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<document id="E4BA7B2C8187B285CBE65ECE9E0B3D4C" ID-DOI="10.5852/ejt.2023.915.2379" ID-ISSN="2118-9773" ID-Zenodo-Dep="10410126" ID-ZooBank="78346F83-1177-4EAE-AFFD-D4DDCC1BFB83" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703073859257" checkinUser="plazi" docAuthor="Balkenohl, Michael" docDate="2023" docId="4A6487ECCF0BFFF2FE32FA8BFD972F79" docLanguage="en" docName="EJT.2023.915.1-57.pdf" docOrigin="European Journal of Taxonomy 915 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2379/10451" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sulciclivina attenuata subsp. semireticulata Balkenohl 2023, subsp. nov." docType="treatment" docUuid="02E61455-C50B-4400-AABE-83F99A14C256" docUuidSource="ZooBank" docVersion="2" lastPageNumber="14" masterDocId="B65DFF94CF07FFFFFFC5FFC1FF8F2A5D" masterDocTitle="Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="13" updateTime="1703074388710" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B3CEEDDA7DBCB51107B7EA7AD3E68555">Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)</mods:title>
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<mods:namePart id="04865903D1ACA651FE0BBEB08EC6B916">Balkenohl, Michael</mods:namePart>
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<mods:date id="1C26DA5718D8B3183A2770239493F1DE">2023</mods:date>
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<treatment id="4A6487ECCF0BFFF2FE32FA8BFD972F79" ID-DOI="http://doi.org/10.5281/zenodo.10410130" ID-Zenodo-Dep="10410130" LSID="urn:lsid:zoobank.org:act:02E61455-C50B-4400-AABE-83F99A14C256" httpUri="http://treatment.plazi.org/id/4A6487ECCF0BFFF2FE32FA8BFD972F79" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="8AD76571CF0BFFF3FE32FA8BFB4C2FD6" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="C27236FACF0BFFF3FE32FA8BFBB22F3B" blockId="12.[369,1219,1354,1456]" box="[503,1085,1354,1382]" pageId="12" pageNumber="13">
<heading id="993A8196CF0BFFF3FE32FA8BFBB22F3B" box="[503,1085,1354,1382]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="05CD4D79CF0BFFF3FE32FA8BFC3D2F39" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[503,946,1354,1381]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="attenuata" status="subsp. nov." subSpecies="semireticulata">
<emphasis id="F0B9EAE8CF0BFFF3FE32FA8BFC3D2F39" bold="true" box="[503,946,1354,1381]" italics="true" pageId="12" pageNumber="13">Sulciclivina attenuata semireticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF0BFFF3FC7FFA8AFBB22F3B" box="[954,1085,1355,1382]" pageId="12" pageNumber="13" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="C27236FACF0BFFF3FEB4FAB1FB4C2FD6" blockId="12.[369,1219,1354,1456]" box="[369,1219,1392,1419]" pageId="12" pageNumber="13">
<uri id="B65C3AF8CF0BFFF3FEB4FAB1FB4C2FD6" box="[369,1219,1392,1419]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="8AD76571CF0BFFF3FD0CFA54FCE52FED" box="[713,874,1429,1456]" pageId="12" pageNumber="13" type="description">
<paragraph id="C27236FACF0BFFF3FD0CFA54FCE52FED" blockId="12.[369,1219,1354,1456]" box="[713,874,1429,1456]" pageId="12" pageNumber="13">
<figureCitation id="5AF62A7FCF0BFFF3FD0CFA54FC9B2FF2" box="[713,788,1429,1455]" captionStart="Figs 411" captionStartId="11.[189,241,1857,1883]" captionTargetBox="[204,1380,703,1763]" captionTargetId="figure-113@11.[189,1398,670,1822]" captionTargetPageId="11" captionText="Figs 411. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 45. S. attenuata attenuata (Herbst, 1806). 4. ♀, lectotype (MFNB). 5. ♂ from Swat (CMBB). 6. S. attenuata semireticulata subsp. nov., ♂, holotype (MNHN). 7. S. bhamoensis (Bates, 1892), ♀, paralectotype (RBINS). 8. S. mikirensis sp. nov., ♀, holotype (NHMB). 9. S. basiangusta sp. nov., ♂, holotype (NHMB). 10. S. coxisetosa sp. nov., ♀, holotype (NHMB). 11. S. splendida sp. nov., ♂, holotype (CADW)." figureDoi="http://doi.org/10.5281/zenodo.10410132" httpUri="https://zenodo.org/record/10410132/files/figure.png" pageId="12" pageNumber="13">Figs 6</figureCitation>
, 27, 53
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF0BFFF3FF78FA19FBCF2CCF" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="C27236FACF0BFFF3FF78FA19FEB92FAF" blockId="12.[189,310,1496,1522]" box="[189,310,1496,1522]" pageId="12" pageNumber="13">
<heading id="993A8196CF0BFFF3FF78FA19FEB92FAF" bold="true" box="[189,310,1496,1522]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="F0B9EAE8CF0BFFF3FF78FA19FEB92FAF" bold="true" box="[189,310,1496,1522]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF0BFFF3FF78F9C8FBCF2CCF" blockId="12.[189,1399,1545,1682]" pageId="12" pageNumber="13">The subspecies differs from the nominotypical subspecies mainly by interval seven hanging over laterally behind the humerus and covering interval eight nearly completely in dorsal view. In addition, the reticulation on interval eight is indistinct in general and covers only half of the width of the interval, and the lateral lobe of the clypeus is completely fused with the clypeal wing.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF0BFFF3FF78F97AFDEA2D77" pageId="12" pageNumber="13" type="etymology">
<paragraph id="C27236FACF0BFFF3FF78F97AFECB2C88" blockId="12.[189,324,1723,1749]" box="[189,324,1723,1749]" pageId="12" pageNumber="13">
<heading id="993A8196CF0BFFF3FF78F97AFECB2C88" bold="true" box="[189,324,1723,1749]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="F0B9EAE8CF0BFFF3FF78F97AFECB2C88" bold="true" box="[189,324,1723,1749]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF0BFFF3FF78F92AFDEA2D77" blockId="12.[189,1398,1771,1835]" pageId="12" pageNumber="13">The name refers to the reticulation on interval eight of the elytron which covers half of the interval by contrast to the nominate subspecies.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF0BFFF2FF78F895FE582B30" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="C27236FACF0BFFF3FF78F895FEE32D33" blockId="12.[189,364,1876,1902]" box="[189,364,1876,1902]" pageId="12" pageNumber="13">
<heading id="993A8196CF0BFFF3FF78F895FEE32D33" bold="true" box="[189,364,1876,1902]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="F0B9EAE8CF0BFFF3FF78F895FEE32D33" bold="true" box="[189,364,1876,1902]" pageId="12" pageNumber="13">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF0BFFF3FF2FF845FED32DC3" blockId="12.[189,1399,1924,2025]" box="[234,348,1924,1950]" pageId="12" pageNumber="13">
<emphasis id="F0B9EAE8CF0BFFF3FF2FF845FED32DC3" bold="true" box="[234,348,1924,1950]" pageId="12" pageNumber="13">
<typeStatus id="1D768858CF0BFFF3FF2FF845FED32DC3" box="[234,348,1924,1950]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C27236FACF0BFFF3FF78F868FE912DB4" blockId="12.[189,1399,1924,2025]" pageId="12" pageNumber="13">
<materialsCitation id="72A53CA7CF0BFFF3FF78F868FE912DB4" collectionCode="MNHN" country="Indonesia" location="Reiche" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur" typeStatus="holotype">
<collectingCountry id="BADA766ACF0BFFF3FF78F868FED22D9E" box="[189,349,1961,1987]" name="Indonesia" pageId="12" pageNumber="13">INDONESIA</collectingCountry>
<specimenCount id="D4CBFD73CF0BFFF3FEB2F86BFE1F2D99" box="[375,400,1962,1988]" count="1" pageId="12" pageNumber="13" type="male"></specimenCount>
; with two yellow labels, both handwritten in black ink and data: “
<collectingRegion id="0009F818CF0BFFF3FB63F868FB512D9E" box="[1190,1246,1961,1987]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="13">Java</collectingRegion>
” / “
<location id="C7126021CF0BFFF3FACBF868FAE82D9E" LSID="urn:lsid:plazi:treatment:4A6487ECCF0BFFF2FE32FA8BFD972F79:C7126021CF0BFFF3FACBF868FAE82D9E" box="[1294,1383,1961,1987]" country="Indonesia" name="Reiche" pageId="12" pageNumber="13" stateProvince="Jawa Timur">Reiche</location>
”;
<collectionCode id="A4DCAE3FCF0BFFF3FF78F80EFE942DB4" box="[189,283,1999,2025]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="13">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FEC8FE582B30" blockId="13.[189,1398,265,365]" pageId="13" pageNumber="14">
<emphasis id="F0B9EAE8CF0AFFF2FF78FEC8FEA42B7E" bold="true" box="[189,299,265,291]" pageId="13" pageNumber="14">Remark.</emphasis>
The left front leg and in the right front leg the four terminal tarsomeres are missing. The right elytron and the pronotum show pinholes and the pronotum a substantial crack. The specimen is very brittle and rotten inside.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF0AFFF2FF78FE54FC832EEC" pageId="13" pageNumber="14" type="description">
<paragraph id="C27236FACF0AFFF2FF78FE54FEC12BF2" blockId="13.[189,334,405,431]" box="[189,334,405,431]" pageId="13" pageNumber="14">
<heading id="993A8196CF0AFFF2FF78FE54FEC12BF2" bold="true" box="[189,334,405,431]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="F0B9EAE8CF0AFFF2FF78FE54FEC12BF2" bold="true" box="[189,334,405,431]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FE07FF672858" blockId="13.[188,1399,454,517]" pageId="13" pageNumber="14">
MEASUREMENTS.
<typeStatus id="1D768858CF0AFFF2FE47FE07FE7C2BBD" box="[386,499,454,480]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
. Body length
<quantity id="05359B1FCF0AFFF2FD57FE07FD722BBC" box="[658,765,454,481]" metricMagnitude="-3" metricUnit="m" metricValue="8.83" pageId="13" pageNumber="14" unit="mm" value="8.83">8.83 mm</quantity>
, width
<quantity id="05359B1FCF0AFFF2FC95FE07FC342BBC" box="[848,955,454,481]" metricMagnitude="-3" metricUnit="m" metricValue="2.49" pageId="13" pageNumber="14" unit="mm" value="2.49">2.49 mm</quantity>
; L/W of pronotum 1.01; L/W of elytra 1.9.
</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FDEFFA8A2815" blockId="13.[189,1285,558,584]" box="[189,1285,558,584]" pageId="13" pageNumber="14">The subspecies differs from the nominotypical subspecies mainly in the following characters:</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FDB0FC7928D6" blockId="13.[189,1014,625,651]" box="[189,1014,625,651]" pageId="13" pageNumber="14">HEAD. Clypeal wing completely fused with lateral lobe of the clypeus.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FD75FA8F28AE" blockId="13.[189,1397,692,756]" pageId="13" pageNumber="14">PRONOTUM. Lateral margin of pronotum nearly straight, slightly concave behind anterior setigerous puncture. Anterior transverse line more distinctly punctured than in the nominate subspecies.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FCDDFE7C29DD" blockId="13.[188,1400,796,896]" pageId="13" pageNumber="14">ELYTRON. Interval seven hanging over laterally behind the humerus and covering interval eight nearly completely in dorsal view. The reticulation on interval eight covers only half of the width of the interval and is generally indistinct.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FC68FCD9299E" blockId="13.[189,854,937,963]" box="[189,854,937,963]" pageId="13" pageNumber="14">HIND WINGS. In the HT, the alae are completely missing.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FC2DFB7C2E5A" blockId="13.[189,1267,1004,1031]" box="[189,1267,1004,1031]" pageId="13" pageNumber="14">LOWER SURFACE. Sternites of abdomen less densely punctured including the apical segment.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FBEEFE302E33" blockId="13.[188,1398,1071,1134]" pageId="13" pageNumber="14">MALE GENITALIA (Fig. 27). The median lobe distinctly and long acuminated in the apical third, the apical lamella parallel sided.</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FB56FC832EEC" blockId="13.[189,780,1175,1202]" box="[189,780,1175,1202]" pageId="13" pageNumber="14">FEMALE GONOCOXITE AND EPIPLEURITE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF0AFFF2FF78FB1BFD972F79" box="[189,536,1242,1317]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="C27236FACF0AFFF2FF78FB1BFED92EA9" blockId="13.[189,342,1242,1268]" box="[189,342,1242,1268]" pageId="13" pageNumber="14">
<heading id="993A8196CF0AFFF2FF78FB1BFED92EA9" bold="true" box="[189,342,1242,1268]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="F0B9EAE8CF0AFFF2FF78FB1BFED92EA9" bold="true" box="[189,342,1242,1268]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF0AFFF2FF78FACAFD972F79" blockId="13.[189,536,1290,1317]" box="[189,536,1290,1317]" pageId="13" pageNumber="14">
Known from
<collectingRegion id="0009F818CF0AFFF2FE9EFACAFE1D2F78" box="[347,402,1291,1317]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="14">Java</collectingRegion>
,
<collectingCountry id="BADA766ACF0AFFF2FE5BFACBFD9B2F79" box="[414,532,1290,1316]" name="Indonesia" pageId="13" pageNumber="14">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>