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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="1DEBA93351915CB0A3F935EC3E44A6A0" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Orionella Jedlicka 1963" docType="treatment" docVersion="3" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="76" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="74" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:date>2013</mods:date>
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<mods:number>2013-04-04</mods:number>
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<mods:number>284</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
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<treatment ID-GBIF-Taxon="182235609" LSID="urn:lsid:plazi:treatment:1DEBA93351915CB0A3F935EC3E44A6A0" httpUri="http://treatment.plazi.org/id/1DEBA93351915CB0A3F935EC3E44A6A0" lastPageId="75" lastPageNumber="76" pageId="73" pageNumber="74">
<subSubSection pageId="73" pageNumber="74" type="nomenclature">
<paragraph pageId="73" pageNumber="74">
Genus
<taxonomicName LSID="1DEBA933-5191-5CB0-A3F9-35EC3E44A6A0" authority="Jedlicka, 1963" authorityName="Jedlicka" authorityYear="1963" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Orionella" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
Orionella
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1963
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="73" pageNumber="74" type="reference_group">
<paragraph pageId="73" pageNumber="74">
<taxonomicName authorityName="Jedlicka" authorityYear="1963" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">Orionella</taxonomicName>
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 307;
<bibRefCitation author="Habu, A" journalOrPublisher="Proceedings of the Japanese Society of Systematic Zoology" pageId="87" pageNumber="88" refId="B50" refString="Habu, A, 1979b. Notes on Endynomena Chaudoir and Orionella Jedlicka (Coleoptera, Carabidae). Entomological Review of Japan 33 (1/2): 61-65." title="Notes on Endynomena Chaudoir and Orionella Jedlicka (Coleoptera, Carabidae). Entomological Review of Japan 33 (1 / 2): 61 - 65." year="1979 b">Habu 1979b</bibRefCitation>
: 64.
</paragraph>
<paragraph pageId="73" pageNumber="74">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">Endynomena</taxonomicName>
auctt. [not
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Annales de Societe Entomologique de Belgique" pageId="86" pageNumber="87" pagination="97 - 204" refId="B34" refString="Chaudoir, BM, 1872. Monographie des Callidides. Annales de Societe Entomologique de Belgique 16: 97 - 204" title="Monographie des Callidides." volume="16" year="1872">Chaudoir 1872</bibRefCitation>
]:
<bibRefCitation author="Habu, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="87" pageNumber="88" refId="B48" refString="Habu, A, 1967. Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." title="Fauna Japonica, Carabidae Truncatipennes Group (Insecta: Coleoptera). Biogeographical Society of Japan. Japan, 338 pp." year="1967">Habu 1967</bibRefCitation>
: 131. [Synonym]
</paragraph>
</subSubSection>
<subSubSection pageId="73" pageNumber="74" type="type-species">
<paragraph pageId="73" pageNumber="74">Type-species:</paragraph>
<paragraph pageId="73" pageNumber="74">
<taxonomicName authorityName="Jedlicka" authorityYear="1964" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="obenbergeri">
<emphasis italics="true" pageId="73" pageNumber="74">Orionella obenbergeri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1963 [=
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena lewisii</emphasis>
</taxonomicName>
Bates], by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="73" pageNumber="74" type="diagnosis">
<paragraph pageId="73" pageNumber="74">Diagnosis.</paragraph>
<paragraph pageId="73" pageNumber="74">Dorsal side evenly pubescent, elytra with accessory setae on all intervals; mandibles strongly widened; labrum and outer scrobe of mandibles nearly glabrous; pronotum with basal foveae wide and shallow; elytral intervals slightly convex; median lobe of aedeagus slender and straight, not setose around apical orifice, apical lamella more or less bent backwards.</paragraph>
<paragraph pageId="73" pageNumber="74">
This genus strongly resembles
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
, but is not closely allied with it given the differences in the male genitalia and male secondary sexual characters. Beyond the remarkable differences in male genitalia (
<figureCitation captionStart="Figure 91" captionStartId="F36" captionText="Figure 91. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Orionella lewisii (Bates), a specimen from Zhejiang." httpUri="https://binary.pensoft.net/fig/29709" pageId="73" pageNumber="74">Figs 91</figureCitation>
<figureCitation captionStart="Figure 94" captionStartId="F39" captionText="Figure 94. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29715" pageId="73" pageNumber="74">-94</figureCitation>
),
<taxonomicName authorityName="Jedlicka" authorityYear="1963" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Orionella</emphasis>
</taxonomicName>
is distinguishable from
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
in having the following characters: (1) striae deeper, intervals slightly convex, striae distinct in apical half, but in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
striae shallow, intervals barely convex, striae nearly effaced in apical half; (2) 7th interval without several long setae, but in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
with such setae; (3) pronotal base weakly but distinctly lobed, but in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
pronotal base hardly lobed; (4) tempora less swollen than in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
; (5) apex of mentum tooth truncate, but always rounded in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
; (6) males with adhesive hairs only present on 1st and 2nd protarsomeres, but present on 1st to 3rd protarsomeres in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="73" pageNumber="74">Endynomena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="generic characters">
<paragraph pageId="73" pageNumber="74">Generic characters.</paragraph>
<paragraph lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
Body length 8.3-9.2 mm. Dorsal side reddish brown to dark brown, elytra unicolored or bicolored, without metallic luster; body rather flat; dorsal side densely pubescent, pubescence short, not erect.
<emphasis bold="true" pageId="73" pageNumber="74">Head.</emphasis>
eyes hemispherical, strongly prominent; tempora short, half length of eyes, slightly tumid, gradually narrowed behind eyes; vertex not distinctly tumid. Antennae barely extended to elytral base. Labrum widened to apex, apical margin distinctly emarginate, with faint isodiametric microsculpture, glabrous or with a few very fine short secondary setae; mandibles distinctly widened, outer margins rounded, glabrous on outer scrobes, with some setae arranged along dorsal ridge; terminal maxillary palpomeres slender in both sexes, api
<pageBreakToken pageId="74" pageNumber="75" start="start">ces</pageBreakToken>
slightly truncate; terminal labial palpomeres slightly widened in both sexes, males somewhat wider, but not typical securiform, apices slightly truncate; ligula with apex slightly projected, with four setae; paraglossae membranous, not longer than ligula, slightly widened, adnate; mentum tooth widened, trapezoid, apex truncate, with two long setae at base, sometimes with a few additional short setae; submentum with two long setae; genae with long setae beneath eyes; gula nearly glabrous except apical part.
<emphasis bold="true" pageId="74" pageNumber="75">Pronotum</emphasis>
transverse, surface densely pubescent; lateral margins with setae along full length, a few long setae present at front angles, four long setae at the greatest width, primary setae on hind angles slightly longer than others; basal foveae wide and shallow; pronotal base weakly but distinctly lobed; lateral margins completely rounded in middle, more or less sinuate before hind angles; hind angles sharp or not.
<emphasis bold="true" pageId="74" pageNumber="75">Elytra</emphasis>
wide, slightly expanded to apex; apex truncate, sutural angles not projected, outer angles completely rounded; basal margination only reaching 3rd interval; basal pores large; striae shallow but distinct, indistinctly punctate; intervals slightly convex; all intervals evenly pubescent, pubescence fine and longish, not erect; primary setigerous pores small, 3rd interval with four pores, 5th interval with one pore near base; 3rd to 6th intervals slightly depressed near middle, 7th and 8th intervals slightly tumid near apex.
<emphasis bold="true" pageId="74" pageNumber="75">Ventral side</emphasis>
with dense pubescence; males with apex of terminal sternum moderately emarginate, with two pairs of setae; females with apex of terminal sternum straight, with two pairs of setae.
<emphasis bold="true" pageId="74" pageNumber="75">Legs</emphasis>
short; protibiae with cleaning spur well developed, distant from inner margin; tarsi widened, 4th tarsomere bifid, claws pectinate. Males with adhesive hairs well developed (two whole rows) on 1st and 2nd protarsomeres and apical half of 1st mesotarsomere; rudimentary (very few near apex, sometimes absent) on 2nd mesotarsomere.
<emphasis bold="true" pageId="74" pageNumber="75">Male genitalia</emphasis>
with median lobe of aedeagus slender, not twisted, nearly straight in lateral view, slightly bent to right side in dorsal view; apical orifice opened apically; apical lamella small, rounded, more or less bent backward; right surface with some fine setae subapically; internal sac with main flagellum slightly thickened, apex distant from apical orifice, trumpet-form expansion small; an elongate scaled area subparallel to main flagellum; apical bursa present; secondary flagellum present.
<emphasis bold="true" pageId="74" pageNumber="75">Female genitalia.</emphasis>
Spermatheca tubular, with ring-sculpture, inserted on bursa copulatrix; spermathecal gland much longer than spermatheca, inserted near base of spermatheca; spermatheca not distinctly bent. Apical segment of ovipositor slender and straight, apex sharp, base gradually widened, apex with membranous extension short and rounded.
</paragraph>
</subSubSection>
<subSubSection lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="distribution">
<paragraph pageId="74" pageNumber="75">Distribution</paragraph>
<paragraph pageId="74" pageNumber="75">
(
<figureCitation captionStart="Map 8" captionStartId="F52" captionText="Map 8. Known distribution of genus Orionella: ● Orionella lewisii (Bates) ■ Orionella discoidalis (Bates) ▲ Orionella kathmanduensis (Kirschenhofer)." httpUri="https://binary.pensoft.net/fig/29730" pageId="74" pageNumber="75">Map 8</figureCitation>
). Japan, Korea, China, Burma, Nepal.
</paragraph>
<paragraph pageId="74" pageNumber="75">
<emphasis bold="true" pageId="74" pageNumber="75">Monophyly and relationships</emphasis>
. From general appearance, this genus could be allied with
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="74" pageNumber="75">Lachnoderma</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="74" pageNumber="75">Dasiosoma</emphasis>
</taxonomicName>
, but genital characters suggest that it could be rather isolated within
<taxonomicName genus="Physoderina" lsidName="" pageId="74" pageNumber="75" rank="genus">Physoderina</taxonomicName>
. The monophyly of
<taxonomicName authorityName="Jedlicka" authorityYear="1963" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="74" pageNumber="75">Orionella</emphasis>
</taxonomicName>
can be inferred from the following apomorphic character states: (1) median lobe of aedeagus tubular, not laterally compressed; (2) internal sac with main flagellum abruptly terminated before apical orifice; (3) apical lamella bent backward; (4) apical segment of ovipositor straight, not curved outward.
</paragraph>
<paragraph pageId="75" pageNumber="76">
<emphasis bold="true" pageId="75" pageNumber="76">
<pageBreakToken pageId="75" pageNumber="76" start="start">Key</pageBreakToken>
to species of
<taxonomicName authorityName="Jedlicka" authorityYear="1963" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="75" pageNumber="76">Orionella</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
</emphasis>
</paragraph>
<paragraph pageId="75" pageNumber="76">
<table inLine="true" pageId="75" pageNumber="76">
<tr pageId="75" pageNumber="76">
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">1</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">
Elytra reddish brown, with base and lateral margins broadly darkened forming distinct band (
<figureCitation captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Habitus, scale bars = 2.0 mm: 25 Dasiosoma bellum (Habu), a specimen from Guangdong 26 Dasiosoma bellum (Habu), a specimen from Sri Lanka 27 Orionella lewisii (Bates), a specimen from Japan 28 Orionella discoidalis (Bates), a specimen from Yunnan 29 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 30 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29678" pageId="75" pageNumber="76">Fig. 28</figureCitation>
); pronotum with lateral margins strongly sinuate before hind angles, hind angles very sharp, forming an acute angle; Yunnan and Myanmar
</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis italics="true" pageId="75" pageNumber="76">Orionella discoidalis</emphasis>
</taxonomicName>
(Bates)
</td>
</tr>
<tr pageId="75" pageNumber="76">
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">-</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">
Elytra unicolored (
<figureCitation captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Habitus, scale bars = 2.0 mm: 25 Dasiosoma bellum (Habu), a specimen from Guangdong 26 Dasiosoma bellum (Habu), a specimen from Sri Lanka 27 Orionella lewisii (Bates), a specimen from Japan 28 Orionella discoidalis (Bates), a specimen from Yunnan 29 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 30 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29678" pageId="75" pageNumber="76">Fig. 27</figureCitation>
) or disc with indistinct reddish patch (
<figureCitation captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Habitus, scale bars = 2.0 mm: 25 Dasiosoma bellum (Habu), a specimen from Guangdong 26 Dasiosoma bellum (Habu), a specimen from Sri Lanka 27 Orionella lewisii (Bates), a specimen from Japan 28 Orionella discoidalis (Bates), a specimen from Yunnan 29 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 30 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29678" pageId="75" pageNumber="76">Fig. 29</figureCitation>
), without distinct marginal band; pronotum with lateral margins nearly straight or slightly sinuate before hind angles, hind angles not so sharp
</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">2</td>
</tr>
<tr pageId="75" pageNumber="76">
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">2</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">Head and pronotum reddish brown, elytra dark brown, disc with indistinct reddish patch; pronotal hind angles slightly sharp, subrectangular; internal sac of aedeagus with scaled area less than one-fourth of median lobe; Nepal</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Kirschenhofer" baseAuthorityYear="1994" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="kathmanduensis">
<emphasis italics="true" pageId="75" pageNumber="76">Orionella kathmanduensis</emphasis>
</taxonomicName>
Kirschenhofer
</td>
</tr>
<tr pageId="75" pageNumber="76">
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">-</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">Dorsal side uniformly brownish; pronotal hind angles nearly completely rounded; internal sac of aedeagus with scaled area half length of median lobe; Japan, Korea, East China</td>
<td colspan="1" pageId="75" pageNumber="76" rowspan="1">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Orionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis italics="true" pageId="75" pageNumber="76">Orionella lewisii</emphasis>
</taxonomicName>
(Bates)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>