294 lines
19 KiB
XML
294 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.56927" ID-GBIF-Dataset="7a0ded71-ddb1-49e5-9c2c-10c01fd03a1f" ID-PMC="PMC7990858" ID-Pensoft-Pub="1313-2970-1024-157" ID-Pensoft-UUID="086BAB631B8D5B888571137F0CEFF686" ID-PubMed="33786006" ID-ZooBank="FDD37EA991214385B67D51AD313CB49E" ModsDocID="1313-2970-1024-157" checkinTime="1615979483707" checkinUser="pensoft" docAuthor="Nestor-Arriola, Jorge Ismael, Toledo-Hernandez, Victor Hugo, Solis, Angel, Gonzalez, Guillermo & Vetrovec, Jaroslav" docDate="2021" docId="F7B73900624C5E39A84B9E769E45EEC4" docLanguage="en" docName="ZooKeys 1024: 157-196" docOrigin="ZooKeys 1024" docPubDate="2021-03-16" docSource="http://dx.doi.org/10.3897/zookeys.1024.56927" docTitle="Brachiacantha tica Nestor-Arriola, Toledo-Hernandez & Solis 2021, sp. nov." docType="treatment" docUuid="8180E24F-BD61-4463-80C1-0E6D19F8DFC4" docUuidSource="ZooBank" docVersion="5" id="086BAB631B8D5B888571137F0CEFF686" lastPageNumber="157" masterDocId="086BAB631B8D5B888571137F0CEFF686" masterDocTitle="The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America" masterLastPageNumber="196" masterPageNumber="157" pageNumber="157" updateTime="1668149905347" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nestor-Arriola, Jorge Ismael</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2394-1586</mods:nameIdentifier>
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<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
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<mods:nameIdentifier type="email">jorge.nestorarr@uaem.edu.mx</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Toledo-Hernandez, Victor Hugo</mods:namePart>
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<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Solis, Angel</mods:namePart>
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<mods:affiliation>Departamento de Historia Natural, Museo Nacional de Costa Rica. Santo Domingo de Heredia, Costa Rica</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gonzalez, Guillermo</mods:namePart>
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<mods:affiliation>La Reina, Santiago, Chile</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vetrovec, Jaroslav</mods:namePart>
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<mods:affiliation>Buzulucka, Hradec Kralove, Ceska Republika</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-03-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1024</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>157</mods:start>
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<mods:end>196</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1024.56927</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.56927</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1024-157</mods:identifier>
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<mods:identifier type="ZooBank">FDD37EA991214385B67D51AD313CB49E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">086BAB631B8D5B888571137F0CEFF686</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="180585844" LSID="urn:lsid:zoobank.org:act:8180E24F-BD61-4463-80C1-0E6D19F8DFC4" httpUri="http://treatment.plazi.org/id/F7B73900624C5E39A84B9E769E45EEC4" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection pageId="0" pageNumber="157" type="nomenclature">
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<paragraph pageId="0" pageNumber="157">
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<taxonomicName LSID="http://zoobank.org/8180E24F-BD61-4463-80C1-0E6D19F8DFC4" authority="Nestor-Arriola, Toledo-Hernández & Solís" authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha tica" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="tica" status="sp. nov.">
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Brachiacantha tica Nestor-Arriola,
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<normalizedToken originalValue="Toledo-Hernández">Toledo-Hernandez</normalizedToken>
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&
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<normalizedToken originalValue="Solís">Solis</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="157">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 69, 70" captionStartId="F16" captionText="Figures 69, 70. Brachiacantha tica sp. nov., female 69 Lateral view 70 Protibia." figureDoi="10.3897/zookeys.1024.56927.figures69-70" httpUri="https://binary.pensoft.net/fig/520881" pageId="0" pageNumber="157">Figures 69-70</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="holotype">
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<paragraph pageId="0" pageNumber="157">
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<typeStatus>Holotype</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<materialsCitation ID-GBIF-Occurrence="3063616387" collectingDate="1992-09" collectionCode="MNCR" collectorName="A. Marin" country="Costa Rica" elevation="200" location="Original" municipality="Female" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
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<collectingMunicipality>Female</collectingMunicipality>
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, pinned.
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<location LSID="urn:lsid:plazi:treatment:F7B73900624C5E39A84B9E769E45EEC4:C553797B68D33DEA1F0A7780C34E319B" country="Costa Rica" municipality="Female" name="Original" stateProvince="Puntarenas">Original</location>
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label: "
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<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
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,
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<collectingRegion country="Costa Rica" name="Puntarenas">PUNTARENAS</collectingRegion>
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,
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<normalizedToken originalValue="Península">Peninsula</normalizedToken>
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de Osa, Rancho Quemado,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
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<date value="1992-09">
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<collectingDate value="1992-09">Sep. 1992</collectingDate>
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</date>
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, col.
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<collectorName>
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A.
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<normalizedToken originalValue="Marín">Marin</normalizedToken>
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</collectorName>
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, L-S 292500_511000" (
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="paratypes">
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<paragraph pageId="0" pageNumber="157">
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<typeStatus>Paratypes</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<materialsCitation ID-GBIF-Occurrence="3063616339" collectingDate="1993-09-10" collectingDateMax="1993-10-10" collectingDateMin="1993-09-10" collectionCode="MNCR" country="Costa Rica" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
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<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
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•
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<specimenCount type="female">1♀</specimenCount>
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;
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<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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, same data than the
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<typeStatus>holotype</typeStatus>
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;
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<date value="1993-09-10" valueMax="1993-10-10" valueMin="1993-09-10">
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<collectingDate value="1993-09-10" valueMax="1993-10-10" valueMin="1993-09-10">10 Sep.-10 Oct. 1993</collectingDate>
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</date>
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(
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3063616316" collectingDate="1992-12" collectionCode="MNCR" collectorName="F. A. Quesada" country="Costa Rica" elevation="200" location="Original" municipality="Female" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
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<specimenCount type="female">1♀</specimenCount>
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; same data as holotype;
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<date value="1992-12">
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<collectingDate value="1992-12">Dec. 1992</collectingDate>
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</date>
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;
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<collectorName>F. A. Quesada</collectorName>
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leg.; (
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3063616359" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MNCR" collectorName="P. N. Corcovado" country="Costa Rica" location="Rio Claro" specimenCount="2" specimenCount-female="2" stateProvince="Puntarenas" typeStatus="paratype">
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<specimenCount type="female">2♀</specimenCount>
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;
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<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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,
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<collectorName>P. N. Corcovado</collectorName>
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, send.
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<location LSID="urn:lsid:plazi:treatment:F7B73900624C5E39A84B9E769E45EEC4:FB467B4E62DECAE9CF9614EAD951FD6F" country="Costa Rica" name="Rio Claro" stateProvince="Puntarenas">Rio Claro</location>
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; 1991; L-S 508300_270500 (
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3063616343" collectingDate="1991-03" collectingDateMax="1991-06-31" collectingDateMin="1991-03" collectionCode="MNCR" collectorName="P. N. Corcovado & Est. Sirena" country="Costa Rica" elevation="50" location="Puntarenas" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
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<specimenCount type="female">1♀</specimenCount>
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;
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<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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,
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<collectorName>P. N. Corcovado</collectorName>
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,
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<collectorName>Est. Sirena</collectorName>
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;
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<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
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<elevation metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0-100 m</elevation>
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</quantity>
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;
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<date value="1991-03" valueMax="1991-06-31" valueMin="1991-03-00">
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<collectingDate value="1991-03" valueMax="1991-06-31" valueMin="1991-03-00">Mar.-June 1991</collectingDate>
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</date>
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; L-S 270500_508300 (
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3063616333" collectingDate="1991-06-17" collectingDateMax="1991-09-04" collectingDateMin="1991-06-17" collectionCode="MNCR" collectorName="P. N. Corcovado & Est. Sirena" country="Costa Rica" elevation="50" location="Puntarenas" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
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<specimenCount type="female">1♀</specimenCount>
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; same data,
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<date value="1991-06-17" valueMax="1991-09-04" valueMin="1991-06-17">
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<collectingDate value="1991-06-17" valueMax="1991-09-04" valueMin="1991-06-17">17 June-4 Sep. 1991</collectingDate>
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</date>
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;
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<collectingMethod>malaise trap</collectingMethod>
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; (
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<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="diagnosis">
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<paragraph pageId="0" pageNumber="157">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="157">Dorsal color black; pronotum anterior angles pale yellow; each elytron with three pale yellow spots, one near the scutellar shield and two near the apex, no humeral or mid-lateral spots. Sponda large and conspicuous.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="description">
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<paragraph pageId="0" pageNumber="157">Description.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<emphasis bold="true" pageId="0" pageNumber="157">
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<emphasis italics="true" pageId="0" pageNumber="157">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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Female. Length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.3" unit="mm" value="2.3">2.3 mm</quantity>
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, width
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.7" unit="mm" value="1.7">1.7 mm</quantity>
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; body oval, convex. Dorsal color black except head pale yellow; pronotum with lateral angles pale yellow; elytron with three pale yellow oval spots: scutellar shield spot small, small discal spot in the apical half of elytra, sub-apical spot large and oval (Fig.
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<figureCitation captionStart="Figures 69, 70" captionStartId="F16" captionText="Figures 69, 70. Brachiacantha tica sp. nov., female 69 Lateral view 70 Protibia." figureDoi="10.3897/zookeys.1024.56927.figures69-70" httpUri="https://binary.pensoft.net/fig/520881" pageId="0" pageNumber="157">69</figureCitation>
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). Ventral surface with head, prosternum, metaventrite and abdomen dark brown; legs, mouthparts and antennae yellow. Head punctures small, separated by their diameter, each puncture as large as an eye facet; pronotal punctures larger than head punctures, separated by their diameter; elytral punctures as large as those on pronotum, separated by their diameter; metaventral punctures larger than elytral punctures, separated by their diameter to two diameters. Clypeus slightly emarginated. Carinae on prosternal process straight, larger than half prosternum, convergent, joined near the anterior margin of prosternum but not touching it. Protibia not flanged, basal tooth small, sponda obvious (Fig.
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<figureCitation captionStart="Figures 69, 70" captionStartId="F16" captionText="Figures 69, 70. Brachiacantha tica sp. nov., female 69 Lateral view 70 Protibia." figureDoi="10.3897/zookeys.1024.56927.figures69-70" httpUri="https://binary.pensoft.net/fig/520881" pageId="0" pageNumber="157">70</figureCitation>
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). Basal abdominal ventrite without setal tuft. Abdomen with postcoxal line on basal abdominal ventrite curved, only touching the posterior ventrite margin, ventrite with sparse, long pubescence and small, dense punctures. Ventrites II-VI pubescent throughout, punctures fine, dense. Genitalia not examined.
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</paragraph>
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<paragraph pageId="0" pageNumber="157">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="variation">
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<paragraph pageId="0" pageNumber="157">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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Length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.1-2.6mm</quantity>
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. Width
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5-1.8 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="etymology">
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<paragraph pageId="0" pageNumber="157">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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The species name is a popular name for the inhabitants of
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<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="type locality">
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<paragraph pageId="0" pageNumber="157">
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<typeStatus>Type</typeStatus>
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locality.
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</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<materialsCitation ID-GBIF-Occurrence="3063616337" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Costa Rica" latitude="8.681911" location="Rancho Quemado" longLatPrecision="1" longitude="-83.559395" municipality="Peninsula de Osa" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
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<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
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,
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<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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:
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<collectingMunicipality>
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<normalizedToken originalValue="Península">Peninsula</normalizedToken>
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de Osa
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</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:F7B73900624C5E39A84B9E769E45EEC4:4484F33943DBABFE4EABD91C952AF96D" country="Costa Rica" latitude="8.681911" longLatPrecision="1" longitude="-83.559395" municipality="Peninsula de Osa" name="Rancho Quemado" stateProvince="Puntarenas">Rancho Quemado</location>
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,
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<geoCoordinate degrees="8" direction="north" minutes="40" orientation="latitude" precision="1" seconds="54.88" value="8.681911">8°40'54.88"N</geoCoordinate>
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,
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<geoCoordinate degrees="83" direction="west" minutes="33" orientation="longitude" precision="1" seconds="33.81" value="-83.559395">83°33'33.81"W</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="distribution">
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<paragraph pageId="0" pageNumber="157">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="discussion">
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<paragraph pageId="0" pageNumber="157">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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Although having three spots on each elytron is common in several
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha</emphasis>
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</taxonomicName>
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species, the particular pattern of
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<taxonomicName lsidName="B. tica" pageId="0" pageNumber="157" rank="species" species="tica">
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<emphasis italics="true" pageId="0" pageNumber="157">B. tica</emphasis>
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</taxonomicName>
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, with one spot in the basal half of the elytron and two in the apical half, is not shared with any other species of the genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |