ventral view roughly rectangular, sides weakly diverging posteriorly; subpetiolar process short, lobate, apex rounded, bearing several long, straight, white setae; petiolar spiracles very large and subcircular in ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF99FCA4F9F8BFBF0439" blockId="34.[818,1444,267,1958]" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCA4F9F8BF3005D3" box="[850,1058,1645,1665]" italics="true" pageId="34" pageNumber="35">Abdominal segment 3</emphasis>
|
||
almost thickly cylindrical, only slightly widening posteriorly, tergite weakly peaked just anterad end of segment; sternite evenly and quite shallowly curved in profile;
|
||
<collectingCountry id="F3675DCCE577FF99FAFFF93DBE3705E9" box="[1289,1317,1704,1723]" name="American Samoa" pageId="34" pageNumber="35">AS</collectingCountry>
|
||
3 without carinate prora, but still with anterior face distinctly depressed and anterior margin of ventral face fairly deeply arcuate in ventral view;AT4 1.2 times longer than AT3 (ASI 124); AT4 large, bulbous, hemidemispherical;
|
||
<collectingCountry id="F3675DCCE577FF99FCC4F88BB85E0463" box="[818,844,1822,1841]" name="American Samoa" pageId="34" pageNumber="35">AS</collectingCountry>
|
||
4 with broad, prominent anterior lip, overlapping most of the width of
|
||
<collectingCountry id="F3675DCCE577FF99FCBCF8AEB875041C" box="[842,871,1851,1870]" name="American Samoa" pageId="34" pageNumber="35">AS</collectingCountry>
|
||
3, anterior margin straight in ventral view; successive abdominal segments short, telescopic, often concealed.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE577FF9FFCA4F8E0BA0701BC" blockId="34.[818,1444,267,1958]" lastBlockId="36.[146,772,657,750]" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
|
||
<emphasis id="B904C14EE577FF99FCA4F8E0B8BB04DA" box="[850,937,1909,1928]" italics="true" pageId="34" pageNumber="35">Sculpture</emphasis>
|
||
on gena and ventral head surface foveolate-reticulate, becoming punctate toward front of head, punctae smaller on frontal
|
||
<emphasis id="B904C14EE571FF9FFF64FD04BBCA01F0" bold="true" box="[146,216,657,674]" pageId="36" pageNumber="37">Model 5.</emphasis>
|
||
3D surface model of
|
||
<taxonomicName id="4C7066DFE571FF9FFE7AFD04BAF701F0" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[396,485,657,674]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera" status="sp. nov.">
|
||
<emphasis id="B904C14EE571FF9FFE7AFD04BAF701F0" box="[396,485,657,674]" italics="true" pageId="36" pageNumber="37">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E571FF9FFE1FFD04B90501F0" box="[489,535,657,674]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
|
||
<typeStatus id="54CBA3FEE571FF9FFDEDFD04B97101F0" box="[539,611,657,674]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
(CASENT0235471). An interactive version of this model is available in the HTML version of this article online and at https://sketchfab.com/3d-models/a9c567dc04c44205a6f 5dee0ea
|
||
<date id="FFCE3B9CE571FF9FFF25FD48BA1D01BC" box="[211,271,733,750]" pageId="36" pageNumber="37" value="18-08-05">080518</date>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0F4DD4E576FF98FF64F963BAE4043F" ID-DOI="http://doi.org/10.5281/zenodo.3542190" ID-Zenodo-Dep="3542190" httpUri="https://zenodo.org/record/3542190/files/figure.png" pageId="35" pageNumber="36" startId="35.[146,173,1782,1799]" targetBox="[202,1386,147,1747]" targetPageId="35">
|
||
<paragraph id="8BCF1D5CE576FF98FF64F963BAE4043F" blockId="35.[146,1444,1782,1901]" pageId="35" pageNumber="36">
|
||
<emphasis id="B904C14EE576FF98FF64F963BBDC0455" bold="true" box="[146,206,1782,1799]" pageId="35" pageNumber="36">Fig. 28.</emphasis>
|
||
Still images from shaded surface display volume renderings of
|
||
<taxonomicName id="4C7066DFE576FF98FD2BF963B8240455" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[733,822,1782,1799]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="chimera" status="sp. nov.">
|
||
<emphasis id="B904C14EE576FF98FD2BF963B8240455" box="[733,822,1782,1799]" italics="true" pageId="35" pageNumber="36">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E576FF98FCCCF963B87A0455" box="[826,872,1782,1799]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
|
||
holotype (CASENT0235471) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF1D5CE571FF9FFF64FC8FB97A00ED" blockId="36.[146,770,793,1370]" pageId="36" pageNumber="37">area posterad frontal lamella; mandible rather roughly sculptured with dense piligerous punctae; mesosomal dorsum, petiole, and abdominal segment 3 shallowly, evenly punctate-reticulate; lateral mesosoma irregularly punctate to rugulose on ventrolateral and posteroventral portions of propodeum; AT4 distinctly shinier and smoother than AT3, with fine piligerous punctulae.</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFF44FC5CB9F5066E" blockId="36.[146,770,793,1370]" pageId="36" pageNumber="37">
|
||
<emphasis id="B904C14EE571FF9FFF44FC5CBBEF008E" box="[178,253,969,988]" italics="true" pageId="36" pageNumber="37">Setation</emphasis>
|
||
on head, antenna, and mesosoma a fairly consistent but short, velvety layer of appressed white pubescence, best developed on mesosomal dorsum; front of head just posterad frontal lamella with longer, more distinct setae directed towards midline; longer hairs, some decumbent, present on gena, best seen in frontal view; ectal face of mandible with numerous long, curved, appressed to suberect setae; masticatory margin without row of straight setae; ventral margin of petiolar sternite with several long suberect hairs; abdominal segment 3 densely clothed with long, appressed pilosity, obscuring sculpture; noticeably longer on sternite than tergite; AT4 with long, appressed pilosity, denser than on mesoma but less so than on AT3; successive abdominal segments with dense, white, standing pilosity; legs densely and more or less evenly pubescent.
|
||
</paragraph>
|
||
<caption id="DF0F4DD4E571FF9FFCC4FBB0B8B7073B" ID-DOI="http://doi.org/10.5281/zenodo.3542192" ID-Zenodo-Dep="3542192" httpUri="https://zenodo.org/record/3542192/files/figure.png" pageId="36" pageNumber="37" startId="36.[818,845,1061,1078]" targetBox="[819,1440,147,1026]" targetPageId="36">
|
||
<paragraph id="8BCF1D5CE571FF9FFCC4FBB0B8B7073B" blockId="36.[818,1441,1061,1129]" pageId="36" pageNumber="37">
|
||
<emphasis id="B904C14EE571FF9FFCC4FBB0B8640764" bold="true" box="[818,886,1061,1078]" pageId="36" pageNumber="37">Fig. 29.</emphasis>
|
||
Stacked digital color images of
|
||
<taxonomicName id="4C7066DFE571FF9FFB5DFBB0BE1B0764" authority="Hita-Garcia & Lieberman & Audisio & Liu & Economo, 2019" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[1195,1289,1061,1078]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="damato" status="sp. nov.">
|
||
<emphasis id="B904C14EE571FF9FFB5DFBB0BE1B0764" box="[1195,1289,1061,1078]" italics="true" pageId="36" pageNumber="37">D. damato</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377C35E571FF9FFAE0FBB0BE5F0764" box="[1302,1357,1061,1078]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
|
||
holotype (CASENT0247362). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF1D5CE571FF9FFF44FAD3B920060B" blockId="36.[146,770,793,1370]" box="[178,562,1350,1370]" pageId="36" pageNumber="37">
|
||
<emphasis id="B904C14EE571FF9FFF44FAD3BBF5060B" box="[178,231,1350,1369]" italics="true" pageId="36" pageNumber="37">Color</emphasis>
|
||
testaceous, appendages dull yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E571FF9FFF64FA1CBBFD0540" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="8BCF1D5CE571FF9FFF64FA1CBBE706CE" blockId="36.[146,770,1417,1554]" box="[146,245,1417,1436]" pageId="36" pageNumber="37">
|
||
<heading id="D087AA30E571FF9FFF64FA1CBBE706CE" bold="true" box="[146,245,1417,1436]" fontSize="8" level="3" pageId="36" pageNumber="37" reason="6">
|
||
<emphasis id="B904C14EE571FF9FFF64FA1CBBE706CE" bold="true" box="[146,245,1417,1436]" pageId="36" pageNumber="37">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFF64FA33BBFD0540" blockId="36.[146,770,1417,1554]" pageId="36" pageNumber="37">The chimera was a mythological monster whose body was an amalgam of various creatures. The species is named for its singular combination of characters. The specific epithet is given as an appositive noun.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E571FF9FFF64F9D1BAF9059E" pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph id="8BCF1D5CE571FF9FFF64F9D1BA6A0505" blockId="36.[146,770,1604,1741]" box="[146,376,1604,1623]" pageId="36" pageNumber="37">
|
||
<heading id="D087AA30E571FF9FFF64F9D1BA6A0505" bold="true" box="[146,376,1604,1623]" fontSize="8" level="3" pageId="36" pageNumber="37" reason="6">
|
||
<emphasis id="B904C14EE571FF9FFF64F9D1BA6A0505" bold="true" box="[146,376,1604,1623]" pageId="36" pageNumber="37">Distribution and Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFF64F9F4BAF9059E" blockId="36.[146,770,1604,1741]" pageId="36" pageNumber="37">
|
||
Presently the new species is known only from a single specimen collected from leaf litter in the Mamiwa-Kisara Forest Reserve in
|
||
<collectingCountry id="F3675DCCE571FF9FFF64F909BBF705FD" box="[146,229,1692,1711]" name="Tanzania" pageId="36" pageNumber="37">Tanzania</collectingCountry>
|
||
(
|
||
<figureCitation id="134B01D9E571FF9FFF02F909BA2A05FD" box="[244,312,1692,1711]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="36" pageNumber="37">Fig. 4E</figureCitation>
|
||
), which is a humid montane forest.
|
||
<taxonomicName id="4C7066DFE571FF9FFD65F909BBC8059E" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE571FF9FFD65F909BBC8059E" italics="true" pageId="36" pageNumber="37">Discothyrea chimera</emphasis>
|
||
</taxonomicName>
|
||
was collected from leaf litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4ED7E571FF9FFF64F969BF9C0508" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="8BCF1D5CE571FF9FFF64F969BBE4045D" blockId="36.[146,770,1788,1954]" box="[146,246,1788,1807]" pageId="36" pageNumber="37">
|
||
<heading id="D087AA30E571FF9FFF64F969BBE4045D" bold="true" box="[146,246,1788,1807]" fontSize="8" level="3" pageId="36" pageNumber="37" reason="6">
|
||
<emphasis id="B904C14EE571FF9FFF64F969BBE4045D" bold="true" box="[146,246,1788,1807]" pageId="36" pageNumber="37">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFF64F88FB88C0685" blockId="36.[146,770,1788,1954]" lastBlockId="36.[818,1442,1183,1495]" pageId="36" pageNumber="37">
|
||
<collectingRegion id="49B4D3BEE571FF9FFF64F88FBBA5047F" box="[146,183,1818,1837]" country="Cameroon" name="North" pageId="36" pageNumber="37">Not</collectingRegion>
|
||
considering the other characters given in the diagnosis above, the presence of a conspicuous subapical mandibular tooth distinguishes
|
||
<taxonomicName id="4C7066DFE571FF9FFF2FF8C1BA2C0435" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[217,318,1876,1895]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE571FF9FFF2FF8C1BA2C0435" box="[217,318,1876,1895]" italics="true" pageId="36" pageNumber="37">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
from all other Afrotropical species. While a small and acute subapical denticle is present in
|
||
<emphasis id="B904C14EE571FF9FFDE6F8E7B99304D6" box="[528,641,1905,1924]" italics="true" pageId="36" pageNumber="37">
|
||
<taxonomicName id="4C7066DFE571FF9FFDE6F8E7B96C04D6" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[528,638,1905,1924]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gryphon">D. gryphon</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
it is not comparable to the tooth seen in
|
||
<taxonomicName id="4C7066DFE571FF9FFE6BF81AB91104F3" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[413,515,1934,1953]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE571FF9FFE6BF81AB91104F3" box="[413,515,1934,1953]" italics="true" pageId="36" pageNumber="37">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, both species appear morphologically closer to each other than to the remainder of the Afrotropical fauna, but can be well separated by eye size, the shape of the petiole, and pilosity.
|
||
<taxonomicName id="4C7066DFE571FF9FFB90FB4FBE3007BF" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[1126,1314,1242,1261]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE571FF9FFB90FB4FBE3007BF" box="[1126,1314,1242,1261]" italics="true" pageId="36" pageNumber="37">Discothyrea chimera</emphasis>
|
||
</taxonomicName>
|
||
has relatively larger eyes (OI 6), a dorsally much thinner petiole, and lacks erect setae on its dorsal surfaces, whereas
|
||
<taxonomicName id="4C7066DFE571FF9FFB7FFA80BFE60675" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[1161,1268,1300,1319]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE571FF9FFB7FFA80BFE60675" box="[1161,1268,1300,1319]" italics="true" pageId="36" pageNumber="37">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
has either no eyes or smaller eyes (OI 0–4), has a dorsally thicker petiole, and conspicuous standing setae through its body. Nevertheless, the similarity of
|
||
<taxonomicName id="4C7066DFE571FF9FFCBDFAF8B8A2062D" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[843,944,1388,1407]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="chimera">
|
||
<emphasis id="B904C14EE571FF9FFCBDFAF8B8A2062D" box="[843,944,1388,1407]" italics="true" pageId="36" pageNumber="37">D. chimera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7066DFE571FF9FFC16FAF8BF58062D" authorityName="Hita-Garcia & Lieberman & Audisio & Liu & Economo" authorityYear="2019" box="[992,1098,1388,1407]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gryphon">
|
||
<emphasis id="B904C14EE571FF9FFC16FAF8BF58062D" box="[992,1098,1388,1407]" italics="true" pageId="36" pageNumber="37">D. gryphon</emphasis>
|
||
</taxonomicName>
|
||
regarding the characters of the head, as well as in general habitus, suggest a possible close phylogenetic relationship, but, without molecular phylogenetic data, this is rather speculative.
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFCC4F999B89A054D" blockId="36.[818,1442,1548,1626]" box="[818,904,1548,1567]" pageId="36" pageNumber="37">
|
||
<heading id="D087AA30E571FF9FFCC4F999B89A054D" bold="true" box="[818,904,1548,1567]" fontSize="8" level="3" pageId="36" pageNumber="37" reason="6">
|
||
<emphasis id="B904C14EE571FF9FFCC4F999B89A054D" bold="true" box="[818,904,1548,1567]" pageId="36" pageNumber="37">Variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCF1D5CE571FF9FFCC4F9BFBF9C0508" blockId="36.[818,1442,1548,1626]" pageId="36" pageNumber="37">
|
||
There is no information about intraspecific variation since the species is only known from the
|
||
<typeStatus id="54CBA3FEE571FF9FFBC1F9D2BF990508" box="[1079,1163,1607,1626]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |