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<document id="D49B6A4BDA8F8F0865C46D1DDE5525AF" ID-DOI="10.5281/zenodo.190308" ID-GBIF-Dataset="b1039475-ade6-49f6-8ea7-c92c35b61c52" ID-ISSN="1175-5326" ID-Zenodo-Dep="190308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461111372257" checkinUser="plazi" docAuthor="Maduwage, Kalana, Silva, Anjana, Manamendra-Arachchi, Kelum &amp; Pethiyagoda, Rohan" docDate="2009" docId="038D1C750D6A7D1A028AFB1F14F8FB7A" docLanguage="en" docName="zt02232p028.pdf" docOrigin="Zootaxa 2232" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hypnale nepa Laurenti 1768" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFB4640D0D637D14021DFFCA165CFFE5" masterDocTitle="A taxonomic revision of the South Asian hump-nosed pit vipers (Squamata: Viperidae: Hypnale)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="10" updateTime="1698594252685" updateUser="plazi">
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<mods:title id="A931412555CA54E77962B205D717CD50">A taxonomic revision of the South Asian hump-nosed pit vipers (Squamata: Viperidae: Hypnale)</mods:title>
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<mods:namePart id="908EAC392E4698108EAFF254C0796F0D">Maduwage, Kalana</mods:namePart>
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<mods:namePart id="A9F06DA63EBD7C88B73AB93E41F935CF">Silva, Anjana</mods:namePart>
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<mods:namePart id="79AEC172587DF25A1CBF2C42E8FEA495">Manamendra-Arachchi, Kelum</mods:namePart>
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<mods:namePart id="E23DEFA79FA76172AD58D94EFE39EDF5">Pethiyagoda, Rohan</mods:namePart>
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<mods:date id="1C21E50DA5D4ACC7A7CF6D93AEB47F15">2009</mods:date>
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<treatment id="038D1C750D6A7D1A028AFB1F14F8FB7A" ID-DOI="http://doi.org/10.5281/zenodo.6212585" ID-GBIF-Taxon="119624057" ID-Zenodo-Dep="6212585" LSID="urn:lsid:plazi:treatment:038D1C750D6A7D1A028AFB1F14F8FB7A" httpUri="http://treatment.plazi.org/id/038D1C750D6A7D1A028AFB1F14F8FB7A" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10">
<subSubSection id="C33EFEE80D6A7D1D028AFB1F17F4FAF3" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9BAD630D6A7D1D028AFB1F144DFB0A" blockId="9.[151,529,1237,1302]" box="[151,529,1237,1263]" pageId="9" pageNumber="10">
<heading id="D0D31A0F0D6A7D1D028AFB1F144DFB0A" bold="true" box="[151,529,1237,1263]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="4C24D6E00D6A7D1D028AFB1F144DFB0A" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" box="[151,529,1237,1263]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6A7D1D028AFB1F144DFB0A" bold="true" box="[151,529,1237,1263]" pageId="9" pageNumber="10">
<emphasis id="B95071710D6A7D1D028AFB1F171CFB0A" bold="true" box="[151,320,1237,1263]" italics="true" pageId="9" pageNumber="10">Hypnale nepa</emphasis>
(
<bibRefCitation id="EFB5D0920D6A7D1D0353FB1F1455FB0A" author="Laurenti" box="[334,521,1237,1263]" pageId="9" pageNumber="27" refString="Laurenti, J. N. (1768) Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium Austriacorum. Trattnern, Viennae, 1 - 2, 1 - 214, [1], Tab. I - V." type="book chapter" year="1768">Laurenti, 1768</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9BAD630D6A7D1D028AFB3617F4FAF3" blockId="9.[151,529,1237,1302]" box="[151,424,1276,1302]" pageId="9" pageNumber="10">
(
<figureCitation id="131FB1E60D6A7D1D0282FB3616BBFAF3" box="[159,231,1276,1302]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
;
<figureCitation id="131FB1E60D6A7D1D02E9FB361767FAF3" box="[244,315,1276,1302]" captionStart="FIGURE 5" captionStartId="12.[151,255,1957,1981]" captionTargetBox="[180,1406,194,1931]" captionTargetId="figure@12.[180,1408,194,1933]" captionTargetPageId="12" captionText="FIGURE 5. Hypnale nepa: a, iconotype, male, reproduced from Seba (1734); b, WHT 6515, neotype, male, 367.0 mm SVL, Agra Arboretum, Agrapatana, Sri Lanka; c, topotypical specimen showing different coloration, not preserved." httpUri="https://zenodo.org/record/190313/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
; Table 2)
</paragraph>
</subSubSection>
<subSubSection id="C33EFEE80D6A7D1D028AFA801512FA14" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8B9BAD630D6A7D1D028AFA80179AFA65" blockId="9.[151,460,1353,1409]" pageId="9" pageNumber="10">
<taxonomicName id="4C24D6E00D6A7D1D028AFA801790FA85" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" box="[151,460,1353,1377]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6A7D1D028AFA801775FA84" box="[151,297,1354,1377]" italics="true" pageId="9" pageNumber="10">Coluber nepa</emphasis>
<bibRefCitation id="EFB5D0920D6A7D1D0332FA831790FA85" author="Laurenti" box="[303,460,1353,1376]" pageId="9" pageNumber="27" refString="Laurenti, J. N. (1768) Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium Austriacorum. Trattnern, Viennae, 1 - 2, 1 - 214, [1], Tab. I - V." type="book chapter" year="1768">Laurenti, 1768</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C24D6E00D6A7D1D028AFAA017E5FA65" authority="Gloyd, 1977" authorityName="Gloyd" authorityYear="1977" box="[151,441,1385,1409]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="walli">
<emphasis id="B95071710D6A7D1D028AFAA01772FA64" box="[151,302,1386,1409]" italics="true" pageId="9" pageNumber="10">Hypnale walli</emphasis>
<bibRefCitation id="EFB5D0920D6A7D1D0329FAA317E5FA65" author="Gloyd" box="[308,441,1385,1408]" pageId="9" pageNumber="26" refString="Gloyd, H. K. (1977 b) Descriptions of new taxa of crotalid snakes from China and Ceylon (Sri Lanka). Proceedings of the Biological Society of Washington, 90, 1002 - 1015." type="journal article" year="1977" yearSuffix="b">Gloyd, 1977</bibRefCitation>
</taxonomicName>
b
</paragraph>
<paragraph id="8B9BAD630D6A7D1D028AFA7A1512FA14" blockId="9.[151,1437,1456,2007]" pageId="9" pageNumber="10">
<emphasis id="B95071710D6A7D1D028AFA7A1756FA2F" bold="true" box="[151,266,1456,1482]" pageId="9" pageNumber="10">
<typeStatus id="549F13C10D6A7D1D028AFA7A1759FA2F" box="[151,261,1456,1482]" pageId="9" pageNumber="10" type="neotype">Neotype</typeStatus>
.
</emphasis>
(Here designated), WHT 6515, male,
<quantity id="4CDC00860D6A7D1D00F7FA7A1513FA2F" box="[746,847,1456,1482]" metricMagnitude="-1" metricUnit="m" metricValue="3.67" pageId="9" pageNumber="10" unit="mm" value="367.0">367 mm</quantity>
SVL, Agra Arboretum, Agarapatana,
<geoCoordinate id="EE10CBA40D6A7D1D0730FA7A13C4FA2F" box="[1325,1432,1456,1482]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="6.8333335">06º50N</geoCoordinate>
,
<geoCoordinate id="EE10CBA40D6A7D1D028AFA1D16A3FA14" box="[151,255,1495,1521]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="80.666664">80º40 E</geoCoordinate>
, alt.
<quantity id="4CDC00860D6A7D1D0325FA1D17C8FA14" box="[312,404,1495,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" pageId="9" pageNumber="10" unit="m" value="1665.0">1665 m</quantity>
, coll. S. V. Nanayakkara, 0
<date id="FF9A8BA30D6A7D1D00CCFA1D1515FA14" box="[721,841,1495,1521]" pageId="9" pageNumber="10" value="2006-03-04">4 III 2006</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33EFEE80D6A7D1D02DBFA37120AF8C3" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B9BAD630D6A7D1D02DBFA37120AF8C3" blockId="9.[151,1437,1456,2007]" pageId="9" pageNumber="10">
<emphasis id="B95071710D6A7D1D02DBFA3717E7F9F2" bold="true" box="[198,443,1533,1559]" pageId="9" pageNumber="10">Measured material.</emphasis>
Males, 2 ex. (WHT 5836,
<collectionCode id="ED3535A60D6A7D1D00E8FA37151CF9F2" box="[757,832,1533,1559]" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="9" pageNumber="10">MMB</collectionCode>
,
<date id="FF9A8BA30D6A7D1D0151FA371588F9F2" box="[844,980,1533,1559]" pageId="9" pageNumber="10" value="2000-09-15">15 IX 2000</date>
; WHT 5953,
<collectionCode id="ED3535A60D6A7D1D066EFA3712E2F9F2" box="[1139,1214,1533,1559]" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="9" pageNumber="10">MMB</collectionCode>
, 0
<date id="FF9A8BA30D6A7D1D06C4FA371307F9F2" box="[1241,1371,1533,1559]" pageId="9" pageNumber="10" value="2001-12-05">5 XII 2001</date>
),
<quantity id="4CDC00860D6A7D1D076DFA371694F9DB" metricMagnitude="-1" metricUnit="m" metricValue="2.38" pageId="9" pageNumber="10" unit="mm" value="238.0">238 mm</quantity>
,
<quantity id="4CDC00860D6A7D1D02C8F9EE1767F9DB" box="[213,315,1572,1598]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" pageId="9" pageNumber="10" unit="mm" value="245.0">245 mm</quantity>
, SVL, respectively; females, 6 ex. (WHT 5833, DG,
<date id="FF9A8BA30D6A7D1D01A4F9EE1206F9DB" box="[953,1114,1572,1598]" pageId="9" pageNumber="10" value="1998-08-10">10 VIII 1998</date>
; WHT 5834,
<collectionCode id="ED3535A60D6A7D1D06E3F9EE1316F9DB" box="[1278,1354,1572,1598]" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="9" pageNumber="10">MMB</collectionCode>
,
<date id="FF9A8BA30D6A7D1D074AF9EE168AF981" pageId="9" pageNumber="10" value="2001-06-20">20 VI 2001</date>
; WHT 5845, DG,
<date id="FF9A8BA30D6A7D1D03ADF980145CF981" box="[432,512,1610,1636]" pageId="9" pageNumber="10" value="1999-01">I 1999</date>
; WHT 6081,
<collectionCode id="ED3535A60D6A7D1D00BEF98014B2F981" box="[675,750,1610,1636]" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="9" pageNumber="10">MMB</collectionCode>
,
<date id="FF9A8BA30D6A7D1D00E6F98015DAF981" box="[763,902,1610,1636]" pageId="9" pageNumber="10" value="2004-04-10">10 IV 2004</date>
, WHT 6082,
<collectionCode id="ED3535A60D6A7D1D0634F9801228F981" box="[1065,1140,1610,1636]" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="9" pageNumber="10">MMB</collectionCode>
, 0
<date id="FF9A8BA30D6A7D1D068CF980135EF981" box="[1169,1282,1610,1636]" pageId="9" pageNumber="10" value="2004-02-03">3 II 2004</date>
; WHT 5954, KM,
<date id="FF9A8BA30D6A7D1D02C4F9BB1732F96E" box="[217,366,1649,1675]" pageId="9" pageNumber="10" value="2003-11-25">25 XI 2003</date>
),
<quantity id="4CDC00860D6A7D1D0398F9BB17B7F96E" box="[389,491,1649,1675]" metricMagnitude="-1" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="10" unit="mm" value="142.0">142 mm</quantity>
SVL,
<quantity id="4CDC00860D6A7D1D005DF9BB14FBF96E" box="[576,679,1649,1675]" metricMagnitude="-1" metricUnit="m" metricValue="3.42" pageId="9" pageNumber="10" unit="mm" value="342.0">342 mm</quantity>
SVL,
<quantity id="4CDC00860D6A7D1D00E6F9BB153EF96E" box="[763,866,1649,1675]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="9" pageNumber="10" unit="mm" value="235.0">235 mm</quantity>
SVL,
<quantity id="4CDC00860D6A7D1D01ABF9BB1241F96E" box="[950,1053,1649,1675]" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="9" pageNumber="10" unit="mm" value="315.0">315 mm</quantity>
SVL,
<quantity id="4CDC00860D6A7D1D066FF9BB1284F96E" box="[1138,1240,1649,1675]" metricMagnitude="-1" metricUnit="m" metricValue="2.92" pageId="9" pageNumber="10" unit="mm" value="292.0">292 mm</quantity>
SVL, damaged specimen, respectively, Agra Arboretum, Agarapatana,
<geoCoordinate id="EE10CBA40D6A7D1D013EF95215D4F957" box="[803,904,1688,1714]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="6.8333335">06º50N</geoCoordinate>
,
<geoCoordinate id="EE10CBA40D6A7D1D0189F95215A1F957" box="[916,1021,1688,1714]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="80.666664">80º40 E</geoCoordinate>
, alt.
<quantity id="4CDC00860D6A7D1D062BF95212CEF957" box="[1078,1170,1688,1714]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" pageId="9" pageNumber="10" unit="m" value="1665.0">1665 m</quantity>
. Females, 3 ex. (WHT 2002.S, DG, 0
<date id="FF9A8BA30D6A7D1D0320F97517E9F93C" box="[317,437,1727,1753]" pageId="9" pageNumber="10" value="1993-11-08">8 XI 1993</date>
; WHT 2002.L, DG, 0
<date id="FF9A8BA30D6A7D1D00AEF9751570F93C" box="[691,812,1727,1753]" pageId="9" pageNumber="10" value="1993-11-08">8 XI 1993</date>
; WHT 5838, SB and KW,
<date id="FF9A8BA30D6A7D1D067EF97512B1F93C" box="[1123,1261,1727,1753]" pageId="9" pageNumber="10" value="2000-04-20">20 IV 2000</date>
),
<quantity id="4CDC00860D6A7D1D071CF9751339F93C" box="[1281,1381,1727,1753]" metricMagnitude="-1" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="10" unit="mm" value="263.0">263 mm</quantity>
,
<quantity id="4CDC00860D6A7D1D076DF9751694F91A" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="9" pageNumber="10" unit="mm" value="315.0">315 mm</quantity>
,
<quantity id="4CDC00860D6A7D1D02C8F92F1764F91A" box="[213,312,1765,1791]" metricMagnitude="-1" metricUnit="m" metricValue="2.78" pageId="9" pageNumber="10" unit="mm" value="278.0">278 mm</quantity>
SVL, respectively, Adams Peak,
<geoCoordinate id="EE10CBA40D6A7D1D00C5F92F1563F91A" box="[728,831,1765,1791]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="6.8166666">06º49N</geoCoordinate>
,
<geoCoordinate id="EE10CBA40D6A7D1D0151F92F15E4F91A" box="[844,952,1765,1791]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="80.5">80º30 E</geoCoordinate>
, alt.
<quantity id="4CDC00860D6A7D1D01E9F92F120FF91A" box="[1012,1107,1765,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="9" pageNumber="10" unit="m" value="1830.0">1830 m</quantity>
. Female 1 ex. (WHT 5837, KM and
<collectingCountry id="F333EDF30D6A7D1D02E2F8C6177BF8C3" box="[255,295,1804,1830]" name="American Samoa" pageId="9" pageNumber="10">AS</collectingCountry>
, 0
<date id="FF9A8BA30D6A7D1D035FF8C617EEF8C3" box="[322,434,1804,1830]" pageId="9" pageNumber="10" value="2003-02-07">7 II 2003</date>
),
<quantity id="4CDC00860D6A7D1D03D8F8C6147BF8C3" box="[453,551,1804,1830]" metricMagnitude="-1" metricUnit="m" metricValue="2.92" pageId="9" pageNumber="10" unit="mm" value="292.0">292 mm</quantity>
SVL, Hakgala,
<geoCoordinate id="EE10CBA40D6A7D1D00F9F8C61515F8C3" box="[740,841,1804,1830]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="6.9166665">06º55N</geoCoordinate>
,
<geoCoordinate id="EE10CBA40D6A7D1D0148F8C615E2F8C3" box="[853,958,1804,1830]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="80.816666">80º49 E</geoCoordinate>
, alt.
<quantity id="4CDC00860D6A7D1D01EBF8C6120EF8C3" box="[1014,1106,1804,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="10" unit="m" value="1850.0">1850 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33EFEE80D6A7D1E02DBF8F81304FB15" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B9BAD630D6A7D1E02DBF8F815C2FEBA" blockId="9.[151,1437,1456,2007]" lastBlockId="10.[151,1437,152,1079]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B95071710D6A7D1D02DBF8F8171AF8A9" bold="true" box="[198,326,1842,1868]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
<taxonomicName id="4C24D6E00D6A7D1D034BF8F9145FF8A9" box="[342,515,1843,1868]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6A7D1D034BF8F9145FF8A9" box="[342,515,1843,1868]" italics="true" pageId="9" pageNumber="10">Hypnale nepa</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of the following characters: snout tip flattened or moderately raised, with 515 minute scales forming a wart-like protuberance at tip of snout (
<figureCitation id="131FB1E60D6A7D1D0353F8B217CAF877" box="[334,406,1912,1938]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
a, b); 1225, heterogeneous scales on internasal-prefrontal region (
<figureCitation id="131FB1E60D6A7D1D06ACF8B212A6F877" box="[1201,1274,1912,1938]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
b); 69 scales surround eye (supraocular + 2 postoculars + 03 suboculars + 2 preoculars + postfoveal) (
<figureCitation id="131FB1E60D6A7D1D06FEF8501372F851" box="[1251,1326,1946,1972]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
a); 4 or 5 scales surround maxillary pit—lower preocular + postfoveal + lacunal + upper edge of second supralabial, or lower preocular + postfoveal + lacunal + upper posterior border of second supralabial and a small scale above lacunal (see
<figureCitation id="131FB1E60D697D1E0333FF70172BFF31" box="[302,375,186,212]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
a); 13 scales between postfoveal and 2nd and 3rd supralabials (
<figureCitation id="131FB1E60D697D1E0663FF701294FF31" box="[1150,1224,186,212]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
a); postocular, 1st lower temporal, and 4th and 5th supralabials surround a rhomboid (diamond-shaped) scale of similar size (
<figureCitation id="131FB1E60D697D1E02BDFECA16B5FEFF" box="[160,233,256,282]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
a). Supralabials 7 or 8; a lacunal scale present; mid-dorsal scales smooth (
<figureCitation id="131FB1E60D697D1E064AFECA12FCFEFF" box="[1111,1184,256,282]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="10" pageNumber="11">
Fig.
<quantity id="4CDC00860D697D1E0692FECA12FCFEFF" box="[1167,1184,256,282]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="10" pageNumber="11" unit="g" value="4.0">4</quantity>
</figureCitation>
g); ventrals 122134; subcaudal series divided medially, with 2741 scales. Hemipenial lobe with large spines on proximal twothirds of its length; distal third with calyces, not spinous (
<figureCitation id="131FB1E60D697D1E0125FE8F15DCFEBA" box="[824,896,325,351]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
d).
</paragraph>
<paragraph id="8B9BAD630D697D1E02DBFEA217E8FDCA" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="11">
<taxonomicName id="4C24D6E00D697D1E02DBFEA21736FE64" box="[198,362,360,385]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D697D1E02DBFEA21736FE64" box="[198,362,360,385]" italics="true" pageId="10" pageNumber="11">Hypnale nepa</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C24D6E00D697D1E006CFEA214AEFE64" box="[625,754,360,385]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hypnale">
<emphasis id="B95071710D697D1E006CFEA214AEFE64" box="[625,754,360,385]" italics="true" pageId="10" pageNumber="11">H. hypnale</emphasis>
</taxonomicName>
by possessing (
<emphasis id="B95071710D697D1E01B0FEA2159AFE64" box="[941,966,360,385]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
lacking) a wart-like protuberance at the tip of the snout; having 13 (
<emphasis id="B95071710D697D1E03F4FE40145EFE46" box="[489,514,394,419]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
no) scales between the postfoveal and the 2nd and 3rd supralabials; having the postocular, 1st lower temporal, and 4th and 5th supralabials separated by a rhomboid scale of similar size (
<emphasis id="B95071710D697D1E0799FE6713C0FE23" box="[1412,1436,429,454]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
in contact); having the mid-dorsal scales smooth (
<emphasis id="B95071710D697D1E00EDFE1A1555FE0C" box="[752,777,464,489]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
all costal scales keeled); possessing 122134 (
<emphasis id="B95071710D697D1E0722FE1A1304FE0C" box="[1343,1368,464,489]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
141 158) ventrals; and having the hemipenial lobes with large spines on their proximal two-thirds (
<emphasis id="B95071710D697D1E0724FE38130FFDEE" box="[1337,1363,498,523]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
lobes smooth, lacking spines).
</paragraph>
<paragraph id="8B9BAD630D697D1E02DBFDF214A7FCC7" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="11">
It is distinguished from
<taxonomicName id="4C24D6E00D697D1E03C1FDF2146FFDB4" box="[476,563,568,593]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="zara">
<emphasis id="B95071710D697D1E03C1FDF2146FFDB4" box="[476,563,568,593]" italics="true" pageId="10" pageNumber="11">H. zara</emphasis>
</taxonomicName>
by having the snout tip flattened or only slightly raised (
<emphasis id="B95071710D697D1E06D4FDF212BDFDB4" box="[1225,1249,568,593]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
snout distinctly elevated); 1225 (
<emphasis id="B95071710D697D1E039EFD9017C2FD96" box="[387,414,602,627]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
1839) heterogeneous scales on internasal-prefrontal region; 45 (
<emphasis id="B95071710D697D1E0715FD90137FFD96" box="[1288,1315,602,627]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
3) scales surrounding maxillary pit; 13 (
<emphasis id="B95071710D697D1E0001FDB71469FD73" box="[540,565,637,662]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
no) scales between postfoveal and 2nd and 3rd supralabials; having the lower postocular, 1st lower temporal, and 4th and 5th supralabials surrounding a rhomboid scale of similar size (
<emphasis id="B95071710D697D1E02CBFD0816B3FD3E" box="[214,239,706,731]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
in contact, no scale between them); a lacunal scale present (
<emphasis id="B95071710D697D1E01A0FD081589FD3E" box="[957,981,706,731]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
absent); the mid-dorsal scales smooth (
<emphasis id="B95071710D697D1E02BCFD2F16E6FD1B" box="[161,186,741,766]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
all costal scales keeled); 122134 (
<emphasis id="B95071710D697D1E0071FD2F14D9FD1B" box="[620,645,741,766]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
134157) ventrals; and the hemipenial lobe with large spines (
<emphasis id="B95071710D697D1E079EFD2F13C0FD1B" box="[1411,1436,741,766]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
without spines) on proximal two-thirds of its length.
</paragraph>
<paragraph id="8B9BAD630D697D1E02DBFCE01497FBD2" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="11">
It is distinguished from
<taxonomicName id="4C24D6E00D697D1E03F6FCE01412FCA6" box="[491,590,810,835]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B95071710D697D1E03F6FCE01412FCA6" box="[491,590,810,835]" italics="true" pageId="10" pageNumber="11">Hypnale</emphasis>
</taxonomicName>
sp. amal by possessing (
<emphasis id="B95071710D697D1E0189FCE015F1FCA6" box="[916,941,810,835]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
lacking) a wart-like protuberance on the snout tip; having 13 (
<emphasis id="B95071710D697D1E0383FC8717EAFC83" box="[414,438,845,870]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
no) scales between postfoveal and 2nd and 3rd supralabials; the lower postocular, 1st lower temporal, and 4th and 5th supralabials separated by a rhomboid scale of similar size (
<emphasis id="B95071710D697D1E06C7FCBA12AFFC6C" box="[1242,1267,880,905]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
in contact, no scale between them); 7 or 8 (
<emphasis id="B95071710D697D1E03EEFC581450FC4E" box="[499,524,914,939]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
9) supralabials; first infralabials on either side separated by a median suture (
<emphasis id="B95071710D697D1E02BCFC7F16E7FC2B" box="[161,187,949,974]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
medially fused); mid-dorsal scales smooth (
<emphasis id="B95071710D697D1E00F1FC7F155AFC2B" box="[748,774,949,974]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
all costal scales keeled); 122134 (
<emphasis id="B95071710D697D1E06D1FC7F12BBFC2B" box="[1228,1255,949,974]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
151) ventrals; hemipenial lobe with large spines on proximal two-thirds of its length (
<emphasis id="B95071710D697D1E01F8FC1215A2FC14" box="[997,1022,984,1009]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
not spinous); two rows of distinct, bilaterally symmetrical blotches meeting on dorsal midline (
<emphasis id="B95071710D697D1E0171FC3015D9FBF6" box="[876,901,1018,1043]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
a distinct purple-brown paravertebral stripe, about 3 scales wide, from neck to level of vent).
</paragraph>
<paragraph id="8B9BAD630D697D1E028AFBA81304FB15" blockId="10.[151,1437,1122,1264]" pageId="10" pageNumber="11">
<emphasis id="B95071710D697D1E028AFBA81746FB99" bold="true" box="[151,282,1122,1148]" pageId="10" pageNumber="11">TABLE 2:</emphasis>
Measurements as a proportion of snout-vent length and head length of
<emphasis id="B95071710D697D1E067DFBA91336FB99" box="[1120,1386,1123,1148]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C24D6E00D697D1E067DFBA9135AFB99" box="[1120,1286,1123,1148]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nepa">Hypnale nepa</taxonomicName>
,
<taxonomicName id="4C24D6E00D697D1E070FFBA91336FB99" box="[1298,1386,1123,1148]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="zara">H. zara</taxonomicName>
</emphasis>
and
<emphasis id="B95071710D697D1E028AFB4316E8FB47" box="[151,180,1161,1186]" italics="true" pageId="10" pageNumber="11">H.</emphasis>
sp. amal. Abbreviations: SVL, snout-vent length;TL, tail length; HL, head length; HW, head width; SL, snout legnth; FL, frontal legth; FW, frontal width; PL, parietal length; PL, parietal width; SU, supraocular length; SW, supraocular width; SS, frontal supraocular suture. See text for details of material measured.
</paragraph>
</subSubSection>
<subSubSection id="C33EFEE80D697D1A035AFA8F14F8FB7A" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11" type="description">
<paragraph id="8B9BAD630D697D1E035AFA8F1365FA9E" blockId="10.[327,1337,1349,1403]" pageId="10" pageNumber="11">
<taxonomicName id="4C24D6E00D697D1E035AFA8F17C7FAB9" box="[327,411,1349,1372]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D697D1E035AFA8F1706FAB9" box="[327,346,1349,1372]" italics="true" pageId="10" pageNumber="11">H</emphasis>
.
<emphasis id="B95071710D697D1E037AFA8F17C7FAB9" box="[359,411,1349,1372]" italics="true" pageId="10" pageNumber="11">nepa</emphasis>
</taxonomicName>
<taxonomicName id="4C24D6E00D697D1E0026FA8F14D7FAB9" box="[571,651,1349,1372]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="zara">
H.
<emphasis id="B95071710D697D1E0041FA8F14D7FAB9" box="[604,651,1349,1372]" italics="true" pageId="10" pageNumber="11">zara</emphasis>
</taxonomicName>
<taxonomicName id="4C24D6E00D697D1E0124FA8F15D2FAB9" box="[825,910,1349,1372]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nepa">
H.
<emphasis id="B95071710D697D1E0147FA8F15D2FAB9" box="[858,910,1349,1372]" italics="true" pageId="10" pageNumber="11">nepa</emphasis>
</taxonomicName>
<taxonomicName id="4C24D6E00D697D1E062CFA8F12DCFAB9" box="[1073,1152,1349,1372]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="zara">
H.
<emphasis id="B95071710D697D1E064CFA8F12DCFAB9" box="[1105,1152,1349,1372]" italics="true" pageId="10" pageNumber="11">zara</emphasis>
</taxonomicName>
H. sp. amal
</paragraph>
<paragraph id="8B9BAD630D697D1E0349FA6B1360FA5D" blockId="10.[340,1340,1441,1464]" box="[340,1340,1441,1464]" pageId="10" pageNumber="11">male male female female male</paragraph>
<paragraph id="8B9BAD630D697D1E02E2FA1412AFFA10" blockId="10.[255,1267,1502,1525]" box="[255,1267,1502,1525]" pageId="10" pageNumber="11">N mean range s.d. Nmean range s.d. N mean range s.d. N mean range s.d.</paragraph>
<paragraph id="8B9BAD630D697D1E02CCF9D61362F995" pageId="10" pageNumber="11">
<table id="F9245FC30D6982EB02CCF9D61362F995" box="[209,1342,1564,1648]" colsContinueIn="10.[209,1342,1686,1985]" gridcols="12" gridrows="3" pageId="10" pageNumber="11">
<tr id="3514AF210D6982EB02CCF9D61362F9D6" box="[209,1342,1564,1587]" gridrow="0" pageId="10" pageNumber="11" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C5C65D0D6982EB02CCF9D6171AF9D6" box="[209,326,1564,1587]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">% SVL</th>
</tr>
<tr id="3514AF210D6982EB02CCF9F01362F9B4" box="[209,1342,1594,1617]" gridrow="1" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF9F0171AF9B4" box="[209,326,1594,1617]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">TL 6 20.9</th>
<td id="76C5C65D0D6982EB0352F9F017BAF9B4" box="[335,486,1594,1617]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">19.2 23.3 1.4</td>
<td id="76C5C65D0D6982EB03E4F9F014EDF9B4" box="[505,689,1594,1617]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">9 19.1 15.721.6</td>
<td id="76C5C65D0D6982EB00A7F9F01487F9B4" box="[698,731,1594,1617]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">1.8</td>
<td id="76C5C65D0D6982EB00F3F9F01555F9B4" box="[750,777,1594,1617]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">14</td>
<td id="76C5C65D0D6982EB0108F9F01518F9B4" box="[789,836,1594,1617]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">16.1</td>
<td id="76C5C65D0D6982EB0150F9F015BFF9B4" box="[845,995,1594,1617]" gridcol="6" gridrow="1" pageId="10" pageNumber="11">14.718.6 1.1</td>
<td id="76C5C65D0D6982EB01EBF9F0124DF9B4" box="[1014,1041,1594,1617]" gridcol="7" gridrow="1" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F9F01210F9B4" box="[1053,1100,1594,1617]" gridcol="8" gridrow="1" pageId="10" pageNumber="11">16.5</td>
<td id="76C5C65D0D6982EB064BF9F01298F9B4" box="[1110,1220,1594,1617]" gridcol="9" gridrow="1" pageId="10" pageNumber="11">11.920.5</td>
<td id="76C5C65D0D6982EB06D0F9F012B2F9B4" box="[1229,1262,1594,1617]" gridcol="10" gridrow="1" pageId="10" pageNumber="11">1.9</td>
<td id="76C5C65D0D6982EB0712F9F01362F9B4" box="[1295,1342,1594,1617]" gridcol="11" gridrow="1" pageId="10" pageNumber="11">18.2</td>
</tr>
<tr id="3514AF210D6982EB02CCF9931362F995" box="[209,1342,1625,1648]" gridrow="2" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF993171AF995" box="[209,326,1625,1648]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">HL 3 6.9</th>
<td id="76C5C65D0D6982EB0352F99317BAF995" box="[335,486,1625,1648]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">6.6 7.1 0.3</td>
<td id="76C5C65D0D6982EB03E4F99314EDF995" box="[505,689,1625,1648]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">5 6.3 5.8 7.3</td>
<td id="76C5C65D0D6982EB00A7F9931487F995" box="[698,731,1625,1648]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">0.7</td>
<td id="76C5C65D0D6982EB00F3F9931555F995" box="[750,777,1625,1648]" gridcol="4" gridrow="2" pageId="10" pageNumber="11">9</td>
<td id="76C5C65D0D6982EB0108F9931518F995" box="[789,836,1625,1648]" gridcol="5" gridrow="2" pageId="10" pageNumber="11">6.9</td>
<td id="76C5C65D0D6982EB0150F99315BFF995" box="[845,995,1625,1648]" gridcol="6" gridrow="2" pageId="10" pageNumber="11">6.2 8.2 0.6</td>
<td id="76C5C65D0D6982EB01EBF993124DF995" box="[1014,1041,1625,1648]" gridcol="7" gridrow="2" pageId="10" pageNumber="11">24</td>
<td id="76C5C65D0D6982EB0600F9931210F995" box="[1053,1100,1625,1648]" gridcol="8" gridrow="2" pageId="10" pageNumber="11">7.4</td>
<td id="76C5C65D0D6982EB064BF9931298F995" box="[1110,1220,1625,1648]" gridcol="9" gridrow="2" pageId="10" pageNumber="11">6.0 8.4</td>
<td id="76C5C65D0D6982EB06D0F99312B2F995" box="[1229,1262,1625,1648]" gridcol="10" gridrow="2" pageId="10" pageNumber="11">0.6</td>
<td id="76C5C65D0D6982EB0712F9931362F995" box="[1295,1342,1625,1648]" gridcol="11" gridrow="2" pageId="10" pageNumber="11">5.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9BAD630D697D1E02CCF95C1362F824" pageId="10" pageNumber="11">
<table id="F9245FC30D6982EB02CCF95C1362F824" box="[209,1342,1686,1985]" colsContinueFrom="10.[209,1342,1564,1648]" gridcols="12" gridrows="10" pageId="10" pageNumber="11">
<tr id="3514AF210D6982EB02CCF95C1362F948" box="[209,1342,1686,1709]" gridrow="0" pageId="10" pageNumber="11" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C5C65D0D6982EB02CCF95C171AF948" box="[209,326,1686,1709]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">% HL</th>
</tr>
<tr id="3514AF210D6982EB02CCF97F1362F929" box="[209,1342,1717,1740]" gridrow="1" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF97F171AF929" box="[209,326,1717,1740]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">HW 3 70.0</th>
<td id="76C5C65D0D6982EB0352F97F17BAF929" box="[335,486,1717,1740]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">67.4 74.8 4.1</td>
<td id="76C5C65D0D6982EB03E4F97F14EDF929" box="[505,689,1717,1740]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">5 61.1 47.871.7</td>
<td id="76C5C65D0D6982EB00A7F97F1487F929" box="[698,731,1717,1740]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">8.9</td>
<td id="76C5C65D0D6982EB00F3F97F1555F929" box="[750,777,1717,1740]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F97F1518F929" box="[789,836,1717,1740]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">72.2</td>
<td id="76C5C65D0D6982EB0150F97F15BFF929" box="[845,995,1717,1740]" gridcol="6" gridrow="1" pageId="10" pageNumber="11">59.284.6 7.3</td>
<td id="76C5C65D0D6982EB01EBF97F124DF929" box="[1014,1041,1717,1740]" gridcol="7" gridrow="1" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F97F1210F929" box="[1053,1100,1717,1740]" gridcol="8" gridrow="1" pageId="10" pageNumber="11">64.5</td>
<td id="76C5C65D0D6982EB0648F97F1298F929" box="[1109,1220,1717,1740]" gridcol="9" gridrow="1" pageId="10" pageNumber="11">53.674.1</td>
<td id="76C5C65D0D6982EB06D0F97F12B2F929" box="[1229,1262,1717,1740]" gridcol="10" gridrow="1" pageId="10" pageNumber="11">5.1</td>
<td id="76C5C65D0D6982EB0712F97F1362F929" box="[1295,1342,1717,1740]" gridcol="11" gridrow="1" pageId="10" pageNumber="11">66.7</td>
</tr>
<tr id="3514AF210D6982EB02CCF91E1362F90E" box="[209,1342,1748,1771]" gridrow="2" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF91E171AF90E" box="[209,326,1748,1771]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">SL 3 34.5</th>
<td id="76C5C65D0D6982EB0352F91E17BAF90E" box="[335,486,1748,1771]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">28.9 38.7 5.0</td>
<td id="76C5C65D0D6982EB03E4F91E14EDF90E" box="[505,689,1748,1771]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">5 38.8 30.841.8</td>
<td id="76C5C65D0D6982EB00A7F91E1487F90E" box="[698,731,1748,1771]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">4.6</td>
<td id="76C5C65D0D6982EB00F3F91E1555F90E" box="[750,777,1748,1771]" gridcol="4" gridrow="2" pageId="10" pageNumber="11">9</td>
<td id="76C5C65D0D6982EB0108F91E1518F90E" box="[789,836,1748,1771]" gridcol="5" gridrow="2" pageId="10" pageNumber="11">37.8</td>
<td id="76C5C65D0D6982EB0150F91E15BFF90E" box="[845,995,1748,1771]" gridcol="6" gridrow="2" pageId="10" pageNumber="11">33.944.7 3.6</td>
<td id="76C5C65D0D6982EB01EBF91E124DF90E" box="[1014,1041,1748,1771]" gridcol="7" gridrow="2" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F91E1210F90E" box="[1053,1100,1748,1771]" gridcol="8" gridrow="2" pageId="10" pageNumber="11">36.6</td>
<td id="76C5C65D0D6982EB0648F91E1298F90E" box="[1109,1220,1748,1771]" gridcol="9" gridrow="2" pageId="10" pageNumber="11">30.239.8</td>
<td id="76C5C65D0D6982EB06D0F91E12B2F90E" box="[1229,1262,1748,1771]" gridcol="10" gridrow="2" pageId="10" pageNumber="11">2.7</td>
<td id="76C5C65D0D6982EB0712F91E1362F90E" box="[1295,1342,1748,1771]" gridcol="11" gridrow="2" pageId="10" pageNumber="11">27.8</td>
</tr>
<tr id="3514AF210D6982EB02CCF9381362F8EC" box="[209,1342,1778,1801]" gridrow="3" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF938171AF8EC" box="[209,326,1778,1801]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">FL 3 19.9</th>
<td id="76C5C65D0D6982EB0352F93817BAF8EC" box="[335,486,1778,1801]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">17.8 22.6 2.5</td>
<td id="76C5C65D0D6982EB03E4F93814EDF8EC" box="[505,689,1778,1801]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">5 20.6 16.923.4</td>
<td id="76C5C65D0D6982EB00A7F9381487F8EC" box="[698,731,1778,1801]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">2.5</td>
<td id="76C5C65D0D6982EB00F3F9381555F8EC" box="[750,777,1778,1801]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F9381518F8EC" box="[789,836,1778,1801]" gridcol="5" gridrow="3" pageId="10" pageNumber="11">21.3</td>
<td id="76C5C65D0D6982EB0150F93815BFF8EC" box="[845,995,1778,1801]" gridcol="6" gridrow="3" pageId="10" pageNumber="11">16.830.9 3.7</td>
<td id="76C5C65D0D6982EB01EBF938124DF8EC" box="[1014,1041,1778,1801]" gridcol="7" gridrow="3" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F9381210F8EC" box="[1053,1100,1778,1801]" gridcol="8" gridrow="3" pageId="10" pageNumber="11">20.2</td>
<td id="76C5C65D0D6982EB0648F9381298F8EC" box="[1109,1220,1778,1801]" gridcol="9" gridrow="3" pageId="10" pageNumber="11">15.227.5</td>
<td id="76C5C65D0D6982EB06D0F93812B2F8EC" box="[1229,1262,1778,1801]" gridcol="10" gridrow="3" pageId="10" pageNumber="11">2.6</td>
<td id="76C5C65D0D6982EB0712F9381362F8EC" box="[1295,1342,1778,1801]" gridcol="11" gridrow="3" pageId="10" pageNumber="11">18.9</td>
</tr>
<tr id="3514AF210D6982EB02CCF8DB1362F8CD" box="[209,1342,1809,1832]" gridrow="4" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF8DB171AF8CD" box="[209,326,1809,1832]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">FW 3 17.1</th>
<td id="76C5C65D0D6982EB0352F8DB17BAF8CD" box="[335,486,1809,1832]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">15.318.3 1.6</td>
<td id="76C5C65D0D6982EB03E4F8DB14EDF8CD" box="[505,689,1809,1832]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">5 16.0 13.418.9</td>
<td id="76C5C65D0D6982EB00A7F8DB1487F8CD" box="[698,731,1809,1832]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">2.0</td>
<td id="76C5C65D0D6982EB00F3F8DB1555F8CD" box="[750,777,1809,1832]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8DB1518F8CD" box="[789,836,1809,1832]" gridcol="5" gridrow="4" pageId="10" pageNumber="11">18.9</td>
<td id="76C5C65D0D6982EB0150F8DB15BFF8CD" box="[845,995,1809,1832]" gridcol="6" gridrow="4" pageId="10" pageNumber="11">16.126.0 3.0</td>
<td id="76C5C65D0D6982EB01EBF8DB124DF8CD" box="[1014,1041,1809,1832]" gridcol="7" gridrow="4" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8DB1210F8CD" box="[1053,1100,1809,1832]" gridcol="8" gridrow="4" pageId="10" pageNumber="11">14.5</td>
<td id="76C5C65D0D6982EB0648F8DB1298F8CD" box="[1109,1220,1809,1832]" gridcol="9" gridrow="4" pageId="10" pageNumber="11">11.017.7</td>
<td id="76C5C65D0D6982EB06D0F8DB12B2F8CD" box="[1229,1262,1809,1832]" gridcol="10" gridrow="4" pageId="10" pageNumber="11">1.4</td>
<td id="76C5C65D0D6982EB0712F8DB1362F8CD" box="[1295,1342,1809,1832]" gridcol="11" gridrow="4" pageId="10" pageNumber="11">15.0</td>
</tr>
<tr id="3514AF210D6982EB02CCF8FA1362F8A2" box="[209,1342,1840,1863]" gridrow="5" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF8FA171AF8A2" box="[209,326,1840,1863]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">PL 3 30.1</th>
<td id="76C5C65D0D6982EB0352F8FA17BAF8A2" box="[335,486,1840,1863]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">28.6 31.5 1.5</td>
<td id="76C5C65D0D6982EB03E4F8FA14EDF8A2" box="[505,689,1840,1863]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">5 25.9 19.030.8</td>
<td id="76C5C65D0D6982EB00A7F8FA1487F8A2" box="[698,731,1840,1863]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">3.7</td>
<td id="76C5C65D0D6982EB00F3F8FA1555F8A2" box="[750,777,1840,1863]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8FA1518F8A2" box="[789,836,1840,1863]" gridcol="5" gridrow="5" pageId="10" pageNumber="11">32.4</td>
<td id="76C5C65D0D6982EB0150F8FA15BFF8A2" box="[845,995,1840,1863]" gridcol="6" gridrow="5" pageId="10" pageNumber="11">26.653.7 7.9</td>
<td id="76C5C65D0D6982EB01EBF8FA124DF8A2" box="[1014,1041,1840,1863]" gridcol="7" gridrow="5" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8FA1210F8A2" box="[1053,1100,1840,1863]" gridcol="8" gridrow="5" pageId="10" pageNumber="11">25.0</td>
<td id="76C5C65D0D6982EB0648F8FA1298F8A2" box="[1109,1220,1840,1863]" gridcol="9" gridrow="5" pageId="10" pageNumber="11">20.134.9</td>
<td id="76C5C65D0D6982EB06D0F8FA12B2F8A2" box="[1229,1262,1840,1863]" gridcol="10" gridrow="5" pageId="10" pageNumber="11">4.1</td>
<td id="76C5C65D0D6982EB0712F8FA1362F8A2" box="[1295,1342,1840,1863]" gridcol="11" gridrow="5" pageId="10" pageNumber="11">25.6</td>
</tr>
<tr id="3514AF210D6982EB02CCF8841362F880" box="[209,1342,1870,1893]" gridrow="6" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF884171AF880" box="[209,326,1870,1893]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">PW 3 18.3</th>
<td id="76C5C65D0D6982EB0352F88417BAF880" box="[335,486,1870,1893]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">16.9 19.0 1.2</td>
<td id="76C5C65D0D6982EB03E4F88414EDF880" box="[505,689,1870,1893]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">5 16.9 12.419.9</td>
<td id="76C5C65D0D6982EB00A7F8841487F880" box="[698,731,1870,1893]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">2.6</td>
<td id="76C5C65D0D6982EB00F3F8841555F880" box="[750,777,1870,1893]" gridcol="4" gridrow="6" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8841518F880" box="[789,836,1870,1893]" gridcol="5" gridrow="6" pageId="10" pageNumber="11">20.1</td>
<td id="76C5C65D0D6982EB0150F88415BFF880" box="[845,995,1870,1893]" gridcol="6" gridrow="6" pageId="10" pageNumber="11">16.726.0 3.4</td>
<td id="76C5C65D0D6982EB01EBF884124DF880" box="[1014,1041,1870,1893]" gridcol="7" gridrow="6" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8841210F880" box="[1053,1100,1870,1893]" gridcol="8" gridrow="6" pageId="10" pageNumber="11">16.2</td>
<td id="76C5C65D0D6982EB0648F8841298F880" box="[1109,1220,1870,1893]" gridcol="9" gridrow="6" pageId="10" pageNumber="11">12.724.1</td>
<td id="76C5C65D0D6982EB06D0F88412B2F880" box="[1229,1262,1870,1893]" gridcol="10" gridrow="6" pageId="10" pageNumber="11">2.5</td>
<td id="76C5C65D0D6982EB0712F8841362F880" box="[1295,1342,1870,1893]" gridcol="11" gridrow="6" pageId="10" pageNumber="11">17.8</td>
</tr>
<tr id="3514AF210D6982EB02CCF8A71362F861" box="[209,1342,1901,1924]" gridrow="7" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF8A7171AF861" box="[209,326,1901,1924]" gridcol="0" gridrow="7" pageId="10" pageNumber="11">SU 3 25.1</th>
<td id="76C5C65D0D6982EB0352F8A717BAF861" box="[335,486,1901,1924]" gridcol="1" gridrow="7" pageId="10" pageNumber="11">21.1 27.4 3.5</td>
<td id="76C5C65D0D6982EB03E4F8A714EDF861" box="[505,689,1901,1924]" gridcol="2" gridrow="7" pageId="10" pageNumber="11">5 22.7 15.925.9</td>
<td id="76C5C65D0D6982EB00A7F8A71487F861" box="[698,731,1901,1924]" gridcol="3" gridrow="7" pageId="10" pageNumber="11">4.0</td>
<td id="76C5C65D0D6982EB00F3F8A71555F861" box="[750,777,1901,1924]" gridcol="4" gridrow="7" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8A71518F861" box="[789,836,1901,1924]" gridcol="5" gridrow="7" pageId="10" pageNumber="11">25.0</td>
<td id="76C5C65D0D6982EB0150F8A715BFF861" box="[845,995,1901,1924]" gridcol="6" gridrow="7" pageId="10" pageNumber="11">20.237.4 5.3</td>
<td id="76C5C65D0D6982EB01EBF8A7124DF861" box="[1014,1041,1901,1924]" gridcol="7" gridrow="7" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8A71210F861" box="[1053,1100,1901,1924]" gridcol="8" gridrow="7" pageId="10" pageNumber="11">23.0</td>
<td id="76C5C65D0D6982EB0648F8A71298F861" box="[1109,1220,1901,1924]" gridcol="9" gridrow="7" pageId="10" pageNumber="11">18.328.3</td>
<td id="76C5C65D0D6982EB06D0F8A712B2F861" box="[1229,1262,1901,1924]" gridcol="10" gridrow="7" pageId="10" pageNumber="11">2.2</td>
<td id="76C5C65D0D6982EB0712F8A71362F861" box="[1295,1342,1901,1924]" gridcol="11" gridrow="7" pageId="10" pageNumber="11">24.4</td>
</tr>
<tr id="3514AF210D6982EB02CCF8461362F846" box="[209,1342,1932,1955]" gridrow="8" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF846171AF846" box="[209,326,1932,1955]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">SW 3 17.0</th>
<td id="76C5C65D0D6982EB0352F84617BAF846" box="[335,486,1932,1955]" gridcol="1" gridrow="8" pageId="10" pageNumber="11">16.1 17.7 0.8</td>
<td id="76C5C65D0D6982EB03E4F84614EDF846" box="[505,689,1932,1955]" gridcol="2" gridrow="8" pageId="10" pageNumber="11">4 18.2 15.420.9</td>
<td id="76C5C65D0D6982EB00A7F8461487F846" box="[698,731,1932,1955]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">2.3</td>
<td id="76C5C65D0D6982EB00F3F8461555F846" box="[750,777,1932,1955]" gridcol="4" gridrow="8" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8461518F846" box="[789,836,1932,1955]" gridcol="5" gridrow="8" pageId="10" pageNumber="11">18.1</td>
<td id="76C5C65D0D6982EB0150F84615BFF846" box="[845,995,1932,1955]" gridcol="6" gridrow="8" pageId="10" pageNumber="11">14.126.0 3.2</td>
<td id="76C5C65D0D6982EB01EBF846124DF846" box="[1014,1041,1932,1955]" gridcol="7" gridrow="8" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8461210F846" box="[1053,1100,1932,1955]" gridcol="8" gridrow="8" pageId="10" pageNumber="11">16.0</td>
<td id="76C5C65D0D6982EB0648F8461298F846" box="[1109,1220,1932,1955]" gridcol="9" gridrow="8" pageId="10" pageNumber="11">11.322.0</td>
<td id="76C5C65D0D6982EB06D0F84612B2F846" box="[1229,1262,1932,1955]" gridcol="10" gridrow="8" pageId="10" pageNumber="11">2.0</td>
<td id="76C5C65D0D6982EB0712F8461362F846" box="[1295,1342,1932,1955]" gridcol="11" gridrow="8" pageId="10" pageNumber="11">21.1</td>
</tr>
<tr id="3514AF210D6982EB02CCF8601362F824" box="[209,1342,1962,1985]" gridrow="9" pageId="10" pageNumber="11">
<th id="76C5C65D0D6982EB02CCF860171AF824" box="[209,326,1962,1985]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">SS 3 15.4</th>
<td id="76C5C65D0D6982EB0352F86017BAF824" box="[335,486,1962,1985]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">11.2 19.4 4.1</td>
<td id="76C5C65D0D6982EB03E4F86014EDF824" box="[505,689,1962,1985]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">5 14.9 13.417.0</td>
<td id="76C5C65D0D6982EB00A7F8601487F824" box="[698,731,1962,1985]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">2.7</td>
<td id="76C5C65D0D6982EB00F3F8601555F824" box="[750,777,1962,1985]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">10</td>
<td id="76C5C65D0D6982EB0108F8601518F824" box="[789,836,1962,1985]" gridcol="5" gridrow="9" pageId="10" pageNumber="11">13.6</td>
<td id="76C5C65D0D6982EB0150F86015BFF824" box="[845,995,1962,1985]" gridcol="6" gridrow="9" pageId="10" pageNumber="11">11.517.9 1.9</td>
<td id="76C5C65D0D6982EB01EBF860124DF824" box="[1014,1041,1962,1985]" gridcol="7" gridrow="9" pageId="10" pageNumber="11">26</td>
<td id="76C5C65D0D6982EB0600F8601210F824" box="[1053,1100,1962,1985]" gridcol="8" gridrow="9" pageId="10" pageNumber="11">14.2</td>
<td id="76C5C65D0D6982EB0648F8601298F824" box="[1109,1220,1962,1985]" gridcol="9" gridrow="9" pageId="10" pageNumber="11">11.820.2</td>
<td id="76C5C65D0D6982EB06D0F86012B2F824" box="[1229,1262,1962,1985]" gridcol="10" gridrow="9" pageId="10" pageNumber="11">2.0</td>
<td id="76C5C65D0D6982EB0712F8601362F824" box="[1295,1342,1962,1985]" gridcol="11" gridrow="9" pageId="10" pageNumber="11">16.1</td>
</tr>
</table>
</paragraph>
<caption id="DF5BFDEB0D687D1F028AF8D5155BF87B" httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="11" pageNumber="12" targetBox="[160,1415,209,1784]" targetPageId="11">
<paragraph id="8B9BAD630D687D1F028AF8D5155BF87B" blockId="11.[151,1437,1823,1951]" pageId="11" pageNumber="12">
<emphasis id="B95071710D687D1F028AF8D51743F8D2" bold="true" box="[151,287,1823,1847]" pageId="11" pageNumber="12">FIGURE 4.</emphasis>
<taxonomicName id="4C24D6E00D687D1F033BF8EA179EF8D2" box="[294,450,1824,1847]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D687D1F033BF8EA179EF8D2" box="[294,450,1824,1847]" italics="true" pageId="11" pageNumber="12">Hypnale nepa</emphasis>
</taxonomicName>
, WHT 6515, neotype, male, 367.0 mm SVL:
<emphasis id="B95071710D687D1F01DAF8EA1251F8D2" box="[967,1037,1824,1847]" italics="true" pageId="11" pageNumber="12">a, b, c</emphasis>
, lateral, dorsal and ventral aspects, respectively, of head;
<emphasis id="B95071710D687D1F0397F88817CBF8BC" box="[394,407,1858,1881]" italics="true" pageId="11" pageNumber="12">d</emphasis>
,
<emphasis id="B95071710D687D1F03BAF88817EFF8BC" box="[423,435,1858,1881]" italics="true" pageId="11" pageNumber="12">e</emphasis>
, everted left hemipenis, showing
<emphasis id="B95071710D687D1F0137F8881518F8BC" box="[810,836,1858,1881]" italics="true" pageId="11" pageNumber="12">(f)</emphasis>
enlarged view of a hemipenial lobe;
<emphasis id="B95071710D687D1F06FCF88812B2F8BC" box="[1249,1262,1858,1881]" italics="true" pageId="11" pageNumber="12">g</emphasis>
, dorsal view of paravertebral scale;
<emphasis id="B95071710D687D1F0371F8AF1725F899" box="[364,377,1893,1916]" italics="true" pageId="11" pageNumber="12">h</emphasis>
, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and
<emphasis id="B95071710D687D1F0660F8AF12D8F899" box="[1149,1156,1893,1916]" italics="true" pageId="11" pageNumber="12">i</emphasis>
, distribution in Sri Lanka:
<taxonomicName id="4C24D6E00D687D1F028AF84216B7F87A" box="[151,235,1928,1951]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D687D1F028AF84216B7F87A" box="[151,235,1928,1951]" italics="true" pageId="11" pageNumber="12">H. nepa</emphasis>
</taxonomicName>
(solid circles),
<taxonomicName id="4C24D6E00D687D1F038CF84217BBF87A" box="[401,487,1928,1951]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="walli">
<emphasis id="B95071710D687D1F038CF84217BBF87A" box="[401,487,1928,1951]" italics="true" pageId="11" pageNumber="12">H. walli</emphasis>
</taxonomicName>
type locality (open circle).
</paragraph>
</caption>
<caption id="DF5BFDEB0D6F7D18028AF86F1337F807" httpUri="https://zenodo.org/record/190313/files/figure.png" pageId="12" pageNumber="13" targetBox="[180,1406,194,1931]" targetPageId="12">
<paragraph id="8B9BAD630D6F7D18028AF86F1337F807" blockId="12.[151,1436,1957,2018]" pageId="12" pageNumber="13">
<emphasis id="B95071710D6F7D18028AF86F1745F858" bold="true" box="[151,281,1957,1981]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
<taxonomicName id="4C24D6E00D6F7D18033CF86F17EBF859" box="[289,439,1957,1980]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6F7D18033CF86F17EBF859" box="[289,439,1957,1980]" italics="true" pageId="12" pageNumber="13">Hypnale nepa</emphasis>
</taxonomicName>
:
<emphasis id="B95071710D6F7D1803DBF86F178FF859" box="[454,467,1957,1980]" italics="true" pageId="12" pageNumber="13">a</emphasis>
, iconotype, male, reproduced from Seba (1734);
<emphasis id="B95071710D6F7D1801FAF86F15A7F859" box="[999,1019,1957,1980]" italics="true" pageId="12" pageNumber="13">b,</emphasis>
WHT 6515, neotype, male, 367.0 mm SVL, Agra Arboretum, Agrapatana, Sri Lanka;
<emphasis id="B95071710D6F7D18008EF80114FAF807" box="[659,678,1995,2018]" italics="true" pageId="12" pageNumber="13">c,</emphasis>
topotypical specimen showing different coloration, not preserved.
</paragraph>
</caption>
<paragraph id="8B9BAD630D6E7D1902DBFF5D126FFBE1" blockId="13.[151,1437,151,1028]" pageId="13" pageNumber="14">
<emphasis id="B95071710D6E7D1902DBFF5D1703FF54" bold="true" box="[198,351,151,177]" pageId="13" pageNumber="14">Description.</emphasis>
Based on
<typeStatus id="549F13C10D6E7D1903FCFF521462FF57" box="[481,574,152,178]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
(WHT 6515). Snout tip distinctly raised, with 5 small scales forming a wart-like protuberance at its tip; 16 scales on internasal-prefrontal region, heterogeneous, irregularly arranged (
<figureCitation id="131FB1E60D6E7D1902BDFF2F16B5FF1A" box="[160,233,229,255]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a, b); internasals smaller than prefrontals, distinct (
<figureCitation id="131FB1E60D6E7D190153FF2F15C4FF1A" box="[846,920,229,255]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b); prefrontals distinct (
<figureCitation id="131FB1E60D6E7D1906AEFF2F12A0FF1A" box="[1203,1276,229,255]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b), somewhat displaced; internasals separated from each other and from rostral by a scaly, wart-like protuberance (
<figureCitation id="131FB1E60D6E7D19072BFEC61321FEC3" box="[1334,1405,268,294]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b); prefrontals distinguishable, anteriorly in contact with each other, posteriorly not in contact (
<figureCitation id="131FB1E60D6E7D190730FEF81320FEA9" box="[1325,1404,306,332]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b); prefrontals separated from frontal and supraoculars by 7 smaller scales (
<figureCitation id="131FB1E60D6E7D1901F7FE93126DFE96" box="[1002,1073,345,371]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
b); upper preocular, loreal, and nasals extend over canthal ridge, visible from above (
<figureCitation id="131FB1E60D6E7D190138FE4A152CFE7F" box="[805,880,384,410]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); rostral longer than wide, obtusely pointed, scarcely visible from above (
<figureCitation id="131FB1E60D6E7D1903E5FE6C141EFE25" box="[504,578,422,448]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); nasal entire (
<figureCitation id="131FB1E60D6E7D19011BFE6C150DFE25" box="[774,849,422,448]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); loreal acutely pointed anteriorly, its posterior margin broad, extending dorsally across canthus (
<figureCitation id="131FB1E60D6E7D1900F1FE07156AFE02" box="[748,822,461,487]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); 2 preoculars (
<figureCitation id="131FB1E60D6E7D19061DFE071216FE02" box="[1024,1098,461,487]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); 2 postoculars (
<figureCitation id="131FB1E60D6E7D19073DFE071335FE02" box="[1312,1385,461,487]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); a single subocular, reaching postfoveal; eye encircled by 7 scales: 2 preoculars + supraocular + 2 postoculars + subocular + postfoveal (
<figureCitation id="131FB1E60D6E7D1903D7FDD01444FDD1" box="[458,536,538,564]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a). Lacunal present (
<figureCitation id="131FB1E60D6E7D190107FDD01534FDD1" box="[794,872,538,564]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a). Maxillary pit bordered by 4 scales: lower preocular, postfoveal, lacunal and upper posterior border of 2nd supralabial (
<figureCitation id="131FB1E60D6E7D190648FD8B12FEFDBE" box="[1109,1186,577,603]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a). Two small scales between postfoveal and 3rd supralabial (
<figureCitation id="131FB1E60D6E7D190072FDA214EAFD67" box="[623,694,616,642]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); supralabials 7, not reaching eye (
<figureCitation id="131FB1E60D6E7D190644FDA212FCFD67" box="[1113,1184,616,642]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a); infralabials 8 (
<figureCitation id="131FB1E60D6E7D190771FDA216F4FD4D" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
c); 1st supralabial triangular, its apex divided into a small scale on left side (
<figureCitation id="131FB1E60D6E7D190601FD44123EFD4D" box="[1052,1122,654,680]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
a), undivided on right side; temporals 2+2+2, smooth, those on lower row greatly enlarged. Lower postocular, 1st lower temporal, and 4th and 5th supralabials surround a similar-sized rhomboid (diamond-shaped) scale. Numerous small scales present behind parietals. Mental subtriangular. A single pair of chin shields, slightly longer than wide (
<figureCitation id="131FB1E60D6E7D190771FCC816F4FCA6" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
c); infralabials I, II and III in contact with chin shields; mental and chin shields separated from each other by infralabial I. Gular scales in median series, 4; on lateral series, 6 (
<figureCitation id="131FB1E60D6E7D1901DCFC9A1251FC8F" box="[961,1037,848,874]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
c). Costal scales at mid-body 17. Dorsal scales smooth (
<figureCitation id="131FB1E60D6E7D1903B3FCBC17A6FC75" box="[430,506,886,912]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">
Fig.
<quantity id="4CDC00860D6E7D1903F5FCBC17A6FC75" box="[488,506,886,912]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="13" pageNumber="14" unit="g" value="4.0">4</quantity>
</figureCitation>
g). Paired apical pits present, inconspicuous. Ventrals 122; subcaudal series divided medially, with 41 scales; anal scale partially divided; 2 sub-anal scales on each side. Rows of costal scales up to 9th ventral on right side and 8th ventral on left, 19; 17 rows thereafter (including mid-body) up to 109th ventral on right side, 102nd on left, following which 15 rows up to anus.
</paragraph>
<paragraph id="8B9BAD630D6E7D1902DBFBF2142CFBB7" blockId="13.[198,624,1080,1106]" box="[198,624,1080,1106]" pageId="13" pageNumber="14">Costodorsal reduction (WHT 6515):</paragraph>
<paragraph id="8B9BAD630D6E7D1902D7FB4F178CFB7A" blockId="13.[151,581,1157,1260]" box="[202,464,1157,1183]" pageId="13" pageNumber="14">4+5(9) 3+4(109)</paragraph>
<paragraph id="8B9BAD630D6E7D19028AFB661419FB23" blockId="13.[151,581,1157,1260]" box="[151,581,1196,1222]" pageId="13" pageNumber="14">19 ————17—————15 (122)</paragraph>
<paragraph id="8B9BAD630D6E7D1902D7FB18178CFB09" blockId="13.[151,581,1157,1260]" box="[202,464,1234,1260]" pageId="13" pageNumber="14">4+5(8) 3+4(102)</paragraph>
<paragraph id="8B9BAD630D6E7D1902DBFAEA1308FA62" blockId="13.[151,1437,1312,2034]" pageId="13" pageNumber="14">
Hemipenes moderately long, slender, deeply forked; prominent spines present on proximal two-thirds of length of each lobe, the proximal-most ones longer than the distal-most ones; spines absent on distal third (
<figureCitation id="131FB1E60D6E7D190282FAA716BAFA62" box="[159,230,1389,1415]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
d); denticulated calyces cover distal portion of lobe; sulcus spermaticus shallow, its wall smooth.
</paragraph>
<paragraph id="8B9BAD630D6E7D1902DBFA5914C4F9C7" blockId="13.[151,1437,1312,2034]" pageId="13" pageNumber="14">
<emphasis id="B95071710D6E7D1902DBFA591720FA48" bold="true" box="[198,380,1427,1453]" pageId="13" pageNumber="14">Mensural data</emphasis>
(see also Table 2). Measurements of
<typeStatus id="549F13C10D6E7D190130FA5E15D2FA4B" box="[813,910,1428,1454]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
: SVL,
<quantity id="4CDC00860D6E7D1901C3FA5E121DFA4B" box="[990,1089,1428,1454]" metricMagnitude="-1" metricUnit="m" metricValue="3.67" pageId="13" pageNumber="14" unit="mm" value="367.0">367 mm</quantity>
; TL,
<quantity id="4CDC00860D6E7D190662FA5E128DFA4B" box="[1151,1233,1428,1454]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="mm" value="80.0">80 mm</quantity>
(21.8% of SVL); HL,
<quantity id="4CDC00860D6E7D1902D1FA701768FA31" box="[204,308,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="13" pageNumber="14" unit="mm" value="24.2">24.2 mm</quantity>
(6.6% of SVL); SL, 7.0 mm (28.9% of HL); FL,
<quantity id="4CDC00860D6E7D190169FA701592FA31" box="[884,974,1466,1492]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="13" pageNumber="14" unit="mm" value="4.3">4.3 mm</quantity>
(17.8% of HL); FW,
<quantity id="4CDC00860D6E7D1906DAFA70137DFA31" box="[1223,1313,1466,1492]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="14" unit="mm" value="3.7">3.7 mm</quantity>
(15.3% of HL); PL,
<quantity id="4CDC00860D6E7D190313FA2B1737FA1E" box="[270,363,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="14" unit="mm" value="7.3">7.3 mm</quantity>
(30.2% of HL); PW,
<quantity id="4CDC00860D6E7D190068FA2B148EFA1E" box="[629,722,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="14" unit="mm" value="4.1">4.1 mm</quantity>
(16.9% of HL); SU,
<quantity id="4CDC00860D6E7D1901C5FA2B1269FA1E" box="[984,1077,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="14" unit="mm" value="5.1">5.1 mm</quantity>
(21.1% of HL); SW,
<quantity id="4CDC00860D6E7D190722FA2B13C0FA1E" box="[1343,1436,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="14" unit="mm" value="3.9">3.9 mm</quantity>
(16.1% of HL); SS,
<quantity id="4CDC00860D6E7D19039EF9C21781F9C7" box="[387,477,1544,1570]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="14" unit="mm" value="2.7">2.7 mm</quantity>
(11.2% of HL).
</paragraph>
<paragraph id="8B9BAD630D6E7D1A02DBF9E4159FFEC3" blockId="13.[151,1437,1312,2034]" lastBlockId="14.[151,1437,152,1183]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B95071710D6E7D1902DBF9E41765F9AD" bold="true" box="[198,313,1582,1608]" pageId="13" pageNumber="14">Variation</emphasis>
(in material examined). Tip of snout flattened to moderately raised; wart like protuberance comprises 515 small scales; 1225 scales on internasal-prefrontal region. Prefrontals distinguishable or indistinguishable; when distinguishable, prefrontals in contact along their entire length, partially or altogether separated from each other. Prefrontals separated from frontal and supraoculars by 510 small scales. Nasal entire or partially divided above nostril; suboculars absent or 13: when absent, lower postocular reaches postfoveal. Eye encircled by 69 scales; a small scale either present or absent at upper angle of lacunal, below loreal and upper preocular. Maxillary pit bordered by 4 scales (lower preocular, postfoveal, lacunal and upper posterior border of second supralabial) or by 5 scales (lower preocular, postfoveal, lacunal, upper posterior border of second supralabial and small scale above lacunal). Small scales between postfoveal and 3rd supralabial, 13. A small triangular scale either present or absent beneath subocular, between 3rd and 4th supralabials. Supralabials, 7 or 8; apex of 1st supralabial divided or undivided; when undivided, 1st supralabial extends upwards to reach loreal. Infralabials I and II, or I, II and III in contact with chin shields; gular scales in median series, 46; gular scales in lateral series, 69. Costal scales smooth or keeled. Ventrals
<quantity id="4CDC00860D6D7D1A028AFF741741FF3D" box="[151,285,190,216]" metricMagnitude="0" metricUnit="m" metricValue="3.2131" metricValueMax="3.3274" metricValueMin="3.0987999999999998" pageId="14" pageNumber="15" unit="in" value="126.5" valueMax="131.0" valueMin="122.0">122131 in</quantity>
males,
<quantity id="4CDC00860D6D7D1A0367FF74145CFF3D" box="[378,512,190,216]" metricMagnitude="0" metricUnit="m" metricValue="3.2893" metricValueMax="3.4036" metricValueMin="3.175" pageId="14" pageNumber="15" unit="in" value="129.5" valueMax="134.0" valueMin="125.0">125134 in</quantity>
females; subcaudal series with 3741 scales in males,
<quantity id="4CDC00860D6D7D1A0693FF7412ABFF3D" box="[1166,1271,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="10.16" metricValueMin="6.858" pageId="14" pageNumber="15" unit="in" value="33.5" valueMax="40.0" valueMin="27.0">2740 in</quantity>
females; anal scale entire or partially divided; sub-anal scales on each side, 13. In WHT 5953, first two proximal-most scales of subcaudal series undivided, with 35 scales in divided series.
</paragraph>
<paragraph id="8B9BAD630D6D7D1A02DBFEF81228FE25" blockId="14.[151,1437,152,1183]" pageId="14" pageNumber="15">
<emphasis id="B95071710D6D7D1A02DBFEF81709FEA9" bold="true" box="[198,341,306,332]" pageId="14" pageNumber="15">Coloration.</emphasis>
In life, dorsal ground colour varies from yellowish-tan to dark brown. Two rows of distinct sub-oval or sub-triangular blotches meet on dorsal midline, bilaterally symmetrical (especially on tail), sometimes displaced longitudinally on body (see
<figureCitation id="131FB1E60D6D7D1A00EAFE4A1563FE7F" box="[759,831,384,410]" captionStart="FIGURE 5" captionStartId="12.[151,255,1957,1981]" captionTargetBox="[180,1406,194,1931]" captionTargetId="figure@12.[180,1408,194,1933]" captionTargetPageId="12" captionText="FIGURE 5. Hypnale nepa: a, iconotype, male, reproduced from Seba (1734); b, WHT 6515, neotype, male, 367.0 mm SVL, Agra Arboretum, Agrapatana, Sri Lanka; c, topotypical specimen showing different coloration, not preserved." httpUri="https://zenodo.org/record/190313/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
b, c). A dark stripe present across eye and cheek. Overall colour of venter much lighter than that of dorsum, sometimes light ash-grey.
</paragraph>
<paragraph id="8B9BAD630D6D7D1A02DBFE0717BAFD4D" blockId="14.[151,1437,152,1183]" pageId="14" pageNumber="15">
<emphasis id="B95071710D6D7D1A02DBFE071783FE02" bold="true" box="[198,479,461,487]" pageId="14" pageNumber="15">Natural history notes.</emphasis>
<taxonomicName id="4C24D6E00D6D7D1A03F1FE0714C8FE03" box="[492,660,461,486]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6D7D1A03F1FE0714C8FE03" box="[492,660,461,486]" italics="true" pageId="14" pageNumber="15">Hypnale nepa</emphasis>
</taxonomicName>
occurs frequently and widely in the central hills of
<collectingCountry id="F333EDF30D6D7D1A0700FE0713CBFE02" box="[1309,1431,461,487]" name="Sri Lanka" pageId="14" pageNumber="15">Sri Lanka</collectingCountry>
, where it is restricted to an altitudinal range of ~
<quantity id="4CDC00860D6D7D1A00DFFE3E1531FDEB" box="[706,877,500,526]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.85" metricValueMin="1.25" pageId="14" pageNumber="15" unit="m" value="1550.0" valueMax="1850.0" valueMin="1250.0">12501,850 m</quantity>
a.s.l (see
<figureCitation id="131FB1E60D6D7D1A01FFFE3E1275FDEB" box="[994,1065,500,526]" captionStart="FIGURE 4" captionStartId="11.[151,258,1823,1847]" captionTargetBox="[160,1415,209,1784]" captionTargetId="figure@11.[151,1436,194,1799]" captionTargetPageId="11" captionText="FIGURE 4. Hypnale nepa, WHT 6515, neotype, male, 367.0 mm SVL: a, b, c, lateral, dorsal and ventral aspects, respectively, of head; d, e, everted left hemipenis, showing (f) enlarged view of a hemipenial lobe; g, dorsal view of paravertebral scale; h, medial view of right palatine of WHT 6082, male, 292.0 mm SVL; and i, distribution in Sri Lanka: H. nepa (solid circles), H. walli type locality (open circle)." httpUri="https://zenodo.org/record/190312/files/figure.png" pageId="14" pageNumber="15">
Fig.
<date id="FF9A8BA30D6D7D1A0605FE3E1275FDEB" box="[1048,1065,500,526]" pageId="14" pageNumber="15">4</date>
</figureCitation>
i). It was also recorded from tea plantations, and does not appear to be adversely affected by forest loss in the islands central highlands. The only putative record of a specimen outside this range is that of the
<typeStatus id="549F13C10D6D7D1A01AAFD8B1243FDBE" box="[951,1055,577,603]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of
<emphasis id="B95071710D6D7D1A065AFD8B12F3FDBF" box="[1095,1199,577,602]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C24D6E00D6D7D1A065AFD8B12F6FDBF" box="[1095,1194,577,602]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="walli">H. walli</taxonomicName>
,
</emphasis>
stated to have been collected from Kanneliya in the south-western lowlands near Galle, which evidently was a mislabelled specimen (see Introduction).
</paragraph>
<paragraph id="8B9BAD630D6D7D1A02DBFD7F1449FC75" blockId="14.[151,1437,152,1183]" pageId="14" pageNumber="15">
We observed
<taxonomicName id="4C24D6E00D6D7D1A0378FD7F1455FD2B" box="[357,521,693,718]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6D7D1A0378FD7F1455FD2B" box="[357,521,693,718]" italics="true" pageId="14" pageNumber="15">Hypnale nepa</emphasis>
</taxonomicName>
to be more active at night, with specimens found in daytime being under logs and rocks among leaf litter in both undisturbed and secondary forest, more commonly at forest edges. It appears that
<taxonomicName id="4C24D6E00D6D7D1A0333FCC817D0FCFE" box="[302,396,770,795]" class="Reptilia" family="Viperidae" genus="Hypnale" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nepa">
<emphasis id="B95071710D6D7D1A0333FCC817D0FCFE" box="[302,396,770,795]" italics="true" pageId="14" pageNumber="15">H. nepa</emphasis>
</taxonomicName>
also feeds on other snakes: the gut of the
<typeStatus id="549F13C10D6D7D1A0198FCC815BFFCF9" box="[901,995,770,796]" pageId="14" pageNumber="15" type="neotype">neotype</typeStatus>
contained two Common rough-sides (
<taxonomicName id="4C24D6E00D6D7D1A02BFFCE31795FCA7" box="[162,457,809,834]" class="Reptilia" family="Colubridae" genus="Aspidura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="trachyprocta">
<emphasis id="B95071710D6D7D1A02BFFCE31795FCA7" box="[162,457,809,834]" italics="true" pageId="14" pageNumber="15">Aspidura trachyprocta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C24D6E00D6D7D1A03C7FCE3142DFCA6" box="[474,625,809,835]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="family">Colubridae</taxonomicName>
). In a study of gut contents,
<bibRefCitation id="EFB5D0920D6D7D1A01F7FCE312D9FCA6" author="Wall" box="[1002,1157,809,835]" pageId="14" pageNumber="27" refString="Wall, F. (1905) Notes on snakes collected at Hakgala, Ceylon. Spolia Zeylanica, 3, 144 - 147." type="journal article" year="1905">Wall (1905)</bibRefCitation>
noted that
<taxonomicName id="4C24D6E00D6D7D1A0706FCE31765FC8C" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="taprobanense">
<emphasis id="B95071710D6D7D1A0706FCE31765FC8C" italics="true" pageId="14" pageNumber="15">Lygosoma taprobanense</emphasis>
</taxonomicName>
appeared to be the staple food of this species at Hakgala, together with soft-shelled eggs [of serpents], frogs and rough-sides.
</paragraph>
<paragraph id="8B9BAD630D6D7D1A02DBFC5714F8FB7A" blockId="14.[151,1437,152,1183]" pageId="14" pageNumber="15">
A gravid female (WHT
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SVL) collected from near Hakgala on 0
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gave birth to five live young with adult coloration, measuring
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,
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,
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,
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, and
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TTL. Week-old juveniles were aggressive, frequently striking when alarmed. A gravid female (WHT
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SVL) collected at the Agra Arboretum on
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contained eight embryos (
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TBL) with yolk sacs and adult coloration. Another gravid female (
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TBL, WHT 5834), also collected at the Agra Arboretum on
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, contained seven fully-developed young (
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TBL), with the yolk sacs fully absorbed and showing adult coloration.
</paragraph>
</subSubSection>
</treatment>
</document>