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<mods:title id="C224BEE90112B687805C891B53F540C9">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<paragraph id="A230B37B4B75FFB9E7B8FB63FE04FB76" blockId="49.[151,463,1230,1293]" box="[151,463,1230,1257]" pageId="49" pageNumber="50">
<heading id="F97804174B75FFB9E7B8FB63FE04FB76" bold="true" box="[151,463,1230,1257]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
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<taxonomicName id="658FC8F84B75FFB9E7B8FB63FEA1FB76" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[151,362,1230,1257]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B75FFB9E7B8FB63FEA1FB76" bold="true" box="[151,362,1230,1257]" italics="true" pageId="49" pageNumber="50">Alpheus nomurai</emphasis>
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<taxonomicNameLabel id="8BC8D2124B75FFB9E65DFB62FE04FB76" box="[370,463,1231,1257]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="A230B37B4B75FFB9E7B8FB5EFEACFA92" blockId="49.[151,463,1230,1293]" box="[151,359,1266,1293]" pageId="49" pageNumber="50">
(
<figureCitation id="3AB4AFFE4B75FFB9E7B0FB5EFEE3FA92" box="[159,296,1266,1293]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStart-3="FIGURE 28" captionStartId-0="50.[151,250,1940,1964]" captionStartId-1="51.[151,250,1813,1837]" captionStartId-2="52.[151,250,1818,1842]" captionStartId-3="55.[151,250,1814,1838]" captionTargetBox-0="[151,1435,669,1917]" captionTargetBox-1="[167,1425,193,1775]" captionTargetBox-2="[162,1424,195,1781]" captionTargetBox-3="[151,1435,181,1790]" captionTargetId-0="figure-226@50.[151,1435,669,1917]" captionTargetId-1="figure-15@51.[151,1436,181,1789]" captionTargetId-2="figure-21@52.[151,1436,181,1794]" captionTargetId-3="figure-15@55.[151,1435,181,1790]" captionTargetPageId-0="50" captionTargetPageId-1="51" captionTargetPageId-2="52" captionTargetPageId-3="55" captionText-0="FIGURE 25. Alpheus nomurai sp. nov.: holotype, male (cl 8.8 mm) from Tanabe, Wakayama, Japan (MNHN-IU-2018-5590); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) cheliped, lateral view; D—same, mesial view; E— minor (right) cheliped, mesial view; F—same, dorsolateral view; G—same, lateral view. Refer to A for scale." captionText-1="FIGURE 26. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D— antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—fifth pereiopod, lateral view; J—same, propodus and dactylus, mesial view; K—second pleopod, appendices masculina and interna, anterior view; L—left uropod, dorsal view." captionText-2="FIGURE 27. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588) [A–G]; paratype, ovigerous female (cl 8.8 mm) from the same locality (MNHN-IU-2018-5744) [H]; paratype, ovigerous female (cl 11.9 mm) from Shimanto River, Japan (MNHN-IU-2018-5589) [I]; A—major (right) cheliped, chela, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor cheliped, lateral view; same, carpus and chela, lateral view." captionText-3="FIGURE 28. Alpheus nomurai sp. nov.: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D)." figureDoi-0="http://doi.org/10.5281/zenodo.7912372" figureDoi-1="http://doi.org/10.5281/zenodo.7912376" figureDoi-2="http://doi.org/10.5281/zenodo.7912380" figureDoi-3="http://doi.org/10.5281/zenodo.7912382" httpUri-0="https://zenodo.org/record/7912372/files/figure.png" httpUri-1="https://zenodo.org/record/7912376/files/figure.png" httpUri-2="https://zenodo.org/record/7912380/files/figure.png" httpUri-3="https://zenodo.org/record/7912382/files/figure.png" pageId="49" pageNumber="50">Figs. 25–28</figureCitation>
, 
<figureCitation id="3AB4AFFE4B75FFB9E61CFB5FFE94FA92" box="[307,351,1266,1293]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="49" pageNumber="50">51F</figureCitation>
)
</paragraph>
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<paragraph id="A230B37B4B75FFB9E7B8FA9AFCD2FAF1" blockId="49.[151,1437,1335,1574]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="829F94554B75FFB9E7B8FA9AFA56FAD0" box="[151,1437,1335,1359]" pageId="49" pageNumber="50">
<taxonomicName id="658FC8F84B75FFB9E7B8FA9AFE97FAD0" ID-CoL="84Z5Y" authority="Dôtu 1961" box="[151,348,1335,1359]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bis-incisus">
<emphasis id="90FB6F694B75FFB9E7B8FA9AFE97FAD0" box="[151,348,1335,1359]" italics="true" pageId="49" pageNumber="50">Alpheus bis-incisus</emphasis>
</taxonomicName>
(or 
<taxonomicName id="658FC8F84B75FFB9E6A7FA9AFDC2FAD0" ID-CoL="84W6W" authorityName="De Haan" authorityYear="1844" box="[392,521,1335,1359]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bisincisus">
<emphasis id="90FB6F694B75FFB9E6A7FA9AFDC2FAD0" box="[392,521,1335,1359]" italics="true" pageId="49" pageNumber="50">A. bisincisus</emphasis>
</taxonomicName>
).— 
<treatmentCitation id="232E956A4B75FFB9E500FA9AFD1FFAD0" author="Dotu, Y." box="[559,724,1335,1359]" page="136" pageId="49" pageNumber="50" year="1961">
<bibRefCitation id="C61ECE8A4B75FFB9E500FA9AFD1FFAD0" author="Dotu, Y." box="[559,724,1335,1359]" pageId="49" pageNumber="50" pagination="133 - 139" refId="ref71162" refString="Dotu, Y. (1961) The bionomics and life history of the gobioid fish, Apocryptodon bleekeri (Day). Bulletin of the Faculty of Fisheries, Nagasaki University, 10, 133 - 139, pl. 28. [in Japanese]" type="journal article" year="1961">Dôtu 1961: 136</bibRefCitation>
</treatmentCitation>
, fig. 1; 
<treatmentCitation id="232E956A4B75FFB9E40BFA9AFBD8FAD0" author="Kim, H. S. &amp; Park, K. B." box="[804,1043,1335,1359]" page="198" pageId="49" pageNumber="50" year="1972">
<bibRefCitation id="C61ECE8A4B75FFB9E40BFA9AFBD8FAD0" author="Kim, H. S. &amp; Park, K. B." box="[804,1043,1335,1359]" pageId="49" pageNumber="50" pagination="1 - 6" refId="ref72431" refString="Kim, H. S. &amp; Park, K. B. (1972) Faunal studies on the macrurans in Korea. In: Floral studies on some taxa of plants and faunal studies on some taxa of animals in Korea. R- 72 - 82. Ministry of Science and Technology, pp. 185 - 216, pls. 1 - 6. [in Korean with English summary]" type="book chapter" year="1972">Kim &amp; Park 1972: 198</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B75FFB9E331FA9AFB77FAD0" author="Kim, H. S." box="[1054,1212,1335,1359]" page="142" pageId="49" pageNumber="50" year="1976">
<bibRefCitation id="C61ECE8A4B75FFB9E331FA9AFB77FAD0" author="Kim, H. S." box="[1054,1212,1335,1359]" pageId="49" pageNumber="50" pagination="131 - 152" refId="ref72351" refString="Kim, H. S. (1976) A checklist of the Macrura (Crustacea, Decapoda) of Korea. Proceedings of the College of Natural Science, Seoul National University, 1, 131 - 152." type="journal article" year="1976">Kim 1976: 142</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B75FFB9E3E8FA9AFAAFFAD0" author="Kim, H. S." box="[1223,1380,1335,1359]" page="247" pageId="49" pageNumber="50" year="1977">
<bibRefCitation id="C61ECE8A4B75FFB9E3E8FA9AFAAFFAD0" author="Kim, H. S." box="[1223,1380,1335,1359]" pageId="49" pageNumber="50" refId="ref72391" refString="Kim, H. S. (1977) Macrura. Illustrated Flora and Fauna of Korea. Vol. 19. Samwha Publishing Co., Seoul, 414 pp., 56 pls. [in Korean]" type="book" year="1977">Kim 1977: 247</bibRefCitation>
</treatmentCitation>
, figs.
</treatmentCitationGroup>
100, 101, pl. 24, fig. 44 [not 
<taxonomicName id="658FC8F84B75FFB9E6C4FAFBFCC4FAF1" ID-CoL="84W6W" authority="De Haan, 1844" authorityName="De Haan" authorityYear="1844" box="[491,783,1366,1390]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bisincisus">
<emphasis id="90FB6F694B75FFB9E6C4FAFBFDA0FAF1" box="[491,619,1366,1390]" italics="true" pageId="49" pageNumber="50">A. bisincisus</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E55EFAFBFCC4FAF1" author="De Haan, W." box="[625,783,1366,1390]" pageId="49" pageNumber="50" pagination="1 - 55" refId="ref70613" refString="De Haan, W. (1844) [1833 - 1850] Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suspecto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Apud Auctorem, Lugduni-Batavorum, pp. i - xxxi + ix - xvi + 1 - 243, pls. A - J + L - Q + 1 - 55." type="book chapter" year="1844">De Haan, 1844</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="A230B37B4B75FFB9E7B8FAD9FBF2FA12" blockId="49.[151,1437,1335,1574]" box="[151,1081,1396,1421]" pageId="49" pageNumber="50">
<emphasis id="90FB6F694B75FFB9E7B8FAD9FE4BFA13" box="[151,384,1396,1420]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="658FC8F84B75FFB9E7B8FAD9FEA9FA13" ID-CoL="84W7P" authorityName="sensu De Man" authorityYear="1897" box="[151,354,1396,1420]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="euphrosyne">Alpheus euphrosyne</taxonomicName>
.—
</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E6AFFAD8FDB4FA12" author="Jeng, M. S. &amp; Chang, K. H." box="[384,639,1396,1421]" pageId="49" pageNumber="50" pagination="241 - 256" refId="ref71966" refString="Jeng, M. S. &amp; Chang, K. H. (1985) Snapping shrimps (Crustacea: Decapoda: Alpheidae) of Taiwan. Bulletin of Institute of Zoology, Academica Sinica, 24, 241 - 256." type="journal article" year="1985">Jeng &amp; Chang 1985: 248</bibRefCitation>
, fig. 23 [not 
<taxonomicName id="658FC8F84B75FFB9E42EFAD8FBE4FA12" ID-CoL="84W7P" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[769,1071,1396,1421]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B75FFB9E42EFAD8FC59FA13" box="[769,914,1396,1420]" italics="true" pageId="49" pageNumber="50">A. euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E4B7FAD8FBE4FA12" author="De Man, J. G." box="[920,1071,1397,1421]" pageId="49" pageNumber="50" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="A230B37B4B75FFB9E7B8FA3EFD0FFA55" blockId="49.[151,1437,1335,1574]" pageId="49" pageNumber="50">
<emphasis id="90FB6F694B75FFB9E7B8FA3EFE37FA34" box="[151,508,1427,1451]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="658FC8F84B75FFB9E7B8FA3EFE14FA34" ID-CoL="85LNR" authorityName="Yaldwyn" authorityYear="1971" box="[151,479,1427,1451]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">Alpheus euphrosyne richardsoni</taxonomicName>
.—
</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E6D3FA3EFD6BFA34" author="Miya, Y." box="[508,672,1427,1451]" pageId="49" pageNumber="50" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995: 275</bibRefCitation>
, fig. 2; 
<bibRefCitation id="C61ECE8A4B75FFB9E5C2FA39FC15FA34" author="Yang, B. P. &amp; Kim, C. H." box="[749,990,1427,1452]" pageId="49" pageNumber="50" pagination="106 - 114" refId="ref75154" refString="Yang, B. P. &amp; Kim, C. H. (1996) Zoeal stages of Alpheus euphrosyne richardsoni Yaldwyn, 1971 (Decapoda: Macrura: Alpheidae) reared in the laboratory. Korean Journal of Zoology, 39, 106 - 114." type="journal article" year="1996">Yang &amp; Kim 1996: 106</bibRefCitation>
, figs. 1-3; 
<bibRefCitation id="C61ECE8A4B75FFB9E364FA3EFAC6FA34" author="Hayashi, K. I." box="[1099,1293,1427,1451]" pageId="49" pageNumber="50" pagination="289 - 293" refId="ref71727" refString="Hayashi, K. I. (1998) Prawns, shrimps and lobsters from Japan (101). Family Alpheidae - Genus Alpheus (10). Aquabiology, 117 (20), 289 - 293." type="journal article" year="1998">Hayashi 1998: 292</bibRefCitation>
, figs. 353c, g, 354c, 355c, e [not 
<taxonomicName id="658FC8F84B75FFB9E6ABFA1FFD72FA55" ID-CoL="85LNR" authority="Yaldwyn, 1971" authorityName="Yaldwyn" authorityYear="1971" box="[388,697,1458,1482]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B75FFB9E6ABFA1FFDDEFA55" box="[388,533,1458,1482]" italics="true" pageId="49" pageNumber="50">A. richardsoni</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E534FA1FFD72FA55" author="Yaldwyn, J. C." box="[539,697,1458,1482]" pageId="49" pageNumber="50" pagination="85 - 94" refId="ref75115" refString="Yaldwyn, J. C. (1971) Preliminary descriptions of a new genus and twelve new species of natant decapod Crustacea from New Zealand. Records of the Dominion Museum, 7, 85 - 94." type="journal article" year="1971">Yaldwyn, 1971</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="A230B37B4B75FFB9E7B8FA7DFDFDF998" blockId="49.[151,1437,1335,1574]" pageId="49" pageNumber="50">
<emphasis id="90FB6F694B75FFB9E7B8FA7DFE48FA77" box="[151,387,1488,1512]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="658FC8F84B75FFB9E7B8FA7DFEAEFA77" ID-CoL="85LNR" authorityName="Yaldwyn" authorityYear="1971" box="[151,357,1488,1512]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="richardsoni">Alpheus richardsoni</taxonomicName>
.—
</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E6ADFA7DFD98FA76" author="Yoshigou, H." box="[386,595,1488,1513]" pageId="49" pageNumber="50" pagination="221 - 273" refId="ref75338" refString="Yoshigou, H. (2009) A collection of alpheid shrimps (Crustacea; Decapoda; Alpheidae) from the mouth of rivers and anchialine caves of Japan. Miscellaneous Reports of the Hiwa Museum for Natural History, 50, 221 - 273. [in Japanese with English abstract]" type="journal article" year="2009">Yoshigou 2009: 249</bibRefCitation>
, fig. 
<quantity id="65771E9E4B75FFB9E5A7FA7CFD73FA76" box="[648,696,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="49" pageNumber="50" unit="m" value="12.0">12m</quantity>
, 
<quantity id="65771E9E4B75FFB9E5ECFA7CFD38FA76" box="[707,755,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="49" pageNumber="50" unit="m" value="13.0">13m</quantity>
, 
<quantity id="65771E9E4B75FFB9E5D1FA7CFCE6FA76" box="[766,813,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="49" pageNumber="50" unit="m" value="14.0">14m</quantity>
, 
<quantity id="65771E9E4B75FFB9E416FA7DFCA0FA77" box="[825,875,1488,1512]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="50" unit="m" value="15.0">15m</quantity>
,. 
<quantity id="65771E9E4B75FFB9E455FA7DFC62FA77" box="[890,937,1488,1512]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="m" value="16.0">16m</quantity>
, 
<quantity id="65771E9E4B75FFB9E49AFA7CFC28FA76" box="[949,995,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="49" pageNumber="50" unit="m" value="17.0">17m</quantity>
, 
<quantity id="65771E9E4B75FFB9E4C0FA7CFBD4FA76" box="[1007,1055,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="49" pageNumber="50" unit="m" value="18.0">18m</quantity>
, pl. III 1-4; 
<bibRefCitation id="C61ECE8A4B75FFB9E3B2FA7CFA57FA76" author="Koyama, A. &amp; Inui, R. &amp; Sawa, K. &amp; Onikura, N." box="[1181,1436,1488,1513]" pageId="49" pageNumber="50" pagination="131 - 138" refId="ref72628" refString="Koyama, A., Inui, R., Sawa, K. &amp; Onikura, N. (2017) Symbiotic partner specificity and dependency of two gobies (Apocryptodon punctatus and Acentrogobius sp. A) and four alpheid shrimps inhabiting the temperate estuary of southern Japan. Ichthyological Research, 64, 131 - 138. https: // doi. org / 10.1007 / s 10228 - 016 - 0535 - 6" type="journal article" year="2017">
Koyama 
<emphasis id="90FB6F694B75FFB9E3D5FA7CFAE3FA77" box="[1274,1320,1488,1512]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2017: 132
</bibRefCitation>
[not 
<taxonomicName id="658FC8F84B75FFB9E7D9FA42FDE0F998" ID-CoL="85LNR" authority="Yaldwyn, 1971" authorityName="Yaldwyn" authorityYear="1971" box="[246,555,1519,1543]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B75FFB9E7D9FA42FE4CF998" box="[246,391,1519,1543]" italics="true" pageId="49" pageNumber="50">A. richardsoni</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E6A2FA42FDE0F998" author="Yaldwyn, J. C." box="[397,555,1519,1543]" pageId="49" pageNumber="50" pagination="85 - 94" refId="ref75115" refString="Yaldwyn, J. C. (1971) Preliminary descriptions of a new genus and twelve new species of natant decapod Crustacea from New Zealand. Records of the Dominion Museum, 7, 85 - 94." type="journal article" year="1971">Yaldwyn, 1971</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="A230B37B4B75FFB9E7B8F9A3FDA2F9B9" blockId="49.[151,1437,1335,1574]" box="[151,617,1550,1574]" pageId="49" pageNumber="50">
<taxonomicName id="658FC8F84B75FFB9E7B8F9A3FEC7F9BA" box="[151,268,1550,1574]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B75FFB9E7B8F9A3FF22F9B9" box="[151,233,1550,1574]" italics="true" pageId="49" pageNumber="50">Alpheus</emphasis>
sp.
</taxonomicName>
<emphasis id="90FB6F694B75FFB9E623F9A2FEE8F9B9" box="[268,291,1551,1574]" italics="true" pageId="49" pageNumber="50">—</emphasis>
<bibRefCitation id="C61ECE8A4B75FFB9E60CF9A3FDD9F9B9" author="Kirihara, S. &amp; Itani, G. &amp; Nunobe, J. I. &amp; Nomoto, A. &amp; Aldea, K. Q. &amp; Murakami, R. &amp; Sakata, H. &amp; Henmi, Y." box="[291,530,1550,1574]" pageId="49" pageNumber="50" pagination="99 - 102" refId="ref72559" refString="Kirihara, S., Itani, G., Nunobe, J. I., Nomoto, A., Aldea, K. Q., Murakami, R., Sakata, H. &amp; Henmi, Y. (2021) Burrow morphology of an alpheid shrimp at muddy tidal flats in western Japan. Kuroshio Science, 14, 99 - 102." type="journal article" year="2021">
Kirihara 
<emphasis id="90FB6F694B75FFB9E652F9A2FE61F9B9" box="[381,426,1550,1574]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2021: 99
</bibRefCitation>
, fig. 1B.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B75FFBAE7B8F9F9FED7FF66" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="A230B37B4B75FFB9E7B8F9F9FB0FF8BD" blockId="49.[151,1437,1620,2043]" pageId="49" pageNumber="50">
<emphasis id="90FB6F694B75FFB9E7B8F9F9FEF4F9F1" bold="true" box="[151,319,1620,1646]" pageId="49" pageNumber="50">Type material</emphasis>
. 
<materialsCitation id="12E7B9264B75FFB9E662F9F9FCFBF90D" ID-GBIF-Occurrence="4104078533" collectingDate="1993-11-27" collectorName="I. Sakamaki" country="Japan" location="Wakayama" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
<typeStatus id="7D340DD94B75FFB9E662F9F9FE73F9F1" box="[333,440,1620,1646]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
, male (cl 
<quantity id="65771E9E4B75FFB9E507F9F9FD4AF9F0" box="[552,641,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="49" pageNumber="50" unit="mm" value="8.8">8.8 mm</quantity>
, tl 23.0 mm, chl 13.0 mm), MNHN-IU-2018-5590, 
<collectingCountry id="DA98F3EB4B75FFB9E3E3F9F8FAC4F9F0" box="[1228,1295,1621,1647]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
, 
<collectingRegion id="604B7D994B75FFB9E235F9F9FA52F9F1" box="[1306,1433,1620,1646]" country="Japan" name="Wakayama" pageId="49" pageNumber="50">Wakayama</collectingRegion>
, Tanabe, intertidal, leg. 
<collectorName id="0F7AD6AD4B75FFB9E6B7F9D4FDD4F90D" box="[408,543,1656,1683]" pageId="49" pageNumber="50">I. Sakamaki</collectorName>
, 
<date id="D63195BB4B75FFB9E505F9D5FD6CF90D" box="[554,679,1656,1682]" pageId="49" pageNumber="50" value="1993-11-27">
<collectingDate id="C6756C534B75FFB9E505F9D5FD6CF90D" box="[554,679,1656,1682]" pageId="49" pageNumber="50" value="1993-11-27">27.11.1993</collectingDate>
</date>
[SMP-940]
</materialsCitation>
. 
<materialsCitation id="12E7B9264B75FFB9E416F9D4FBDFF929" ID-GBIF-Occurrence="4104078342" collectingDate="1995-08-25" collectorName="K. Nomura" country="Japan" location="Wakayama" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama" typeStatus="paratype">
<typeStatus id="7D340DD94B75FFB9E416F9D4FC61F90C" box="[825,938,1657,1683]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="B48978F24B75FFB9E49AF9D5FC34F90D" box="[949,1023,1656,1682]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B75FFB9E305F9D5FB4FF90D" box="[1066,1156,1656,1683]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="49" pageNumber="50" unit="mm" value="8.5">8.5 mm</quantity>
), MNHN-IU-2018-5588, 
<collectingCountry id="DA98F3EB4B75FFB9E7B8F930FF11F928" box="[151,218,1693,1719]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
, 
<collectingRegion id="604B7D994B75FFB9E7CBF931FEA8F929" box="[228,355,1692,1718]" country="Japan" name="Wakayama" pageId="49" pageNumber="50">Wakayama</collectingRegion>
, Tanabe, intertidal, leg. 
<collectorName id="0F7AD6AD4B75FFB9E540F930FD3AF928" box="[623,753,1693,1719]" pageId="49" pageNumber="50">K. Nomura</collectorName>
, 
<date id="D63195BB4B75FFB9E5D3F931FCB2F928" box="[764,889,1692,1719]" pageId="49" pageNumber="50" value="1995-08-25">
<collectingDate id="C6756C534B75FFB9E5D3F931FCB2F928" box="[764,889,1692,1719]" pageId="49" pageNumber="50" value="1995-08-25">25.08.1995</collectingDate>
</date>
[SMP-1220];
</materialsCitation>
<materialsCitation id="12E7B9264B75FFB9E334F931FC36F945" ID-GBIF-Occurrence="4104078498" collectingDate="1995-08-25" collectorName="K. Nomura" country="Japan" location="Wakayama" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama" typeStatus="paratype">
1 ov. female (cl 
<quantity id="65771E9E4B75FFB9E3FFF931FAE1F929" box="[1232,1322,1692,1718]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="49" pageNumber="50" unit="mm" value="8.8">8.8 mm</quantity>
), MNHN-IU-2018-5744, same collection data as for previous 
<typeStatus id="7D340DD94B75FFB9E5D9F96CFC9DF944" box="[758,854,1729,1755]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
[SMP-1220a]
</materialsCitation>
; 
<materialsCitation id="12E7B9264B75FFB9E323F96DFC19F961" ID-GBIF-Occurrence="4104078525" collectingDate="2002-12-07" collectingDateMax="2011-06-06" collectingDateMin="2002-12-07" collectionCode="OUMNH, ZC" collectorName="K. Nomura" country="Japan" location="Shimanto River" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama" typeStatus="paratype">
<specimenCount id="B48978F24B75FFB9E323F96DFB91F945" box="[1036,1114,1728,1754]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B75FFB9E3A2F96DFB21F945" box="[1165,1258,1728,1754]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="49" pageNumber="50" unit="mm" value="9.6">9.6 mm</quantity>
), 
<collectionCode id="C49E2BBE4B75FFB9E22EF96DFABAF945" box="[1281,1393,1728,1754]" pageId="49" pageNumber="50">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B75FFB9E25BF96DFA52F945" box="[1396,1433,1728,1754]" country="Austria" name="Zoological Collection, University of Vienna" pageId="49" pageNumber="50" type="University or college">ZC</collectionCode>
. 
<date id="D63195BB4B75FFB9E7B8F949FEC2F961" box="[151,265,1764,1790]" pageId="49" pageNumber="50" value="2011-06-06">
<collectingDate id="C6756C534B75FFB9E7B8F949FEC2F961" box="[151,265,1764,1790]" pageId="49" pageNumber="50" value="2011-06-06">2011.06.6</collectingDate>
</date>
, same collection data as for previous 
<typeStatus id="7D340DD94B75FFB9E593F948FCECF960" box="[700,807,1765,1791]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
[SMP-1220b];
</materialsCitation>
<materialsCitation id="12E7B9264B75FFB9E4F4F949FB0BF8BD" ID-GBIF-Occurrence="4104078428" collectingDate="2002-12-07" collectingDateMax="2011-06-06" collectingDateMin="2002-12-07" collectionCode="OUMNH, ZC" collectorName="K. Nomura" country="Japan" location="Shimanto River" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama" typeStatus="paratype">
1 ov. female (cl 
<quantity id="65771E9E4B75FFB9E3B7F949FACAF961" box="[1176,1281,1764,1790]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="49" pageNumber="50" unit="mm" value="11.9">11.9 mm</quantity>
), MNHN-IU-2018-5589, 
<collectingCountry id="DA98F3EB4B75FFB9E630F8A4FEA9F8BC" box="[287,354,1801,1827]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
, Kochi, 
<location id="A750E5A04B75FFB9E690F8A5FDBAF8BD" LSID="urn:lsid:plazi:treatment:2A26026D4B75FFB0E7B8FB63FD50FE4E:A750E5A04B75FFB9E690F8A5FDBAF8BD" box="[447,625,1800,1826]" country="Japan" name="Shimanto River" pageId="49" pageNumber="50">Shimanto River</location>
, intertidal, leg. 
<collectorName id="0F7AD6AD4B75FFB9E430F8A4FC6AF8BC" box="[799,929,1801,1827]" pageId="49" pageNumber="50">K. Nomura</collectorName>
, 
<date id="D63195BB4B75FFB9E484F8A5FBE2F8BD" box="[939,1065,1800,1826]" pageId="49" pageNumber="50" value="2002-12-07">
<collectingDate id="C6756C534B75FFB9E484F8A5FBE2F8BD" box="[939,1065,1800,1826]" pageId="49" pageNumber="50" value="2002-12-07">12.07.2002</collectingDate>
</date>
[SMP-1748]
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B75FFB9E7E8F881FB2DF811" blockId="49.[151,1437,1620,2043]" pageId="49" pageNumber="50">
<emphasis id="90FB6F694B75FFB9E7E8F881FE79F8D9" bold="true" box="[199,434,1836,1862]" pageId="49" pageNumber="50">Additional material</emphasis>
. 
<materialsCitation id="12E7B9264B75FFB9E6EFF880FC47F8F5" ID-GBIF-Occurrence="4104078318" collectingDate="1994-08-08" collectingDateMax="1994-10-17" collectingDateMin="1994-08-08" collectionCode="OUMNH, ZC" collectorName="K. Nomura &amp; A. Kikukawa" country="Japan" location="Okinawa" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<emphasis id="90FB6F694B75FFB9E6EFF880FDCFF8D9" box="[448,516,1837,1862]" italics="true" pageId="49" pageNumber="50">
<collectingCountry id="DA98F3EB4B75FFB9E6EFF880FDCFF8D9" box="[448,516,1837,1862]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
</emphasis>
: 1 ov. female (cl 
<quantity id="65771E9E4B75FFB9E5E9F881FCD5F8D8" box="[710,798,1836,1863]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="49" pageNumber="50" unit="mm" value="9.6">9.6 mm</quantity>
, tl 
<quantity id="65771E9E4B75FFB9E410F881FC6EF8D8" box="[831,933,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="49" pageNumber="50" unit="mm" value="27.5">27.5 mm</quantity>
, chl 12.7), MNHN-IU-2018-5748, OkinawaJima, Sajiki, intertidal, leg. 
<collectorName id="0F7AD6AD4B75FFB9E6F3F8FCFDAAF8F4" box="[476,609,1873,1899]" pageId="49" pageNumber="50">K. Nomura</collectorName>
, 
<date id="D63195BB4B75FFB9E540F8FDFD26F8F5" box="[623,749,1872,1898]" pageId="49" pageNumber="50" value="1994-10-17">
<collectingDate id="C6756C534B75FFB9E540F8FDFD26F8F5" box="[623,749,1872,1898]" pageId="49" pageNumber="50" value="1994-10-17">17.10.1994</collectingDate>
</date>
[SMP-1334];
</materialsCitation>
<materialsCitation id="12E7B9264B75FFB9E4B8F8FDFB29F811" ID-GBIF-Occurrence="4104078367" collectingDate="1994-08-08" collectingDateMax="1994-10-17" collectingDateMin="1994-08-08" collectionCode="OUMNH, ZC" collectorName="K. Nomura &amp; A. Kikukawa" country="Japan" location="Okinawa" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<specimenCount id="B48978F24B75FFB9E4B8F8FDFC2DF8F5" box="[919,998,1872,1898]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B75FFB9E336F8FDFBBCF8F5" box="[1049,1143,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="49" pageNumber="50" unit="mm" value="6.5">6.5 mm</quantity>
), 1 ov. female (
<quantity id="65771E9E4B75FFB9E21EF8FDFA5BF8F5" box="[1329,1424,1872,1898]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="49" pageNumber="50" unit="mm" value="7.7">7.7 mm</quantity>
), 
<collectionCode id="C49E2BBE4B75FFB9E7B8F8D9FECCF811" box="[151,263,1908,1934]" pageId="49" pageNumber="50">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B75FFB9E625F8D9FEE4F811" box="[266,303,1908,1934]" country="Austria" name="Zoological Collection, University of Vienna" pageId="49" pageNumber="50" type="University or college">ZC</collectionCode>
. 2019.06.58, 
<collectingRegion id="604B7D994B75FFB9E6E9F8D9FDE5F811" box="[454,558,1908,1934]" country="Japan" name="Okinawa" pageId="49" pageNumber="50">Okinawa</collectingRegion>
, Sajiki, intertidal, leg. 
<collectorName id="0F7AD6AD4B75FFB9E405F8D9FC09F811" box="[810,962,1908,1934]" pageId="49" pageNumber="50">A. Kikukawa</collectorName>
, 
<date id="D63195BB4B75FFB9E4E2F8D9FB87F811" box="[973,1100,1908,1934]" pageId="49" pageNumber="50" value="1994-08-08">
<collectingDate id="C6756C534B75FFB9E4E2F8D9FB87F811" box="[973,1100,1908,1934]" pageId="49" pageNumber="50" value="1994-08-08">08.08.1994</collectingDate>
</date>
[SMP-1332]
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B75FFB9E7E8F835FDDEF865" blockId="49.[151,1437,1620,2043]" pageId="49" pageNumber="50">
<materialsCitation id="12E7B9264B75FFB9E7E8F835FDDBF865" ID-GBIF-Occurrence="4104078469" collectingDate="2002-06-22" collectorName="H. J. Yang" country="South Korea" county="Garorim Bay" latitude="36.88493" location="Seosan-si" longLatPrecision="1" longitude="126.38985" municipality="Yecheon-dong" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Chungnam">
<emphasis id="90FB6F694B75FFB9E7E8F835FE93F82D" box="[199,344,1944,1970]" italics="true" pageId="49" pageNumber="50">
<collectingCountry id="DA98F3EB4B75FFB9E7E8F835FE93F82D" box="[199,344,1944,1970]" name="South Korea" pageId="49" pageNumber="50">South Korea</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B75FFB9E643F835FE77F82D" box="[364,444,1944,1970]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(
<quantity id="65771E9E4B75FFB9E6FEF835FDF9F82D" box="[465,562,1944,1970]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="49" pageNumber="50" unit="mm" value="8.7">8.7 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B75FFB9E43BF835FCB8F82D" box="[788,883,1944,1970]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="49" pageNumber="50" unit="mm" value="8.2">8.2 mm</quantity>
), MNHN-IU-2018-5587 (ex-SNU-220610-012), 
<collectingCounty id="4B51CBF74B75FFB9E7B8F811FEE1F848" box="[151,298,1980,2007]" pageId="49" pageNumber="50">Garorim Bay</collectingCounty>
, 
<collectingMunicipality id="425429014B75FFB9E61BF811FE12F849" box="[308,473,1980,2006]" pageId="49" pageNumber="50">Yecheon-dong</collectingMunicipality>
, 
<location id="A750E5A04B75FFB9E6CBF811FD99F849" LSID="urn:lsid:plazi:treatment:2A26026D4B75FFB0E7B8FB63FD50FE4E:A750E5A04B75FFB9E6CBF811FD99F849" box="[484,594,1980,2006]" country="South Korea" county="Garorim Bay" latitude="36.88493" longLatPrecision="1" longitude="126.38985" municipality="Yecheon-dong" name="Seosan-si" pageId="49" pageNumber="50" stateProvince="Chungnam">Seosan-si</location>
, 
<collectingRegion id="604B7D994B75FFB9E573F811FC9BF849" box="[604,848,1980,2006]" country="South Korea" name="Chungnam" pageId="49" pageNumber="50">Chungcheongnam-do</collectingRegion>
, 
<geoCoordinate id="C7BBD5BC4B75FFB9E474F811FC32F848" box="[859,1017,1980,2007]" degrees="36" direction="north" minutes="53" orientation="latitude" pageId="49" pageNumber="50" precision="1" seconds="5.74" value="36.88493">36°53’5.74”N</geoCoordinate>
– 
<geoCoordinate id="C7BBD5BC4B75FFB9E326F811FB0AF848" box="[1033,1217,1980,2007]" degrees="126" direction="east" minutes="23" orientation="longitude" pageId="49" pageNumber="50" precision="1" seconds="23.46" value="126.38985">126°23’23.46”E</geoCoordinate>
, collector unknown (probably 
<collectorName id="0F7AD6AD4B75FFB9E624F84CFEB7F865" box="[267,380,2016,2043]" pageId="49" pageNumber="50">H.J. Yang</collectorName>
), 
<date id="D63195BB4B75FFB9E6A0F84DFDDBF865" box="[399,528,2016,2042]" pageId="49" pageNumber="50" value="2002-06-22">
<collectingDate id="C6756C534B75FFB9E6A0F84DFDDBF865" box="[399,528,2016,2042]" pageId="49" pageNumber="50" value="2002-06-22">22.06.2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B76FFBAE7E8FF3AFED7FF66" blockId="50.[151,1437,150,645]" pageId="50" pageNumber="51">
<materialsCitation id="12E7B9264B76FFBAE7E8FF3AFDC5FF2E" ID-GBIF-Occurrence="4104078362" box="[199,526,151,177]" country="Taiwan" location="Taiwan" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1">
<emphasis id="90FB6F694B76FFBAE7E8FF3AFED3FF2F" box="[199,280,151,176]" italics="true" pageId="50" pageNumber="51">
<collectingCountry id="DA98F3EB4B76FFBAE7E8FF3AFED3FF2F" box="[199,280,151,176]" name="Taiwan" pageId="50" pageNumber="51">Taiwan</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B76FFBAE607FF3AFEB8FF2E" box="[296,371,151,177]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B76FFBAE68FFF3AFDCCFF2E" box="[416,519,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="50" pageNumber="51" unit="mm" value="11.8">11.8 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="12E7B9264B76FFBAE535FF3AFE73FF4A" ID-GBIF-Occurrence="4104078319" collectionCode="OUMNH, ZC, T" collectorName="Chang Hua &amp; Y. Chan" country="Taiwan" pageId="50" pageNumber="51" specimenCount="2" specimenCount-female="2">
<specimenCount id="B48978F24B76FFBAE535FF3AFD4DFF2E" box="[538,646,151,177]" pageId="50" pageNumber="51" type="female">2 females</specimenCount>
(cl 5.5, 
<quantity id="65771E9E4B76FFBAE5CBFF3BFCF5FF2E" box="[740,830,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="50" pageNumber="51" unit="mm" value="8.5">8.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B76FFBAE47EFF3AFC0AFF2E" box="[849,961,151,177]" pageId="50" pageNumber="51">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B76FFBAE4EBFF3AFC22FF2E" box="[964,1001,151,177]" country="Austria" name="Zoological Collection, University of Vienna" pageId="50" pageNumber="51" type="University or college">ZC</collectionCode>
. 2019.06.59, 
<collectorName id="0F7AD6AD4B76FFBAE3AFFF3AFAC9FF2E" box="[1152,1282,151,177]" pageId="50" pageNumber="51">Chang Hua</collectorName>
, mudflat, leg. 
<collectionCode id="C49E2BBE4B76FFBAE7B8FF16FF62FF4A" box="[151,169,187,213]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="51">T</collectionCode>
. 
<collectorName id="0F7AD6AD4B76FFBAE783FF16FEC3FF4A" box="[172,264,187,213]" pageId="50" pageNumber="51">Y. Chan</collectorName>
<emphasis id="90FB6F694B76FFBAE63EFF16FE88FF4A" box="[273,323,187,213]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
., 12.1996
</materialsCitation>
; 
<materialsCitation id="12E7B9264B76FFBAE6EBFF16FED3FF66" ID-GBIF-Occurrence="4104078364" collectingDate="1997-11-28" collectionCode="ZRC" collectorName="Y. Cai &amp; C. W. Lin" country="Taiwan" pageId="50" pageNumber="51" specimenCode="ZRC 2014.0676" specimenCount="1">
<specimenCount id="B48978F24B76FFBAE6EBFF16FDCFFF4A" box="[452,516,187,213]" pageId="50" pageNumber="51" type="generic">1 juv.</specimenCount>
male (cl 
<quantity id="65771E9E4B76FFBAE55EFF16FD07FF4A" box="[625,716,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="50" pageNumber="51" unit="mm" value="4.7">4.7 mm</quantity>
), 
<specimenCode id="F2291B004B76FFBAE5CFFF16FC52FF4A" box="[736,921,187,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="50" pageNumber="51">ZRC 2014.0676</specimenCode>
, 
<collectorName id="0F7AD6AD4B76FFBAE48AFF16FBCFFF4A" box="[933,1028,187,213]" pageId="50" pageNumber="51">Shinzhu</collectorName>
, mangrove, leg. 
<collectorName id="0F7AD6AD4B76FFBAE3EFFF16FACCFF4A" box="[1216,1287,187,213]" pageId="50" pageNumber="51">Y. Cai</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B76FFBAE202FF16FA53FF4A" box="[1325,1432,187,213]" pageId="50" pageNumber="51">C.W. Lin</collectorName>
, 
<date id="D63195BB4B76FFBAE7B8FF72FED3FF66" box="[151,280,223,249]" pageId="50" pageNumber="51" value="1997-11-28">
<collectingDate id="C6756C534B76FFBAE7B8FF72FED3FF66" box="[151,280,223,249]" pageId="50" pageNumber="51" value="1997-11-28">28.11.1997</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B76FFBEE7E8FEAFFA85FDFE" lastPageId="54" lastPageNumber="55" pageId="50" pageNumber="51" type="description">
<paragraph id="A230B37B4B76FFBAE7E8FEAFFC0AFE32" blockId="50.[151,1437,150,645]" pageId="50" pageNumber="51">
<emphasis id="90FB6F694B76FFBAE7E8FEAFFE9AFE83" bold="true" box="[199,337,258,284]" pageId="50" pageNumber="51">Description</emphasis>
. See 
<figureCitation id="3AB4AFFE4B76FFBAE6BDFEAEFDD0FE82" box="[402,539,258,285]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStartId-0="50.[151,250,1940,1964]" captionStartId-1="51.[151,250,1813,1837]" captionStartId-2="52.[151,250,1818,1842]" captionTargetBox-0="[151,1435,669,1917]" captionTargetBox-1="[167,1425,193,1775]" captionTargetBox-2="[162,1424,195,1781]" captionTargetId-0="figure-226@50.[151,1435,669,1917]" captionTargetId-1="figure-15@51.[151,1436,181,1789]" captionTargetId-2="figure-21@52.[151,1436,181,1794]" captionTargetPageId-0="50" captionTargetPageId-1="51" captionTargetPageId-2="52" captionText-0="FIGURE 25. Alpheus nomurai sp. nov.: holotype, male (cl 8.8 mm) from Tanabe, Wakayama, Japan (MNHN-IU-2018-5590); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) cheliped, lateral view; D—same, mesial view; E— minor (right) cheliped, mesial view; F—same, dorsolateral view; G—same, lateral view. Refer to A for scale." captionText-1="FIGURE 26. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D— antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—fifth pereiopod, lateral view; J—same, propodus and dactylus, mesial view; K—second pleopod, appendices masculina and interna, anterior view; L—left uropod, dorsal view." captionText-2="FIGURE 27. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588) [A–G]; paratype, ovigerous female (cl 8.8 mm) from the same locality (MNHN-IU-2018-5744) [H]; paratype, ovigerous female (cl 11.9 mm) from Shimanto River, Japan (MNHN-IU-2018-5589) [I]; A—major (right) cheliped, chela, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor cheliped, lateral view; same, carpus and chela, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7912372" figureDoi-1="http://doi.org/10.5281/zenodo.7912376" figureDoi-2="http://doi.org/10.5281/zenodo.7912380" httpUri-0="https://zenodo.org/record/7912372/files/figure.png" httpUri-1="https://zenodo.org/record/7912376/files/figure.png" httpUri-2="https://zenodo.org/record/7912380/files/figure.png" pageId="50" pageNumber="51">Figs. 25–27</figureCitation>
. Medium-sized species of 
<taxonomicName id="658FC8F84B76FFBAE466FEAEFC68FE82" authorityName="Fabricius" authorityYear="1798" box="[841,931,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B76FFBAE466FEAEFC68FE82" box="[841,931,259,285]" italics="true" pageId="50" pageNumber="51">Alpheus</emphasis>
</taxonomicName>
(maximal cl 
<quantity id="65771E9E4B76FFBAE314FEAEFB68FE82" box="[1083,1187,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="50" pageNumber="51" unit="mm" value="12.1">12.1 mm</quantity>
, tl ~ 
<quantity id="65771E9E4B76FFBAE3FAFEAEFAF4FE82" box="[1237,1343,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="50" pageNumber="51" unit="mm" value="36.3">36.3 mm</quantity>
; present study; 
<bibRefCitation id="C61ECE8A4B76FFBAE7C8FE8AFEA1FEDE" author="Miya, Y." box="[231,362,294,321]" pageId="50" pageNumber="51" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
). Carapace glabrous, finely pitted, without pubescence, with slight grooves. Rostrum short, triangular, distally acute, typically reaching 0.3 length of first article of antennular peduncle, sometimes its mid-length; rostral carina present as low elevation, fading posterior to eye level; rostro-orbital furrows poorly demarcated or indistinct. Pterygostomial angle broadly rounded; cardiac notch deep.
</paragraph>
<paragraph id="A230B37B4B76FFBAE7E8FE1AFC4FFD1A" blockId="50.[151,1437,150,645]" pageId="50" pageNumber="51">Pleon smooth, scarcely pitted, without pubescence. Telson moderately broad, ovate-rectangular, gently tapering towards posterior margin, about 1.7–1.8 times as long as maximal width near proximal margin; lateral margins with broad concavity on proximal half; dorsal surface finely pitted, without trace of median depression, with two pairs of short spiniform setae inserted in deep pits at considerable distance from lateral margins, first pair near telson mid-length, second pair between 0.7 and 0.8 of telson length; posterior margin broadly rounded, with two pairs of spiniform setae at posterolateral angles, lateral shorter than mesial.</paragraph>
<caption id="F6F0E3F34B76FFBAE7B8F839FB21F86B" ID-DOI="http://doi.org/10.5281/zenodo.7912372" ID-Zenodo-Dep="7912372" httpUri="https://zenodo.org/record/7912372/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,250,1940,1964]" targetBox="[151,1435,669,1917]" targetPageId="50" targetType="figure">
<paragraph id="A230B37B4B76FFBAE7B8F839FB21F86B" blockId="50.[151,1436,1940,2037]" pageId="50" pageNumber="51">
<emphasis id="90FB6F694B76FFBAE7B8F839FED2F833" bold="true" box="[151,281,1940,1964]" pageId="50" pageNumber="51">FIGURE 25</emphasis>
. 
<taxonomicName id="658FC8F84B76FFBAE609F839FE1FF833" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[294,468,1940,1964]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B76FFBAE609F839FE1FF833" box="[294,468,1940,1964]" italics="true" pageId="50" pageNumber="51">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B76FFBAE6F4F838FDE0F832" bold="true" box="[475,555,1941,1965]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="8BC8D2124B76FFBAE6F4F838FDE0F832" box="[475,555,1941,1965]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: holotype, male (cl 8.8 mm) from Tanabe, Wakayama, Japan (MNHN-IU-2018-5590); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) cheliped, lateral view; D—same, mesial view; E— minor (right) cheliped, mesial view; F—same, dorsolateral view; G—same, lateral view. Refer to A for scale.
</paragraph>
</caption>
<caption id="F6F0E3F34B77FFBBE7B8F8B8FB4FF822" ID-DOI="http://doi.org/10.5281/zenodo.7912376" ID-Zenodo-Dep="7912376" httpUri="https://zenodo.org/record/7912376/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1813,1837]" targetBox="[167,1425,193,1775]" targetPageId="51" targetType="figure">
<paragraph id="A230B37B4B77FFBBE7B8F8B8FB4FF822" blockId="51.[151,1437,1813,1981]" pageId="51" pageNumber="52">
<emphasis id="90FB6F694B77FFBBE7B8F8B8FED1F8B2" bold="true" box="[151,282,1813,1837]" pageId="51" pageNumber="52">FIGURE 26</emphasis>
. 
<taxonomicName id="658FC8F84B77FFBBE608F8B8FE1DF8B2" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[295,470,1813,1837]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B77FFBBE608F8B8FE1DF8B2" box="[295,470,1813,1837]" italics="true" pageId="51" pageNumber="52">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B77FFBBE6F2F8B8FDE5F8B2" bold="true" box="[477,558,1813,1837]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="8BC8D2124B77FFBBE6F2F8B8FDE5F8B2" box="[477,558,1813,1837]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D— antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—fifth pereiopod, lateral view; J—same, propodus and dactylus, mesial view; K—second pleopod, appendices masculina and interna, anterior view; L—left uropod, dorsal view.
</paragraph>
</caption>
<caption id="F6F0E3F34B70FFBCE7B8F8B7FCF8F878" ID-DOI="http://doi.org/10.5281/zenodo.7912380" ID-Zenodo-Dep="7912380" httpUri="https://zenodo.org/record/7912380/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1818,1842]" targetBox="[162,1424,195,1781]" targetPageId="52" targetType="figure">
<paragraph id="A230B37B4B70FFBCE7B8F8B7FCF8F878" blockId="52.[151,1436,1818,2023]" pageId="52" pageNumber="53">
<emphasis id="90FB6F694B70FFBCE7B8F8B7FED6F8AD" bold="true" box="[151,285,1818,1842]" pageId="52" pageNumber="53">FIGURE 27</emphasis>
. 
<taxonomicName id="658FC8F84B70FFBCE602F8B6FE14F8AD" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[301,479,1819,1843]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B70FFBCE602F8B6FE14F8AD" box="[301,479,1819,1843]" italics="true" pageId="52" pageNumber="53">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B70FFBCE6C6F8B6FDF6F8AC" bold="true" box="[489,573,1819,1843]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="8BC8D2124B70FFBCE6C6F8B6FDF6F8AC" box="[489,573,1819,1843]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588) [A–G]; paratype, ovigerous female (cl 8.8 mm) from the same locality (MNHN-IU-2018-5744) [H]; paratype, ovigerous female (cl 11.9 mm) from Shimanto River, Japan (MNHN-IU-2018-5589) [I]; A—major (right) cheliped, chela, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor cheliped, lateral view; same, carpus and chela, lateral view.
</paragraph>
</caption>
<paragraph id="A230B37B4B71FFBDE7E8FF3AFD39FEFC" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">Antennular peduncle with stylocerite broad, markedly convex laterally, with acute tip, latter reaching, but not overreaching distal margin of first article; ventromesial carina with anteriorly directed, distally subacute tooth; second article about 2.3–2.4 times as long as wide. Antennal peduncle with basicerite armed with small, sharp tooth; carpocerite reaching far beyond scaphocerite; scaphocerite with lateral margin nearly straight to very shallowly concave; blade moderately broad, separated from broad distolateral tooth by deep cleft; anterior margin of blade broadly rounded, exceeded by stout distolateral tooth.</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8FEC0FBBDFE6D" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">Third maxilliped with antepenultimate article not particularly broadened; penultimate article elongate, about four times as long as wide, with long setae on ventral margin; ultimate article with dense transverse rows of short serrulate setae on mesial surface and longer setae on dorsal and lateral surfaces, especially near or on apex; coxal lateral plate acutely protruding dorsally; exopod reaching end of antepenultimate article.</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8FE51FDA1FC5C" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">
Major cheliped of 
<taxonomicName id="658FC8F84B71FFBDE6B6FE51FDEDFD89" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[409,550,508,534]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="90FB6F694B71FFBDE6B6FE51FDEDFD89" box="[409,550,508,534]" italics="true" pageId="53" pageNumber="54">A. edwardsii</emphasis>
</taxonomicName>
- 
<typeStatus id="7D340DD94B71FFBDE501FE50FDA8FD88" box="[558,611,509,535]" pageId="53" pageNumber="54">type</typeStatus>
. Merus stout, about twice as long as wide; ventromesial margin unarmed distally. Carpus very short, cup-shaped. Chela large, massive, with fingers about 0.7 length of palm. Mesial face of palm largely smooth, except for small field of minute granules dorsal to ventral shoulder, and with weak mesial transverse ridge in proximal half, extending from proximal edge of subtriangular mesial longitudinal groove to ventral surface of palm. Lateral face of palm smooth; lateral longitudinal groove deep, subrectangular; proximal half without distinct transverse ridge. Dorsal shoulder smooth, gently sloping into broad transverse groove with angle of about 45°. Ventral shoulder pronounced, slightly protruding in lateral view, broadly rounded, smooth. Pollex with mesial face finely granulated over most of its surface, with short subdistal mesial ridge near margin, without proximal protuberance; area ventral to mesial subdistal ridge not grooved; distomesial angle distinctly superior to 90°, blunt; lateral face smooth; distolateral angle superior to 90°. Dactylus with weak dorsal ridge, latter slightly twisted mesially, not protruding proximally; mesial and lateral surfaces smooth; dactylar plunger very stout, large, with anterior margin well delineated, clearly separated from ventral margin of dactylus; distal surface of plunger somewhat truncate. Adhesive discs small.
</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8FC61FE73FAD3" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">
<collectingRegion id="604B7D994B71FFBDE7E8FC61FECAFC79" box="[199,257,972,998]" country="Maldives" name="Maale" pageId="53" pageNumber="54">Male</collectingRegion>
minor cheliped with chela strongly balaeniceps. Merus slightly slenderer than that of major cheliped, about 2.5 times as long as wide; ventral margin with distomesial angle unarmed; distomesial margin slightly rugose. Chela not particularly swollen, slender in lateral view, with fingers about as long as palm; all surfaces smooth, without granulation. Palm subcylindrical, compressed, feebly swollen; both mesial and lateral longitudinal grooves well marked, deep; dorsal shoulder distinct, rounded, sloping almost perpendicularly into dorsal transverse groove; ventral transverse groove present, relatively deep, marked by distinct ventral shoulder on lateral surface. Pollex with mesial surface bearing row of balaeniceps setae on low crest extending from base to about mid-length of pollex; lateral surface with similar row of balaeniceps setae as on mesial surface. Dactylus with conspicuous lateral expansion (balaeniceps expansion) in proximal half, about 1.7 as long as maximal width; lateral and mesial dactylar ridges each with dense rows of balaeniceps setae. Cutting edges of both fingers shallowly excavated on each side of sharp, blade-like ridge.
</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8FAF8FADFF9BD" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">Female minor cheliped simple, not balaeniceps. Chela slender, not particularly swollen, with fingers subequal to, or slightly longer than palm. Palm with surface smooth, not granulated; mesial longitudinal groove feebly developed; lateral longitudinal groove distinct, but shallow; dorsal transverse groove usually distinct, but dorsal shoulder poorly demarcated; ventral transverse groove present in form of deep sinus bordered by feebly developed ventral shoulder, latter sometimes not distinct (especially in younger females or regenerated chelipeds). Fingers simple, with sharp cutting edges, slightly gaping when closed; fingertips strongly curved and crossing.</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8F981FC94F8FB" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">Second pereiopod with ratio of carpal subarticles approximately equal to 4: 2.3: 1: 1: 1.7. Third pereiopod moderately robust; ischium armed with short, stout spiniform seta on ventrolateral surface; merus about 5.2 times as long as maximal width, unarmed; propodus with three or four short spiniform setae on ventral margin and one spiniform seta on distoventral margin adjacent to dactylus; dactylus about half-length of propodus, distinctly spatulate, expanded, wider than propodus in dorsal view, with two rows of setae along ridges. Fourth pereiopod generally similar to third pereiopod, shorter and slenderer; dactylus comparatively shorter, about 0.4 length of propodus. Fifth pereiopod much slenderer than third and fourth pereiopods; ischium unarmed; propodus apparently without spiniform setae, with well-developed setal brush; dactylus subspatulate, much less expanded compared to that of third or fourth pereiopod, about half-length of propodus.</paragraph>
<paragraph id="A230B37B4B71FFBDE7E8F8C0FDD1F86B" blockId="53.[151,1437,151,2036]" pageId="53" pageNumber="54">
<collectingRegion id="604B7D994B71FFBDE7E8F8C0FECAF818" box="[199,257,1901,1927]" country="Maldives" name="Maale" pageId="53" pageNumber="54">Male</collectingRegion>
second pleopod with appendix masculina 0.8 length of appendix interna. Uropod with each protopodal lobe ending in subacute tooth distally; exopod and endopod broad, ovate; diaeresis of exopod complete, almost straight, with broadly subtriangular lobe near small distolateral spiniform seta and adjacent distolateral tooth; lateral margin of exopod broadly convex.
</paragraph>
<paragraph id="A230B37B4B72FFBEE7E8FF3AFBE6FF2E" blockId="54.[151,1437,150,2013]" box="[199,1069,150,177]" pageId="54" pageNumber="55">
Eggs numerous (&gt; 
<quantity id="65771E9E4B72FFBEE6BBFF3AFE17FF2E" box="[404,476,151,177]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="54" pageNumber="55" unit="in" value="100.0">100 in</quantity>
larger females), small; egg diameter about 
<quantity id="65771E9E4B72FFBEE4EDFF3BFBE2FF2E" box="[962,1065,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="54" pageNumber="55" unit="mm" value="0.65">0.65 mm</quantity>
.
</paragraph>
<paragraph id="A230B37B4B72FFBEE7E8FF17FA85FDFE" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FF17FEB6FF4A" bold="true" box="[199,381,186,213]" pageId="54" pageNumber="55">Colour pattern</emphasis>
. Carapace greyish, semi-opaque, with two complete transverse bands of dark brown to dark grey-blue colour, one broad complete band near posterior margin and one narrower, laterally broken band at about carapace mid-length; two smaller, incomplete (dorsally not closing) and more diffuse bands present in anterior half of carapace, one along its anterolateral margin and one lateral to cardiac area; pleon with six moderately broad, dark blue-grey, blue-green or red-brown transverse bands, one across each pleonite and near its posterior margin; broad white spaces between dark bands with small but conspicuous neon-yellow spots, most of them fringing dark bands on carapace and pleon; antennular peduncles and scaphocerite patchily marbled with brown and yellow; antennular and antennal flagella pale yellow or pale greenish; mesial face of major chela olive-green, with paler greenish or orange-yellowish areas, especially on dorsal and ventral transverse grooves; dactylus dark olive-green; distal portion of both fingers pale orange; minor chela olive-green mottled with brown, fingers darker; second to fifth pereiopods pale yellowish; telson and uropods with patchy brown mottling and yellow spots, darker distally; posterior half of uropodal exopod dark blue (
<figureCitation id="3AB4AFFE4B72FFBEE6FCFDEAFDECFDFE" box="[467,551,583,609]" captionStart="FIGURE 28" captionStartId="55.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1790]" captionTargetId="figure-15@55.[151,1435,181,1790]" captionTargetPageId="55" captionText="FIGURE 28. Alpheus nomurai sp. nov.: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7912382" httpUri="https://zenodo.org/record/7912382/files/figure.png" pageId="54" pageNumber="55">Fig. 28</figureCitation>
; see also 
<bibRefCitation id="C61ECE8A4B72FFBEE5BEFDEAFCBEFDFE" author="Jeng, M. S. &amp; Chang, K. H." box="[657,885,582,609]" pageId="54" pageNumber="55" pagination="241 - 256" refId="ref71966" refString="Jeng, M. S. &amp; Chang, K. H. (1985) Snapping shrimps (Crustacea: Decapoda: Alpheidae) of Taiwan. Bulletin of Institute of Zoology, Academica Sinica, 24, 241 - 256." type="journal article" year="1985">Jeng &amp; Chang 1985</bibRefCitation>
: fig. 23; 
<bibRefCitation id="C61ECE8A4B72FFBEE4F9FDEAFB48FDFE" author="Yoshigou, H." box="[982,1155,583,609]" pageId="54" pageNumber="55" pagination="221 - 273" refId="ref75338" refString="Yoshigou, H. (2009) A collection of alpheid shrimps (Crustacea; Decapoda; Alpheidae) from the mouth of rivers and anchialine caves of Japan. Miscellaneous Reports of the Hiwa Museum for Natural History, 50, 221 - 273. [in Japanese with English abstract]" type="journal article" year="2009">Yoshigou 2009</bibRefCitation>
: pl. III, figs. 1–4).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFBEE7E8FDC6FAC4FD1A" box="[199,1295,618,645]" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="A230B37B4B72FFBEE7E8FDC6FAC4FD1A" blockId="54.[151,1437,150,2013]" box="[199,1295,618,645]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FDC6FEABFD1B" bold="true" box="[199,352,618,645]" pageId="54" pageNumber="55">Type locality</emphasis>
. 
<materialsCitation id="12E7B9264B72FFBEE642FDC6FAC0FD1A" ID-GBIF-Occurrence="4104078458" box="[365,1291,619,645]" country="Japan" location="Tanabe" municipality="Honshu" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
<location id="A750E5A04B72FFBEE642FDC6FE0BFD1A" LSID="urn:lsid:plazi:treatment:2A26026D4B75FFB0E7B8FB63FD50FE4E:A750E5A04B72FFBEE642FDC6FE0BFD1A" box="[365,448,619,645]" country="Japan" municipality="Honshu" name="Tanabe" pageId="54" pageNumber="55" stateProvince="Wakayama">Tanabe</location>
, 
<collectingRegion id="604B7D994B72FFBEE6E4FDC6FD0FFD1A" box="[459,708,619,645]" country="Japan" name="Wakayama" pageId="54" pageNumber="55">Wakayama Prefecture</collectingRegion>
, 
<collectingMunicipality id="425429014B72FFBEE5E0FDC6FCE2FD1A" box="[719,809,619,645]" pageId="54" pageNumber="55">Honshu</collectingMunicipality>
, 
<collectingCountry id="DA98F3EB4B72FFBEE41BFDC6FCBFFD1A" box="[820,884,619,645]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
(collection locality of the 
<typeStatus id="7D340DD94B72FFBEE3B3FDC6FAC8FD1A" box="[1180,1283,619,645]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFBEE7E8FD23FAB0FD52" pageId="54" pageNumber="55" type="distribution">
<paragraph id="A230B37B4B72FFBEE7E8FD23FAB0FD52" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FD23FE92FD37" bold="true" box="[199,345,654,680]" pageId="54" pageNumber="55">Distribution</emphasis>
. Temperate and subtropical north-western Pacific (
<figureCitation id="3AB4AFFE4B72FFBEE4B1FD22FBCFFD36" box="[926,1028,654,681]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="54" pageNumber="55">Fig. 51F</figureCitation>
): 
<collectingCountry id="DA98F3EB4B72FFBEE336FD22FB92FD36" box="[1049,1113,655,681]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
(Honshu, Kyushu, Shikoku, 
<collectingRegion id="604B7D994B72FFBEE7B8FD1EFECAFD52" box="[151,257,691,717]" country="Japan" name="Okinawa" pageId="54" pageNumber="55">Okinawa</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B72FFBEE63BFD1EFE6BFD52" box="[276,416,691,717]" name="South Korea" pageId="54" pageNumber="55">South Korea</collectingCountry>
(Garorim Bay, Byeonsan = Pyonsan) and 
<collectingCountry id="DA98F3EB4B72FFBEE455FD1EFC07FD52" box="[890,972,691,717]" name="Taiwan" pageId="54" pageNumber="55">Taiwan</collectingCountry>
(Chang Hua, Shinzhu, Tamsui River).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFBEE7E8FD7AFE25FCA6" pageId="54" pageNumber="55" type="etymology">
<paragraph id="A230B37B4B72FFBEE7E8FD7AFE25FCA6" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FD7AFE83FD6E" bold="true" box="[199,328,727,753]" pageId="54" pageNumber="55">Etymology</emphasis>
. This species is named after the author’s dear colleague, Keiichi Nomura (Sabiura Marine Park Research Station, 
<collectingCountry id="DA98F3EB4B72FFBEE648FD56FE60FC8A" box="[359,427,763,789]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
), who contributed to alpheid taxonomy early in his career and made important material available for the present study.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFBEE7E8FCEFFCF3FCC2" box="[199,824,834,861]" pageId="54" pageNumber="55" type="vernacular_names">
<paragraph id="A230B37B4B72FFBEE7E8FCEFFCF3FCC2" blockId="54.[151,1437,150,2013]" box="[199,824,834,861]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FCEFFE38FCC2" bold="true" box="[199,499,834,861]" pageId="54" pageNumber="55">Common name proposed</emphasis>
. Nomura’s snapping shrimp.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFBEE7E8FCCAFCFFFA02" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="A230B37B4B72FFBEE7E8FCCAFCFFFA02" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FCCAFE77FC1F" bold="true" box="[199,444,870,897]" pageId="54" pageNumber="55">Ecology and biology</emphasis>
. 
<taxonomicName id="658FC8F84B72FFBEE6E4FCCAFD46FC1F" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[459,653,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE6E4FCCAFD46FC1F" box="[459,653,871,897]" italics="true" pageId="54" pageNumber="55">Alpheus nomurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BC8D2124B72FFBEE5B9FCCAFD38FC1E" box="[662,755,871,897]" pageId="54" pageNumber="55" rank="species">
<emphasis id="90FB6F694B72FFBEE5B9FCCAFD27FC1E" bold="true" box="[662,748,871,897]" pageId="54" pageNumber="55">sp. nov</emphasis>
.
</taxonomicNameLabel>
is a marine / estuarine snapping shrimp largely confined to intertidal mudflats, especially those situated inside or close to estuaries. The shrimps usually live in pairs (rarely as singles) in complex tridimensional burrows made in soft mud, with side branches extending from the burrow entrance in a radius of about 
<quantity id="65771E9E4B72FFBEE6FBFC7FFDD3FC72" box="[468,536,978,1005]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="54" pageNumber="55" unit="cm" value="25.0">25 cm</quantity>
and penetrating about 
<quantity id="65771E9E4B72FFBEE43BFC7EFC4AFC72" box="[788,897,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="54" pageNumber="55" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 cm</quantity>
deep into the mud (
<bibRefCitation id="C61ECE8A4B72FFBEE375FC7EFB13FC72" author="Miya, Y." box="[1114,1240,978,1005]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE3D1FC7EFED2FB8E" authorityName="Yaldwyn" authorityYear="1971" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">
<emphasis id="90FB6F694B72FFBEE3D1FC7EFED2FB8E" italics="true" pageId="54" pageNumber="55">A. euphrosyne richardsoni</emphasis>
</taxonomicName>
, based on 
<bibRefCitation id="C61ECE8A4B72FFBEE6B8FC5AFDDFFB8E" author="Dotu, Y." box="[407,532,1015,1041]" pageId="54" pageNumber="55" pagination="133 - 139" refId="ref71162" refString="Dotu, Y. (1961) The bionomics and life history of the gobioid fish, Apocryptodon bleekeri (Day). Bulletin of the Faculty of Fisheries, Nagasaki University, 10, 133 - 139, pl. 28. [in Japanese]" type="journal article" year="1961">Dôtu 1961</bibRefCitation>
: fig. 1). The burrows of 
<taxonomicName id="658FC8F84B72FFBEE41FFC5AFC7BFB8F" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[816,944,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE41FFC5AFC7BFB8F" box="[816,944,1015,1041]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE496FC5AFBDEFB8E" bold="true" box="[953,1045,1015,1041]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE496FC5AFBDEFB8E" box="[953,1045,1015,1041]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are typically composed of a main tunnel with one or two or three openings and multiple branches, either interconnected or ending blindly (see 
<bibRefCitation id="C61ECE8A4B72FFBEE210FBB6FEDAFBC6" author="Kirihara, S. &amp; Itani, G. &amp; Nunobe, J. I. &amp; Nomoto, A. &amp; Aldea, K. Q. &amp; Murakami, R. &amp; Sakata, H. &amp; Henmi, Y." pageId="54" pageNumber="55" pagination="99 - 102" refId="ref72559" refString="Kirihara, S., Itani, G., Nunobe, J. I., Nomoto, A., Aldea, K. Q., Murakami, R., Sakata, H. &amp; Henmi, Y. (2021) Burrow morphology of an alpheid shrimp at muddy tidal flats in western Japan. Kuroshio Science, 14, 99 - 102." type="journal article" year="2021">
Kirihara 
<emphasis id="90FB6F694B72FFBEE7B8FB92FF03FBC6" box="[151,200,1087,1113]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2021
</bibRefCitation>
: figs. 2, 3, as 
<taxonomicName id="658FC8F84B72FFBEE686FB92FDE7FBC6" box="[425,556,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B72FFBEE686FB92FDC8FBC6" box="[425,515,1087,1113]" italics="true" pageId="54" pageNumber="55">Alpheus</emphasis>
sp.
</taxonomicName>
). They often serve as shelter for small gobies, such as 
<taxonomicName id="658FC8F84B72FFBEE3A5FB92FE9CFBE2" authority="Tomiyama, 1934" authorityName="Tomiyama" authorityYear="1934" class="Actinopterygii" family="Gobiidae" genus="Apocryptodon" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="90FB6F694B72FFBEE3A5FB92FA57FBC7" box="[1162,1436,1087,1113]" italics="true" pageId="54" pageNumber="55">Apocryptodon punctatus</emphasis>
<bibRefCitation id="C61ECE8A4B72FFBEE7B8FBCEFE9CFBE2" author="Tomiyama, I." box="[151,343,1123,1149]" pageId="54" pageNumber="55" pagination="325 - 334" refId="ref74678" refString="Tomiyama, I. (1934) Four species of gobies of Japan. Journal of the Tokyo Imperial University, Faculty of Science, Section IV, 3, 325 - 334." type="journal article" year="1934">Tomiyama, 1934</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B72FFBEE64DFBCEFC9CFBE2" authority="(Tomiyama, 1936)" baseAuthorityName="Tomiyama" baseAuthorityYear="1936" box="[354,855,1123,1149]" class="Actinopterygii" family="Gobiidae" genus="Gymnogobius" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="cylindricus">
<emphasis id="90FB6F694B72FFBEE64DFBCEFD4BFBE2" box="[354,640,1123,1149]" italics="true" pageId="54" pageNumber="55">Gymnogobius cylindricus</emphasis>
(
<bibRefCitation id="C61ECE8A4B72FFBEE5A1FBCEFC85FBE2" author="Tomiyama, I." box="[654,846,1123,1149]" pageId="54" pageNumber="55" pagination="37 - 112" refId="ref74712" refString="Tomiyama, I. (1936) Gobiidae of Japan. Japanese Journal of Zoology, 7, 37 - 112." type="journal article" year="1936">Tomiyama, 1936</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="658FC8F84B72FFBEE4A3FBCEFB9EFBE2" box="[908,1109,1123,1149]" class="Actinopterygii" family="Gobiidae" genus="Acentrogobius" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="90FB6F694B72FFBEE4A3FBCEFBE5FBE2" box="[908,1070,1123,1149]" italics="true" pageId="54" pageNumber="55">Acentrogobius</emphasis>
sp.
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B72FFBEE34CFBCEFB29FBE2" author="Miya, Y." box="[1123,1250,1122,1149]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE3C2FBCEFF19FB3E" author="Suzuki, T. &amp; Wada, K." pageId="54" pageNumber="55" pagination="61 - 63" refId="ref74605" refString="Suzuki, T. &amp; Wada, K. (1999) Record of a gobiid fish, Apocryptodon punctatus from the Uchinoura Inlet in Wakayama Prefecture, Japan. Nankiseibutu, 41, 61 - 63." type="journal article" year="1999">Suzuki &amp; Wada 1999</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE7F0FB2AFE46FB3E" author="Yoshigou, H." box="[223,397,1159,1185]" pageId="54" pageNumber="55" pagination="221 - 273" refId="ref75338" refString="Yoshigou, H. (2009) A collection of alpheid shrimps (Crustacea; Decapoda; Alpheidae) from the mouth of rivers and anchialine caves of Japan. Miscellaneous Reports of the Hiwa Museum for Natural History, 50, 221 - 273. [in Japanese with English abstract]" type="journal article" year="2009">Yoshigou 2009</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE6B5FB2AFDB4FB3E" author="Koyama, A. &amp; Inui, R. &amp; Sawa, K. &amp; Onikura, N." box="[410,639,1159,1185]" pageId="54" pageNumber="55" pagination="131 - 138" refId="ref72628" refString="Koyama, A., Inui, R., Sawa, K. &amp; Onikura, N. (2017) Symbiotic partner specificity and dependency of two gobies (Apocryptodon punctatus and Acentrogobius sp. A) and four alpheid shrimps inhabiting the temperate estuary of southern Japan. Ichthyological Research, 64, 131 - 138. https: // doi. org / 10.1007 / s 10228 - 016 - 0535 - 6" type="journal article" year="2017">
Koyama 
<emphasis id="90FB6F694B72FFBEE52DFB2AFDFEFB3E" box="[514,565,1159,1185]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE5A2FB2AFCBBFB3E" author="Kirihara, S. &amp; Itani, G. &amp; Nunobe, J. I. &amp; Nomoto, A. &amp; Aldea, K. Q. &amp; Murakami, R. &amp; Sakata, H. &amp; Henmi, Y." box="[653,880,1159,1185]" pageId="54" pageNumber="55" pagination="99 - 102" refId="ref72559" refString="Kirihara, S., Itani, G., Nunobe, J. I., Nomoto, A., Aldea, K. Q., Murakami, R., Sakata, H. &amp; Henmi, Y. (2021) Burrow morphology of an alpheid shrimp at muddy tidal flats in western Japan. Kuroshio Science, 14, 99 - 102." type="journal article" year="2021">
Kirihara 
<emphasis id="90FB6F694B72FFBEE5DCFB2AFCEDFB3E" box="[755,806,1159,1185]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2021
</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE4B2FB2AFBE9FB3E" box="[925,1058,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B72FFBEE4B2FB2AFC3CFB3E" box="[925,1015,1159,1185]" italics="true" pageId="54" pageNumber="55">Alpheus</emphasis>
sp.
</taxonomicName>
, 
<taxonomicName id="658FC8F84B72FFBEE31FFB2AFA97FB3E" authorityName="Yaldwyn" authorityYear="1971" box="[1072,1372,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">
<emphasis id="90FB6F694B72FFBEE31FFB2AFA97FB3E" box="[1072,1372,1159,1185]" italics="true" pageId="54" pageNumber="55">A. euphrosyne richardsoni</emphasis>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B72FFBEE2ABFB2AFED2FB5A" authorityName="Yaldwyn" authorityYear="1971" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE2ABFB2AFED2FB5A" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
). It must be noted that the association between the snapping shrimp identified as 
<taxonomicName id="658FC8F84B72FFBEE3E7FB06FAA0FB5A" authorityName="Yaldwyn" authorityYear="1971" box="[1224,1387,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE3E7FB06FAA0FB5A" box="[1224,1387,1195,1221]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
and the goby 
<taxonomicName id="658FC8F84B72FFBEE62FFB62FE03FB77" baseAuthorityName="Jordan &amp; Seale" baseAuthorityYear="1907" box="[256,456,1231,1257]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ocyurus">
<emphasis id="90FB6F694B72FFBEE62FFB62FE03FB77" box="[256,456,1231,1257]" italics="true" pageId="54" pageNumber="55">Drombus ocyurus</emphasis>
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B72FFBEE6F8FB62FD07FB76" author="Jordan, D. S. &amp; Seale, A." box="[471,716,1231,1257]" pageId="54" pageNumber="55" pagination="1 - 48" refId="ref72130" refString="Jordan, D. S. &amp; Seale, A. (1907) Fishes of the islands of Luzon and Panay. Bulletin of the Bureau of Fisheries, 26, 1 - 48." type="journal article" year="1907">
<collectingCountry id="DA98F3EB4B72FFBEE6F8FB62FDEFFB76" box="[471,548,1231,1257]" name="Jordan" pageId="54" pageNumber="55">Jordan</collectingCountry>
&amp; Seale, 1907
</bibRefCitation>
) in 
<bibRefCitation id="C61ECE8A4B72FFBEE5D6FB62FC34FB76" author="Kunishima, T. &amp; Palla, H. P. &amp; Tachikara, K. &amp; Maeda, K." box="[761,1023,1231,1257]" pageId="54" pageNumber="55" pagination="129 - 138" refId="ref72705" refString="Kunishima, T., Palla, H. P., Tachikara, K. &amp; Maeda, K. (2022) First records of an estuarine goby Acentrogobius ocyurus (Gobiiformes: Gobiidae) from Japan and the Sulu Sea in the Philippines. Species Diversity, 27, 129 - 138. https: // doi. org / 10.12782 / specdiv. 27.129" type="journal article" year="2022">
Kunishima 
<emphasis id="90FB6F694B72FFBEE453FB62FC7FFB76" box="[892,948,1231,1257]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2022
</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE305FB62FAE6FB77" baseAuthorityName="Jordan &amp; Seale" baseAuthorityYear="1907" box="[1066,1325,1231,1257]" class="Actinopterygii" family="Gobiidae" genus="Acentrogobius" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ocyurus">
<emphasis id="90FB6F694B72FFBEE305FB62FAE6FB77" box="[1066,1325,1231,1257]" italics="true" pageId="54" pageNumber="55">Acentrogobius ocyurus</emphasis>
</taxonomicName>
) involves a species from the 
<taxonomicName id="658FC8F84B72FFBEE642FB5EFDD2FA92" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[365,537,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B72FFBEE642FB5EFDD2FA92" box="[365,537,1267,1293]" italics="true" pageId="54" pageNumber="55">A. malabaricus</emphasis>
</taxonomicName>
complex, possibly 
<taxonomicName id="658FC8F84B72FFBEE5DAFB5EFBB2FA92" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[757,1145,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="dolichodactylus">
<emphasis id="90FB6F694B72FFBEE5DAFB5EFC0DFA92" box="[757,966,1267,1293]" italics="true" pageId="54" pageNumber="55">A. dolichodactylus</emphasis>
<bibRefCitation id="C61ECE8A4B72FFBEE4E1FB5EFBB2FA92" author="Ortmann, A." box="[974,1145,1267,1293]" pageId="54" pageNumber="55" pagination="437 - 542" refId="ref73624" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beru ¨ cksichtigung der von Herrn Dr. D ˆ derlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 5, 437 - 542, pls. 36, 37." type="book chapter" year="1890">Ortmann, 1890</bibRefCitation>
</taxonomicName>
(see under the synonymy of 
<taxonomicName id="658FC8F84B72FFBEE799FABAFE9CFAAE" authorityName="Yaldwyn" authorityYear="1971" box="[182,343,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE799FABAFE9CFAAE" box="[182,343,1303,1329]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
). Whether or not 
<taxonomicName id="658FC8F84B72FFBEE531FABAFD5DFAAF" baseAuthorityName="Jordan &amp; Seale" baseAuthorityYear="1907" box="[542,662,1303,1329]" class="Actinopterygii" family="Gobiidae" genus="Acentrogobius" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ocyurus">
<emphasis id="90FB6F694B72FFBEE531FABAFD5DFAAF" box="[542,662,1303,1329]" italics="true" pageId="54" pageNumber="55">A. ocyurus</emphasis>
</taxonomicName>
also associates with 
<taxonomicName id="658FC8F84B72FFBEE4AEFABAFC34FAAF" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[897,1023,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE4AEFABAFC34FAAF" box="[897,1023,1303,1329]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE329FABAFBABFAAE" bold="true" box="[1030,1120,1303,1329]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE329FABAFBABFAAE" box="[1030,1120,1303,1329]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
remains to be shown. Some shrimps were found in burrows made under large intertidal rocks partly embedded in mud (
<bibRefCitation id="C61ECE8A4B72FFBEE3B9FA96FAB1FACA" author="Jeng, M. S. &amp; Chang, K. H." box="[1174,1402,1338,1365]" pageId="54" pageNumber="55" pagination="241 - 256" refId="ref71966" refString="Jeng, M. S. &amp; Chang, K. H. (1985) Snapping shrimps (Crustacea: Decapoda: Alpheidae) of Taiwan. Bulletin of Institute of Zoology, Academica Sinica, 24, 241 - 256." type="journal article" year="1985">Jeng &amp; Chang 1985</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE7B8FAF2FEFEFAE6" authorityName="sensu De Man" authorityYear="1897" box="[151,309,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B72FFBEE7B8FAF2FEFEFAE6" box="[151,309,1375,1401]" italics="true" pageId="54" pageNumber="55">A. euphrosyne</emphasis>
</taxonomicName>
). The large number of eggs and their small size suggest an extended larval development in 
<taxonomicName id="658FC8F84B72FFBEE20FFAF2FA57FAE7" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1312,1436,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE20FFAF2FA57FAE7" box="[1312,1436,1375,1401]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE7B8FA2EFF3AFA02" bold="true" box="[151,241,1411,1437]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE7B8FA2EFF3AFA02" box="[151,241,1411,1437]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="C61ECE8A4B72FFBEE7D0FA2EFE19FA02" author="Yang, B. P. &amp; Kim, C. H." box="[255,466,1411,1437]" pageId="54" pageNumber="55" pagination="106 - 114" refId="ref75154" refString="Yang, B. P. &amp; Kim, C. H. (1996) Zoeal stages of Alpheus euphrosyne richardsoni Yaldwyn, 1971 (Decapoda: Macrura: Alpheidae) reared in the laboratory. Korean Journal of Zoology, 39, 106 - 114." type="journal article" year="1996">Yang &amp; Kim 1996</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE6D3FA2EFCEFFA02" authorityName="Yaldwyn" authorityYear="1971" box="[508,804,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">
<emphasis id="90FB6F694B72FFBEE6D3FA2EFCEFFA02" box="[508,804,1411,1437]" italics="true" pageId="54" pageNumber="55">A. euphrosyne richardsoni</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B72FFB0E7E8FA0BFD50FE4E" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="discussion">
<paragraph id="A230B37B4B72FFB0E7E8FA0BFF7CFE82" blockId="54.[151,1437,150,2013]" lastBlockId="56.[151,1437,151,465]" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55">
<emphasis id="90FB6F694B72FFBEE7E8FA0BFE73FA5E" bold="true" box="[199,440,1446,1473]" pageId="54" pageNumber="55">Taxonomic remarks</emphasis>
. 
<taxonomicName id="658FC8F84B72FFBEE6E8FA0AFD43FA5F" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[455,648,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE6E8FA0AFD43FA5F" box="[455,648,1447,1473]" italics="true" pageId="54" pageNumber="55">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE5BEFA0AFD20FA5E" bold="true" box="[657,747,1447,1473]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE5BEFA0AFD20FA5E" box="[657,747,1447,1473]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was previously confused with two other species of 
<taxonomicName id="658FC8F84B72FFBEE214FA0AFA5EFA5E" authorityName="Fabricius" authorityYear="1798" box="[1339,1429,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B72FFBEE214FA0AFA5EFA5E" box="[1339,1429,1447,1473]" italics="true" pageId="54" pageNumber="55">Alpheus</emphasis>
</taxonomicName>
, viz. the distantly related 
<taxonomicName id="658FC8F84B72FFBEE697FA66FC51FA7A" authority="De Haan, 1844" authorityName="De Haan" authorityYear="1844" box="[440,922,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="bisincisus">
<emphasis id="90FB6F694B72FFBEE697FA66FD82FA7A" box="[440,585,1483,1509]" italics="true" pageId="54" pageNumber="55">A. bisincisus</emphasis>
<bibRefCitation id="C61ECE8A4B72FFBEE57CFA66FCC8FA7A" author="De Haan, W." box="[595,771,1483,1509]" pageId="54" pageNumber="55" pagination="1 - 55" refId="ref70613" refString="De Haan, W. (1844) [1833 - 1850] Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suspecto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Apud Auctorem, Lugduni-Batavorum, pp. i - xxxi + ix - xvi + 1 - 243, pls. A - J + L - Q + 1 - 55." type="book chapter" year="1844">De Haan, 1844</bibRefCitation>
(
<bibRefCitation id="C61ECE8A4B72FFBEE43AFA66FC59FA7A" author="Dotu, Y." box="[789,914,1483,1509]" pageId="54" pageNumber="55" pagination="133 - 139" refId="ref71162" refString="Dotu, Y. (1961) The bionomics and life history of the gobioid fish, Apocryptodon bleekeri (Day). Bulletin of the Faculty of Fisheries, Nagasaki University, 10, 133 - 139, pl. 28. [in Japanese]" type="journal article" year="1961">Dôtu 1961</bibRefCitation>
)
</taxonomicName>
and the presumably more closely related 
<taxonomicName id="658FC8F84B72FFBEE2ABFA66FED2F996" authorityName="Yaldwyn" authorityYear="1971" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE2ABFA66FED2F996" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B72FFBEE609FA42FE68F996" author="Miya, Y." box="[294,419,1518,1545]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
, as 
<taxonomicName id="658FC8F84B72FFBEE6E5FA42FD24F996" authorityName="Yaldwyn" authorityYear="1971" box="[458,751,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">
<emphasis id="90FB6F694B72FFBEE6E5FA42FD24F996" box="[458,751,1519,1545]" italics="true" pageId="54" pageNumber="55">A. euphrosyne richardsoni</emphasis>
</taxonomicName>
). The morphological differences between 
<taxonomicName id="658FC8F84B72FFBEE3EDFA42FAF4F997" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1218,1343,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE3EDFA42FAF4F997" box="[1218,1343,1519,1545]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE26BFA42FA57F996" bold="true" box="[1348,1436,1519,1545]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE26BFA42FA57F996" box="[1348,1436,1519,1545]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="658FC8F84B72FFBEE7E6F9BEFEA7F9B2" authorityName="Yaldwyn" authorityYear="1971" box="[201,364,1555,1581]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE7E6F9BEFEA7F9B2" box="[201,364,1555,1581]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
are relatively subtle, which explains 
<bibRefCitation id="C61ECE8A4B72FFBEE437F9BEFC72F9B2" author="Miya, Y." box="[792,953,1554,1581]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya’s (1995)</bibRefCitation>
identification of the Japanese and Korean material as 
<taxonomicName id="658FC8F84B72FFBEE632F99AFD80F9CE" authorityName="Yaldwyn" authorityYear="1971" box="[285,587,1591,1617]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="richardsoni">
<emphasis id="90FB6F694B72FFBEE632F99AFD80F9CE" box="[285,587,1591,1617]" italics="true" pageId="54" pageNumber="55">A. euphrosyne richardsoni</emphasis>
</taxonomicName>
, despite the resulting disjunct, antitropical distribution for 
<taxonomicName id="658FC8F84B72FFBEE3DDF99AFA5DF9CE" authorityName="Yaldwyn" authorityYear="1971" box="[1266,1430,1591,1617]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE3DDF99AFA5DF9CE" box="[1266,1430,1591,1617]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
. Nevertheless, 
<taxonomicName id="658FC8F84B72FFBEE619F9F6FE7FF9EB" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[310,436,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE619F9F6FE7FF9EB" box="[310,436,1627,1653]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE695F9F6FDDFF9EA" bold="true" box="[442,532,1627,1653]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE695F9F6FDDFF9EA" box="[442,532,1627,1653]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="658FC8F84B72FFBEE565F9F6FD20F9EA" authorityName="Yaldwyn" authorityYear="1971" box="[586,747,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE565F9F6FD20F9EA" box="[586,747,1627,1653]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
can be separated from each other by the shape of the dactylar plunger of the major chela, which appears to broader and stouter in 
<taxonomicName id="658FC8F84B72FFBEE4BDF9D2FBDAF907" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[914,1041,1663,1689]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE4BDF9D2FBDAF907" box="[914,1041,1663,1689]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE337F9D2FBB9F906" bold="true" box="[1048,1138,1663,1689]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE337F9D2FBB9F906" box="[1048,1138,1663,1689]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(cf. 
<figureCitation id="3AB4AFFE4B72FFBEE388F9D2FAC5F906" box="[1191,1294,1663,1689]" captionStart="FIGURE 27" captionStartId="52.[151,250,1818,1842]" captionTargetBox="[162,1424,195,1781]" captionTargetId="figure-21@52.[151,1436,181,1794]" captionTargetPageId="52" captionText="FIGURE 27. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588) [A–G]; paratype, ovigerous female (cl 8.8 mm) from the same locality (MNHN-IU-2018-5744) [H]; paratype, ovigerous female (cl 11.9 mm) from Shimanto River, Japan (MNHN-IU-2018-5589) [I]; A—major (right) cheliped, chela, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor cheliped, lateral view; same, carpus and chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912380" httpUri="https://zenodo.org/record/7912380/files/figure.png" pageId="54" pageNumber="55">Fig. 27D</figureCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE235F9D2FA53F906" author="Miya, Y." box="[1306,1432,1662,1689]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
: fig. 2C; 
<bibRefCitation id="C61ECE8A4B72FFBEE7D8F90EFDCEF922" author="Banner, D. M. &amp; Banner, A. H." box="[247,517,1699,1725]" pageId="54" pageNumber="55" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 74e); the presence of a small subdistal mesial ridge in 
<taxonomicName id="658FC8F84B72FFBEE39DF90EFAFAF923" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1202,1329,1699,1725]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE39DF90EFAFAF923" box="[1202,1329,1699,1725]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE215F90EFA5EF922" bold="true" box="[1338,1429,1699,1725]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE215F90EFA5EF922" box="[1338,1429,1699,1725]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is not distinct in 
<taxonomicName id="658FC8F84B72FFBEE68FF96AFD89F97E" authorityName="Yaldwyn" authorityYear="1971" box="[416,578,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE68FF96AFD89F97E" box="[416,578,1735,1761]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
(cf. 
<figureCitation id="3AB4AFFE4B72FFBEE559F96AFD17F97E" box="[630,732,1735,1761]" captionStart="FIGURE 27" captionStartId="52.[151,250,1818,1842]" captionTargetBox="[162,1424,195,1781]" captionTargetId="figure-21@52.[151,1436,181,1794]" captionTargetPageId="52" captionText="FIGURE 27. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588) [A–G]; paratype, ovigerous female (cl 8.8 mm) from the same locality (MNHN-IU-2018-5744) [H]; paratype, ovigerous female (cl 11.9 mm) from Shimanto River, Japan (MNHN-IU-2018-5589) [I]; A—major (right) cheliped, chela, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor cheliped, lateral view; same, carpus and chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912380" httpUri="https://zenodo.org/record/7912380/files/figure.png" pageId="54" pageNumber="55">Fig. 27C</figureCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE5C7F96AFC39F97E" author="Banner, D. M. &amp; Banner, A. H." box="[744,1010,1735,1761]" pageId="54" pageNumber="55" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 74c); the size and position of the tooth on the distolateral margin of the antennal basicerite, which is more developed and more ventral in 
<taxonomicName id="658FC8F84B72FFBEE230F946FA57F89B" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1311,1436,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE230F946FA57F89B" box="[1311,1436,1771,1797]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE7B8F8A2FF24F8B6" bold="true" box="[151,239,1807,1833]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE7B8F8A2FF24F8B6" box="[151,239,1807,1833]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(cf. 
<figureCitation id="3AB4AFFE4B72FFBEE60EF8A2FE4DF8B6" box="[289,390,1807,1833]" captionStart="FIGURE 26" captionStartId="51.[151,250,1813,1837]" captionTargetBox="[167,1425,193,1775]" captionTargetId="figure-15@51.[151,1436,181,1789]" captionTargetPageId="51" captionText="FIGURE 26. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D— antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—fifth pereiopod, lateral view; J—same, propodus and dactylus, mesial view; K—second pleopod, appendices masculina and interna, anterior view; L—left uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912376" httpUri="https://zenodo.org/record/7912376/files/figure.png" pageId="54" pageNumber="55">Fig. 26D</figureCitation>
; Miya 1996: fig. 2A; 
<bibRefCitation id="C61ECE8A4B72FFBEE55CF8A2FCBCF8B6" author="Banner, D. M. &amp; Banner, A. H." box="[627,887,1807,1833]" pageId="54" pageNumber="55" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 74b); and the armature of the third pereiopod propodus, which has only three to four spiniform setae in 
<taxonomicName id="658FC8F84B72FFBEE40BF89EFC68F8D3" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[804,931,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE40BF89EFC68F8D3" box="[804,931,1843,1869]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE485F89EFBCFF8D2" bold="true" box="[938,1028,1843,1869]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE485F89EFBCFF8D2" box="[938,1028,1843,1869]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="90FB6F694B72FFBEE323F89EFBE8F8D3" box="[1036,1059,1843,1868]" italics="true" pageId="54" pageNumber="55">vs</emphasis>
. at least six in 
<taxonomicName id="658FC8F84B72FFBEE3E2F89EFAA4F8D2" authorityName="Yaldwyn" authorityYear="1971" box="[1229,1391,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE3E2F89EFAA4F8D2" box="[1229,1391,1843,1869]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
(cf. 
<figureCitation id="3AB4AFFE4B72FFBEE7B8F8FAFF35F8EE" box="[151,254,1879,1905]" captionStart="FIGURE 26" captionStartId="51.[151,250,1813,1837]" captionTargetBox="[167,1425,193,1775]" captionTargetId="figure-15@51.[151,1436,181,1789]" captionTargetPageId="51" captionText="FIGURE 26. Alpheus nomurai sp. nov.: paratype, male (cl 8.5 mm) from Tanabe, Japan (MNHN-IU-2018-5588); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D— antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—fifth pereiopod, lateral view; J—same, propodus and dactylus, mesial view; K—second pleopod, appendices masculina and interna, anterior view; L—left uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912376" httpUri="https://zenodo.org/record/7912376/files/figure.png" pageId="54" pageNumber="55">Fig. 26G</figureCitation>
; 
<bibRefCitation id="C61ECE8A4B72FFBEE626F8FAFE4CF8EE" author="Miya, Y." box="[265,391,1878,1905]" pageId="54" pageNumber="55" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
: fig. 2F; 
<bibRefCitation id="C61ECE8A4B72FFBEE6C4F8FAFD38F8EE" author="Banner, D. M. &amp; Banner, A. H." box="[491,755,1879,1905]" pageId="54" pageNumber="55" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 74n). In addition, 
<taxonomicName id="658FC8F84B72FFBEE4DBF8FAFBB9F8EF" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1012,1138,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE4DBF8FAFBB9F8EF" box="[1012,1138,1879,1905]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE357F8FAFB19F8EE" bold="true" box="[1144,1234,1879,1905]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE357F8FAFB19F8EE" box="[1144,1234,1879,1905]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with the maximal total length not exceeding 
<quantity id="65771E9E4B72FFBEE692F8D7FDEBF80A" box="[445,544,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="54" pageNumber="55" unit="mm" value="36.5">36.5 mm</quantity>
is a much smaller species compared to 
<taxonomicName id="658FC8F84B72FFBEE4F9F8D6FBBCF80A" authorityName="Yaldwyn" authorityYear="1971" box="[982,1143,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B72FFBEE4F9F8D6FBBCF80A" box="[982,1143,1915,1941]" italics="true" pageId="54" pageNumber="55">A. richardsoni</emphasis>
</taxonomicName>
, which may reach 
<quantity id="65771E9E4B72FFBEE268F8D7FA52F80A" box="[1351,1433,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="54" pageNumber="55" unit="mm" value="65.0">65 mm</quantity>
. The most obvious and reliable distinguishing feature between these two species involves the banding pattern of the pleon: in 
<taxonomicName id="658FC8F84B72FFBEE62EF86EFE4BF843" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[257,384,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B72FFBEE62EF86EFE4BF843" box="[257,384,1987,2013]" italics="true" pageId="54" pageNumber="55">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B72FFBEE6A8F86EFE2AF842" bold="true" box="[391,481,1987,2013]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="8BC8D2124B72FFBEE6A8F86EFE2AF842" box="[391,481,1987,2013]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, each pleonite has a simple dark transverse band (
<figureCitation id="3AB4AFFE4B72FFBEE33BF86EFBACF842" box="[1044,1127,1987,2013]" captionStart="FIGURE 28" captionStartId="55.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1790]" captionTargetId="figure-15@55.[151,1435,181,1790]" captionTargetPageId="55" captionText="FIGURE 28. Alpheus nomurai sp. nov.: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7912382" httpUri="https://zenodo.org/record/7912382/files/figure.png" pageId="54" pageNumber="55">Fig. 28</figureCitation>
, disregarding diffuse bands formed by neon-yellow spots), whereas in 
<taxonomicName id="658FC8F84B7CFFB0E558FF3AFCD2FF2E" authorityName="Yaldwyn" authorityYear="1971" box="[631,793,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B7CFFB0E558FF3AFCD2FF2E" box="[631,793,151,177]" italics="true" pageId="56" pageNumber="57">A. richardsoni</emphasis>
</taxonomicName>
, each pleonite has a double or even triple band (
<figureCitation id="3AB4AFFE4B7CFFB0E215FF3AFA5CFF2E" box="[1338,1431,151,177]" captionStart="FIGURE 17" captionStartId="35.[151,250,1930,1954]" captionTargetBox="[151,1435,237,1907]" captionTargetId="figure-46@35.[151,1435,237,1907]" captionTargetPageId="35" captionText="FIGURE 17. Alpheus richardsoni Yaldwyn, 1971: male (cl 8.3 mm) from Loders Creek, Moreton Bay, Queensland, Australia (FLMNH UF 18694) [A]; male (cl 9.3 mm) from Cleveland, Queensland, Australia (OUMNH.ZC 2009-09-0015); A, B— habitus, dorsal view; C—same, lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.7912344" httpUri="https://zenodo.org/record/7912344/files/figure.png" pageId="56" pageNumber="57">Figs. 17</figureCitation>
, 
<figureCitation id="3AB4AFFE4B7CFFB0E7B8FF16FF7DFF4A" box="[151,182,187,213]" captionStart="FIGURE 18" captionStartId="36.[151,250,1909,1933]" captionTargetBox="[221,1366,181,1885]" captionTargetId="figure-21@36.[221,1366,181,1885]" captionTargetPageId="36" captionText="FIGURE 18. Alpheus richardsoni Yaldwyn, 1971: male (cl indet.) from Brunswisk River, New South Wales, Australia [A, B]; male from Parengarenga (Paua), New Zealand [C]; male from Whangarei, New Zealand [D]; A, C—habitus, dorsal view; B—same, lateral view; D—tail fan, dorsal view. Specimens not deposited / deposition unknown. Photographs courtesy of Denis Riek (A, B) and Richard B. Taylor (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7912350" httpUri="https://zenodo.org/record/7912350/files/figure.png" pageId="56" pageNumber="57">18</figureCitation>
). Of course, the two species are also geographically widely separated, with 
<taxonomicName id="658FC8F84B7CFFB0E4DEFF16FBA5FF4B" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1009,1134,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B7CFFB0E4DEFF16FBA5FF4B" box="[1009,1134,187,213]" italics="true" pageId="56" pageNumber="57">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B7CFFB0E35CFF16FB01FF4A" bold="true" box="[1139,1226,187,213]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="8BC8D2124B7CFFB0E35CFF16FB01FF4A" box="[1139,1226,187,213]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="658FC8F84B7CFFB0E3D2FF16FA56FF4A" authorityName="Yaldwyn" authorityYear="1971" box="[1277,1437,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B7CFFB0E3D2FF16FA56FF4A" box="[1277,1437,187,213]" italics="true" pageId="56" pageNumber="57">A. richardsoni</emphasis>
</taxonomicName>
each restricted to temperate and subtropical north-eastern Asia and 
<collectingCountry id="DA98F3EB4B7CFFB0E4AFFF72FC2CFF66" box="[896,999,223,249]" name="Australia" pageId="56" pageNumber="57">Australia</collectingCountry>
/ 
<collectingCountry id="DA98F3EB4B7CFFB0E4D4FF72FB58FF66" box="[1019,1171,223,249]" name="New Zealand" pageId="56" pageNumber="57">New Zealand</collectingCountry>
, respectively (
<figureCitation id="3AB4AFFE4B7CFFB0E21CFF72FF61FE82" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="56" pageNumber="57">Fig. 51C, F</figureCitation>
).
</paragraph>
<caption id="F6F0E3F34B73FFBFE7B8F8BBFB13F804" ID-DOI="http://doi.org/10.5281/zenodo.7912382" ID-Zenodo-Dep="7912382" httpUri="https://zenodo.org/record/7912382/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1814,1838]" targetBox="[151,1435,181,1790]" targetPageId="55" targetType="figure">
<paragraph id="A230B37B4B73FFBFE7B8F8BBFB13F804" blockId="55.[151,1437,1814,1947]" pageId="55" pageNumber="56">
<emphasis id="90FB6F694B73FFBFE7B8F8BBFED3F8B1" bold="true" box="[151,280,1814,1838]" pageId="55" pageNumber="56">FIGURE 28</emphasis>
. 
<taxonomicName id="658FC8F84B73FFBFE60AF8BAFE19F8B1" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[293,466,1815,1839]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B73FFBFE60AF8BAFE19F8B1" box="[293,466,1815,1839]" italics="true" pageId="55" pageNumber="56">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B73FFBFE6F7F8BAFDECF8B0" bold="true" box="[472,551,1815,1839]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="8BC8D2124B73FFBFE6F7F8BAFDECF8B0" box="[472,551,1815,1839]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D).
</paragraph>
</caption>
<paragraph id="A230B37B4B7CFFB0E7E8FE8AFD50FE4E" blockId="56.[151,1437,151,465]" pageId="56" pageNumber="57">
<taxonomicName id="658FC8F84B7CFFB0E7E8FE8AFE41FEDF" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[199,394,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B7CFFB0E7E8FE8AFE41FEDF" box="[199,394,295,321]" italics="true" pageId="56" pageNumber="57">Alpheus nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B7CFFB0E6BCFE8AFE25FEDE" bold="true" box="[403,494,295,321]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="8BC8D2124B7CFFB0E6BCFE8AFE25FEDE" box="[403,494,295,321]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="658FC8F84B7CFFB0E5D2FE8AFC54FEDE" authorityName="sensu De Man" authorityYear="1897" box="[765,927,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B7CFFB0E5D2FE8AFC54FEDE" box="[765,927,295,321]" italics="true" pageId="56" pageNumber="57">A. euphrosyne</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B7CFFB0E4F6FE8AFB41FEDE" authorityName="De Man" authorityYear="1920" box="[985,1162,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B7CFFB0E4F6FE8AFB41FEDE" box="[985,1162,295,321]" italics="true" pageId="56" pageNumber="57">A. eurydactylus</emphasis>
</taxonomicName>
essentially by the same criteria as 
<taxonomicName id="658FC8F84B7CFFB0E626FEE6FE63FEFA" authorityName="Yaldwyn" authorityYear="1971" box="[265,424,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B7CFFB0E626FEE6FE63FEFA" box="[265,424,331,357]" italics="true" pageId="56" pageNumber="57">A. richardsoni</emphasis>
</taxonomicName>
(see above), and from 
<taxonomicName id="658FC8F84B7CFFB0E58DFEE6FCAEFEFA" baseAuthorityName="De Man" baseAuthorityYear="1897" box="[674,869,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B7CFFB0E58DFEE6FCAEFEFA" box="[674,869,331,357]" italics="true" pageId="56" pageNumber="57">A. microrhynchus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B7CFFB0E4B6FEE6FBE7FEFA" authorityName="Yeo &amp; Ng" authorityYear="1996" box="[921,1068,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cyanoteles">
<emphasis id="90FB6F694B7CFFB0E4B6FEE6FBE7FEFA" box="[921,1068,331,357]" italics="true" pageId="56" pageNumber="57">A. cyanoteles</emphasis>
</taxonomicName>
by the conspicuously broadened, spatulate dactylus of the third to fifth pereiopods, which in the other two species is trigonal-subspatulate in shape (cf. 
<bibRefCitation id="C61ECE8A4B7CFFB0E7B8FE3EFEDDFE32" author="Miya, Y." box="[151,278,402,429]" pageId="56" pageNumber="57" pagination="271 - 288" refId="ref73081" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995</bibRefCitation>
: fig. 2F; 
<bibRefCitation id="C61ECE8A4B7CFFB0E656FE3EFDE4FE32" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[377,559,403,429]" pageId="56" pageNumber="57" pagination="37 - 63" refId="ref75202" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (1996) A new species of freshwater snapping shrimp, Alpheus cyanoteles (Decapoda: Caridea: Alpheidae) from Peninsular Malaysia and a redescription of Alpheus paludicola Kemp, 1915. Raffles Bulletin of Zoology, 44, 37 - 63." type="journal article" year="1996">Yeo &amp; Ng 1996</bibRefCitation>
: fig. 6b, d). For separation of 
<taxonomicName id="658FC8F84B7CFFB0E451FE3EFC36FE33" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[894,1021,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B7CFFB0E451FE3EFC36FE33" box="[894,1021,403,429]" italics="true" pageId="56" pageNumber="57">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B7CFFB0E32BFE3EFB95FE32" bold="true" box="[1028,1118,403,429]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="8BC8D2124B7CFFB0E32BFE3EFB95FE32" box="[1028,1118,403,429]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from other morphologically similar species see below and refer to 
<tableCitation id="EF0D86C04B7CFFB0E56DFE1AFD5EFE4E" box="[578,661,439,465]" captionStart="TABLE 1" captionStartId="93.[145,233,465,491]" captionTargetPageId="93" captionText="TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897—A. micro-" httpUri="http://table.plazi.org/id/F6F0E3F34B19FFD5E7BEFE7CF8F8FD90" pageId="56" pageNumber="57" tableUuid="F6F0E3F34B19FFD5E7BEFE7CF8F8FD90">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>