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<document id="8D7C6E7FFB5886A94E8F434FD46D7B33" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FF8E3672FF76FA59BA88FEAC" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Geotrupes splendidus" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="7" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="1F703DB3792E518AE7C45F0D6EA83FA2">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
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<mods:namePart id="6568BBFD5EB915D1DA0C5AE291FD168D">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="D5222F321203520B1B12CD1B081BAF6C">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
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<mods:namePart id="F54B5FF29927C8384F84FED396D1C0BB">Veal, Doug A.</mods:namePart>
<mods:affiliation id="8D9095AA97AFE6E3EE46112D44227C0E">168 Thompson Drive SE Cedar Rapids, Iowa 52403</mods:affiliation>
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<mods:namePart id="CA9B4E56FEDE8FA48C92A0B84B22C69A">Lago, Paul K.</mods:namePart>
<mods:affiliation id="BC41DE935B3C0A933FECE5816BB3D8A1">Biology Department, University of Mississippi University, Mississippi 38677</mods:affiliation>
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<paragraph id="A0763684FF8E3673FF76FA59B9E5FA48" blockId="8.[192,722,1460,1484]" box="[192,722,1460,1484]" pageId="8" pageNumber="7">
<heading id="FB3E81E8FF8E3673FF76FA59B9E5FA48" bold="true" box="[192,722,1460,1484]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<taxonomicName id="67C94D07FF8E3673FF76FA59B9E5FA48" ID-CoL="7H6N4" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[192,722,1460,1484]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="92BDEA96FF8E3673FF76FA59B9E5FA48" bold="true" box="[192,722,1460,1484]" pageId="8" pageNumber="7">
<emphasis id="92BDEA96FF8E3673FF76FA59BAD1FA48" bold="true" box="[192,486,1460,1484]" italics="true" pageId="8" pageNumber="7">Geotrupes splendidus</emphasis>
(Fabricius, 1775)
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<paragraph id="A0763684FF8E3673FF76FA0FBF86FA7D" blockId="8.[192,1201,1505,1529]" box="[192,1201,1505,1529]" pageId="8" pageNumber="7">
<bibRefCitation id="C4584B75FF8E3673FF76FA0FBA5EFA7D" author="Putnam, J. D." box="[192,361,1505,1529]" pageId="8" pageNumber="7" pagination="169 - 173" refId="ref33576" refString="Putnam, J. D. 1876. List of Coleoptera found in the vicinity of Davenport, Iowa. Proceedings of the Davenport Academy of Natural Sciences 1: 169 - 173." type="journal article" year="1876">Putnam 1876</bibRefCitation>
;
<bibRefCitation id="C4584B75FF8E3673FEC3FA0CB918FA7D" author="Wickham, H. F." box="[373,559,1505,1529]" pageId="8" pageNumber="7" pagination="1 - 40" refId="ref34735" refString="Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40." type="journal article" year="1911">Wickham 1911</bibRefCitation>
;
<bibRefCitation id="C4584B75FF8E3673FD8DFA0CB9C2FA7D" author="Howden, H. F." box="[571,757,1505,1529]" pageId="8" pageNumber="7" pagination="151 - 319" refId="ref32123" refString="Howden, H. F. 1955 a. Biology and taxonomy of North American beetles of the subfamily Geotrupinae with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes, and Peltotrupes (Scarabaeidae). Proceedings of the United States National Museum 104 (3342): 151 - 319." type="journal article" year="1955">Howden 1955a</bibRefCitation>
; ISIC; IWUC; DAVC; ELFC; MJPC.
</paragraph>
</subSubSection>
<subSubSection id="E8D3650FFF8E3672FF76F9E4BA88FEAC" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7" type="description">
<paragraph id="A0763684FF8E3673FF76F9E4B97DF9A5" blockId="8.[192,586,1545,1569]" box="[192,586,1545,1569]" pageId="8" pageNumber="7">Collection dates: AprilOctober.</paragraph>
<paragraph id="A0763684FF8E3673FF76F9DDB8FCF9CC" blockId="8.[192,971,1584,1608]" box="[192,971,1584,1608]" pageId="8" pageNumber="7">Specimens were collected from dung, deer carcass, and at light.</paragraph>
<paragraph id="A0763684FF8E3673FF76F9B5BA13F909" blockId="8.[192,1440,1624,1677]" pageId="8" pageNumber="7">
Specimen label data: Sharon Bluffs State Park, Appanoose Co.,
<collectingRegion id="620DF866FF8E3673FC42F9B5BF04F9EB" box="[1012,1075,1624,1647]" country="United States of America" name="Iowa" pageId="8" pageNumber="7">Iowa</collectingRegion>
,
<date id="D4771044FF8E3673FBF4F9B5BE3AF9F4" box="[1090,1293,1624,1648]" pageId="8" pageNumber="7" value="2013-08-20">20 August 2013</date>
, light, ELF (ELFC).
</paragraph>
<paragraph id="A0763684FF8E3673FF76F971BE97F88E" blockId="8.[191,1440,1692,1803]" pageId="8" pageNumber="7">
County records (
<figureCitation id="38F22A01FF8E3673FE24F971B938F930" box="[402,527,1692,1716]" captionStart="Figures 1017" captionStartId="62.[192,287,1847,1869]" captionTargetBox="[147,1439,219,1804]" captionTargetPageId="62" captionText="Figures 1017. Distribution of Scarabaeoidea in Iowa. 10) Eucanthus lazarus (Fabricius, 1775). 11) Geotrupes blackburneus excrementi Say, 1823. 12) Geotrupes horni Blanchard, 1888. 13) Geotrupes opacus Haldeman, 1853. 14) Geotrupes semiopacus Jekel, 1865. 15) Geotrupes splendidus (Fabricius, 1775). 16) Trox aequalis Say, 1831." figureDoi="http://doi.org/10.5281/zenodo.5353796" httpUri="https://zenodo.org/record/5353796/files/figure.png" pageId="8" pageNumber="7">Figure 15</figureCitation>
): Adair, Allamakee, Appanoose, Black Hawk, Bremer, Clayton, Clinton, Dallas, Davis, Decatur,
<collectingRegion id="620DF866FF8E3673FE58F954B950F955" box="[494,615,1721,1745]" country="United States of America" name="Delaware" pageId="8" pageNumber="7">Delaware</collectingRegion>
, Des Moines, Dickinson, Fayette, Fremont, Hancock, Henry, Jasper, Jefferson, Johnson, Keokuk, Kossuth, Lee, Linn, Louisa, Madison, Marshall, Monona, Muscatine, Page, Polk, Pottawattamie, Story, Taylor, Van Buren, Wapello,
<collectingRegion id="620DF866FF8E3673FC7CF91FBF55F88E" box="[970,1122,1778,1802]" country="United States of America" name="Washington" pageId="8" pageNumber="7">Washington</collectingRegion>
, Winnebago, Winneshiek.
</paragraph>
<paragraph id="A0763684FF8F3672FF76FF3AB942FF6B" blockId="9.[192,669,215,296]" box="[192,629,215,239]" pageId="9" pageNumber="8">
Family
<taxonomicName id="67C94D07FF8F3672FEA9FF3AB942FF6B" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[287,629,215,239]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="family">TROGIDAE MacLeay, 1819</taxonomicName>
</paragraph>
<paragraph id="A0763684FF8F3672FF76FF19B9AAFE88" blockId="9.[192,669,215,296]" box="[192,669,244,268]" pageId="9" pageNumber="8">
Subfamily
<taxonomicName id="67C94D07FF8F3672FEF0FF19B9AAFE88" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[326,669,244,268]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Troginae">TROGINAE MacLeay, 1819</taxonomicName>
</paragraph>
<paragraph id="A0763684FF8F3672FF76FEFCBA88FEAC" blockId="9.[192,669,215,296]" box="[192,447,272,296]" pageId="9" pageNumber="8">
<taxonomicName id="67C94D07FF8F3672FF76FEFCBA88FEAC" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[192,447,272,296]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FF8F3672FF76FEFCBBCEFEAC" box="[192,249,273,296]" italics="true" pageId="9" pageNumber="8">Trox</emphasis>
Fabricius, 1775
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