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<mods:title id="5434B4A321A5C5AC5ECBD02F8713B786">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<taxonomicName id="658FC8F84B61FFADE7B8FB7CFDA4FB73" ID-CoL="85LLH" ID-ENA="1240033" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[151,623,1233,1260]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">
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<emphasis id="90FB6F694B61FFADE7B8FB7CFE71FB73" bold="true" box="[151,442,1233,1260]" italics="true" pageId="37" pageNumber="38">Alpheus microrhynchus</emphasis>
<bibRefCitation id="C61ECE8A4B61FFADE6EEFB7FFDA4FB73" author="De Man, J. G." box="[449,623,1234,1260]" pageId="37" pageNumber="38" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
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<paragraph id="A230B37B4B61FFADE7B8FB5BFE70FA8E" blockId="37.[151,623,1233,1297]" box="[151,443,1270,1297]" pageId="37" pageNumber="38">
(
<figureCitation id="3AB4AFFE4B61FFADE7B0FB5BFEE8FA8F" box="[159,291,1270,1296]" captionStart="FIGURE 2" captionStartId="7.[151,250,1827,1851]" captionTargetBox="[180,1403,388,1787]" captionTargetId="figure-118@7.[168,1419,381,1804]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus microrhynchus DeMan, 1897:syntype, incomplete male (cl indet., tl 40.0 mm) from Pontianak, Kalimantan, Indonesia (after De Man 1897, as Alpheus sp. and A. microrhynchus, specimen lost) [AC]; female (cl indet., tl 43.0 mm) from Bangkok, Thailand (after De Man 1898) [D, E]; female (tl 85.0 mm) from Makassar, Sulawesi, Indonesia (after De Man 1911), specimen tentatively assigned to A. takla sp. nov. (see text) [F, G]: A—frontal region, dorsal view; B—male minor (left) chela, dorsal view; C—same, lateral view; D—female minor (right) cheliped, mesial view; E—female major (left) cheliped, lateral view; F—frontal area of carapace, dorsal view; G—telson, dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7912298" httpUri="https://zenodo.org/record/7912298/files/figure.png" pageId="37" pageNumber="38">Figs. 2AE</figureCitation>
,
<figureCitation id="3AB4AFFE4B61FFADE601FB5BFEBCFA8F" box="[302,375,1270,1296]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStart-2="FIGURE 21" captionStart-3="FIGURE 22" captionStartId-0="39.[151,250,1944,1968]" captionStartId-1="40.[151,249,1647,1671]" captionStartId-2="43.[151,250,1724,1748]" captionStartId-3="44.[151,250,1907,1931]" captionTargetBox-0="[159,1427,181,1920]" captionTargetBox-1="[164,1423,199,1623]" captionTargetBox-2="[151,1435,196,1685]" captionTargetBox-3="[226,1361,193,1884]" captionTargetId-0="figure-15@39.[159,1427,181,1920]" captionTargetId-1="figure-21@40.[164,1423,199,1623]" captionTargetId-2="figure-15@43.[151,1435,196,1685]" captionTargetId-3="figure-21@44.[226,1361,181,1884]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="43" captionTargetPageId-3="44" captionText-0="FIGURE 19. Alpheus microrhynchus De Man, 1897: neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped, lateral view; D—same, mesial view; E—minor (left) cheliped, lateral view; F—same, mesial view. Refer to A for scale." captionText-1="FIGURE 20. Alpheus microrhynchus De Man, 1897: male (cl 17.1 mm) from near Ho Chi Minh City, Vietnam (MNHN-IU- 2013-14789) [AD]; neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193) [E]; A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped in situ, mesial view; D—minor (left) cheliped in situ, lateral view; E—minor (left) cheliped, lateral view. Refer to A for scale in BD, and to Fig. 19A for scale in E." captionText-2="FIGURE 21. Alpheus microrhynchus De Man, 1897: ovigerous female (cl indet.) imported to Germany from Thailand [A]; ovigerous female (cl indet.) imported to Singapore from Thailand [BE]: A, B—habitus, lateral view; C—pleon and tail fan, dorsal view; D—pleon, ventrolateral view E—frontal region and chelipeds, dorsal view. Aquarium-kept specimens, probably not preserved / deposited. Note the low number and large size of developing embryos inA and D. Photographs courtesy of Chris Lukhaup (A) and Paul Ng (BE)." captionText-3="FIGURE 22. Alpheus microrhynchus De Man, 1897: female (cl indet.) imported to Singapore from Thailand; A, B—habitus, dorsal view (with one Singapore dollar coin, diameter 22.4 mm, for scale in B); C—detail of frontal margin; D—major (right) chela, mesial view; E—posterior pleonites and tail fan, d orsal view. Aquarium-kept specimen, probably not preserved / deposited. Photographs courtesy of Paul Ng." figureDoi-0="http://doi.org/10.5281/zenodo.7912352" figureDoi-1="http://doi.org/10.5281/zenodo.7912354" figureDoi-2="http://doi.org/10.5281/zenodo.7912358" figureDoi-3="http://doi.org/10.5281/zenodo.7912362" httpUri-0="https://zenodo.org/record/7912352/files/figure.png" httpUri-1="https://zenodo.org/record/7912354/files/figure.png" httpUri-2="https://zenodo.org/record/7912358/files/figure.png" httpUri-3="https://zenodo.org/record/7912362/files/figure.png" pageId="37" pageNumber="38">1922</figureCitation>
,
<figureCitation id="3AB4AFFE4B61FFADE6ADFB5BFE7FFA8E" box="[386,436,1270,1297]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="37" pageNumber="38">51D</figureCitation>
)
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<treatmentCitation id="232E956A4B61FFADE7B8FA97FD86FACD" author="De Man, J. G." box="[151,589,1338,1362]" page="754" pageId="37" pageNumber="38" year="1897">
<taxonomicName id="658FC8F84B61FFADE7B8FA97FD86FACD" ID-CoL="85LLH" authority="De Man 1897: 754" authorityName="sensu De Man" authorityPageNumber="754" authorityYear="1897" box="[151,589,1338,1362]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B61FFADE7B8FA97FE4DFACD" box="[151,390,1338,1362]" italics="true" pageId="37" pageNumber="38">Alpheus microrhynchus</emphasis>
<bibRefCitation id="C61ECE8A4B61FFADE6A3FA96FD86FACD" author="De Man, J. G." box="[396,589,1338,1362]" pageId="37" pageNumber="38" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897: 754</bibRefCitation>
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, pl. 36, fig. 65 (as
<taxonomicName id="658FC8F84B61FFADE425FA97FC4AFACD" box="[778,897,1338,1362]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B61FFADE425FA97FC97FACD" box="[778,860,1338,1362]" italics="true" pageId="37" pageNumber="38">Alpheus</emphasis>
sp.
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).
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<taxonomicName id="658FC8F84B61FFADE7B8FAF4FE42FAEE" ID-CoL="85LLH" authorityName="sensu De Man" authorityYear="1897" box="[151,393,1369,1393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">Alpheus microrhynchus</taxonomicName>
.
</emphasis>
<treatmentCitation id="232E956A4B61FFADE689FAF4FDADFAEE" author="De Man, J. G." box="[422,614,1369,1393]" page="318" pageId="37" pageNumber="38" year="1898">
<bibRefCitation id="C61ECE8A4B61FFADE689FAF4FDADFAEE" author="De Man, J. G." box="[422,614,1369,1393]" pageId="37" pageNumber="38" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man 1898: 318</bibRefCitation>
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, pl. 4, fig. 3;
<treatmentCitation id="232E956A4B61FFADE5C3FAF4FC57FAEE" author="Nobili, G." box="[748,924,1369,1393]" page="479" pageId="37" pageNumber="38" year="1900">
<bibRefCitation id="C61ECE8A4B61FFADE5C3FAF4FC57FAEE" author="Nobili, G." box="[748,924,1369,1393]" pageId="37" pageNumber="38" pagination="473 - 523" refId="ref73484" refString="Nobili, G. (1900) Decapodi e stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale di Genova, 40, 473 - 523." type="journal article" year="1900">Nobili 1900: 479</bibRefCitation>
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;
<treatmentCitation id="232E956A4B61FFADE488FAF4FB86FAEE" author="Johnson, D. S." box="[935,1101,1369,1393]" page="9" pageId="37" pageNumber="38" year="1965">
<bibRefCitation id="C61ECE8A4B61FFADE488FAF4FB86FAEE" author="Johnson, D. S." box="[935,1101,1369,1393]" pageId="37" pageNumber="38" pagination="7 - 11" refId="ref72065" refString="Johnson, D. S. (1965) A review of brackish water prawns of Malaya. Bulletin of the National Museum of Singapore, 33, 7 - 11." type="journal article" year="1965">Johnson 1965: 9</bibRefCitation>
</treatmentCitation>
(part.?);
<treatmentCitation id="232E956A4B61FFADE385FAF4FF26FA0F" author="Banner, A. H. &amp; Banner, D. M." page="133" pageId="37" pageNumber="38" year="1966">
<bibRefCitation id="C61ECE8A4B61FFADE385FAF4FF26FA0F" author="Banner, A. H. &amp; Banner, D. M." pageId="37" pageNumber="38" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966: 133</bibRefCitation>
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(part.?), fig. 50;
<treatmentCitation id="232E956A4B61FFADE6B5FAD5FD87FA0F" author="Johnson, D. S." box="[410,588,1400,1424]" page="39" pageId="37" pageNumber="38" year="1976">
<bibRefCitation id="C61ECE8A4B61FFADE6B5FAD5FD87FA0F" author="Johnson, D. S." box="[410,588,1400,1424]" pageId="37" pageNumber="38" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">Johnson 1976: 39</bibRefCitation>
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(part.?);
<treatmentCitation id="232E956A4B61FFADE585FAD5FC77FA0F" author="Banner, D. M. &amp; Banner, A. H." box="[682,956,1399,1424]" page="24" pageId="37" pageNumber="38" year="1985">
<bibRefCitation id="C61ECE8A4B61FFADE585FAD5FC77FA0F" author="Banner, D. M. &amp; Banner, A. H." box="[682,956,1399,1424]" pageId="37" pageNumber="38" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. &amp; Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II. Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner &amp; Banner 1985: 24</bibRefCitation>
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(footnote);
<treatmentCitation id="232E956A4B61FFADE31AFAD5FAC2FA0F" author="Naiyanetr, P." box="[1077,1289,1400,1424]" page="173" pageId="37" pageNumber="38" year="2007">
<bibRefCitation id="C61ECE8A4B61FFADE31AFAD5FAC2FA0F" author="Naiyanetr, P." box="[1077,1289,1400,1424]" pageId="37" pageNumber="38" refId="ref73355" refString="Naiyanetr, P. (2007) Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr 2007: 173</bibRefCitation>
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.
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<treatmentCitationGroup id="829F94554B61FFADE7B8FA3AFD91FA31" box="[151,602,1430,1454]" pageId="37" pageNumber="38">
<emphasis id="90FB6F694B61FFADE7B8FA3AFE60FA31" box="[151,427,1430,1454]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="658FC8F84B61FFADE7B8FA3AFE45FA31" authority="Suvatti 1937" box="[151,398,1430,1454]" class="Malacostraca" family="Alpheidae" genus="Crangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">Crangon microrhynchus</taxonomicName>
.—
</emphasis>
<treatmentCitation id="232E956A4B61FFADE684FA3BFD9EFA31" author="Suvatti, C." box="[427,597,1430,1454]" page="48" pageId="37" pageNumber="38" year="1937">
<bibRefCitation id="C61ECE8A4B61FFADE684FA3BFD9EFA31" author="Suvatti, C." box="[427,597,1430,1454]" pageId="37" pageNumber="38" refId="ref74577" refString="Suvatti, C. (1937) A check-list of aquatic fauna of Siam (excluding fishes). Bureau of Fisheries, Bangkok, 116 pp." type="book" year="1937">Suvatti 1937: 48</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B61FFADE7B8FA18FC52FA52" blockId="37.[151,1437,1338,1577]" box="[151,921,1461,1485]" pageId="37" pageNumber="38">
(?)
<emphasis id="90FB6F694B61FFADE795FA18FE4EFA52" box="[186,389,1461,1485]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="658FC8F84B61FFADE795FA18FE49FA52" ID-CoL="85LMD" baseAuthorityName="Yeo &amp; Ng" baseAuthorityYear="1996" box="[186,386,1461,1485]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="paludicola">Alpheus paludicola</taxonomicName>
.
</emphasis>
<bibRefCitation id="C61ECE8A4B61FFADE6B1FA18FD83FA52" author="Kemp, S." box="[414,584,1461,1485]" pageId="37" pageNumber="38" pagination="81 - 102" refId="ref72298" refString="Kemp, S. (1918) Crustacea Decapoda of the Inle Lake Basin. Records of the Indian Museum, 14, 81 - 102, pls. 24 + 25. https: // doi. org / 10.26515 / rzsi / v 14 / i 4 / 1918 / 163580" type="journal article" year="1918">Kemp 1918: 273</bibRefCitation>
(not
<taxonomicName id="658FC8F84B61FFADE552FA18FC46FA52" ID-CoL="85LMD" authority="Kemp, 1915" authorityName="Kemp" authorityYear="1915" box="[637,909,1461,1485]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="90FB6F694B61FFADE552FA18FCCDFA52" box="[637,774,1461,1485]" italics="true" pageId="37" pageNumber="38">A. paludicola</emphasis>
<bibRefCitation id="C61ECE8A4B61FFADE423FA18FC46FA52" author="Kemp, S." box="[780,909,1461,1485]" pageId="37" pageNumber="38" pagination="201 - 325" refId="ref72246" refString="Kemp, S. (1915) Fauna of the Chilka Lake. No. 3. Crustacea Decapoda. Memoirs of the Indian Museum, 5, 201 - 325, pls. 12 - 13. https: // doi. org / 10.5962 / bhl. title. 10414" type="journal article" year="1915">Kemp, 1915</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="A230B37B4B61FFADE7B8FA79FEC9F9B7" blockId="37.[151,1437,1338,1577]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="829F94554B61FFADE7B8FA79FEC9F9B7" pageId="37" pageNumber="38">
<collectingRegion id="604B7D994B61FFADE7B8FA79FF76FA74" box="[151,189,1492,1515]" country="Maldives" name="Uthuru" pageId="37" pageNumber="38">Not</collectingRegion>
<emphasis id="90FB6F694B61FFADE7EBFA79FE71FA73" box="[196,442,1492,1516]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="658FC8F84B61FFADE7EBFA79FE7DFA73" ID-CoL="85LLH" authorityName="sensu De Man" authorityYear="1897" box="[196,438,1492,1516]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">Alpheus microrhynchus</taxonomicName>
.
</emphasis>
<treatmentCitation id="232E956A4B61FFADE6FCFA79FD5AFA73" author="De Man, J. G." box="[467,657,1492,1516]" page="413" pageId="37" pageNumber="38" year="1911">
<bibRefCitation id="C61ECE8A4B61FFADE6FCFA79FD5AFA73" author="De Man, J. G." box="[467,657,1492,1516]" pageId="37" pageNumber="38" pagination="133 - 465" refId="ref70992" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga-Expeditie, 39 a 1 (2), 133 - 465; (1915) Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article" year="1911">De Man 1911: 413</bibRefCitation>
</treatmentCitation>
(part.), pl. 23, fig. 99 [=
<taxonomicName id="658FC8F84B61FFADE4BDFA79FC2AFA73" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[914,993,1492,1516]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B61FFADE4BDFA79FC2AFA73" box="[914,993,1492,1516]" italics="true" pageId="37" pageNumber="38">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B61FFADE4C7FA79FBF2FA73" bold="true" box="[1000,1081,1492,1516]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="8BC8D2124B61FFADE4C7FA79FBF2FA73" box="[1000,1081,1492,1516]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
];
<treatmentCitation id="232E956A4B61FFADE360FA79FB3DFA73" author="Johnson, D. S." box="[1103,1270,1491,1516]" page="9" pageId="37" pageNumber="38" year="1965">
<bibRefCitation id="C61ECE8A4B61FFADE360FA79FB3DFA73" author="Johnson, D. S." box="[1103,1270,1491,1516]" pageId="37" pageNumber="38" pagination="7 - 11" refId="ref72065" refString="Johnson, D. S. (1965) A review of brackish water prawns of Malaya. Bulletin of the National Museum of Singapore, 33, 7 - 11." type="journal article" year="1965">Johnson 1965: 9</bibRefCitation>
</treatmentCitation>
(part.);
<treatmentCitation id="232E956A4B61FFADE266FA79FEEAF995" author="Johnson, D. S." page="39" pageId="37" pageNumber="38" year="1976">
<bibRefCitation id="C61ECE8A4B61FFADE266FA79FEEAF995" author="Johnson, D. S." pageId="37" pageNumber="38" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">Johnson 1976: 39</bibRefCitation>
</treatmentCitation>
(part.?) [possibly =
<taxonomicName id="658FC8F84B61FFADE6DDFA5EFD89F995" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[498,578,1522,1546]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B61FFADE6DDFA5EFD89F995" box="[498,578,1522,1546]" italics="true" pageId="37" pageNumber="38">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B61FFADE566FA5FFD51F995" bold="true" box="[585,666,1522,1546]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="8BC8D2124B61FFADE566FA5FFD51F995" box="[585,666,1522,1546]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and/or
<taxonomicName id="658FC8F84B61FFADE5C6FA5EFCA8F995" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[745,867,1522,1546]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B61FFADE5C6FA5EFCA8F995" box="[745,867,1522,1546]" italics="true" pageId="37" pageNumber="38">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B61FFADE445FA5FFC70F995" bold="true" box="[874,955,1522,1546]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="8BC8D2124B61FFADE445FA5FFC70F995" box="[874,955,1522,1546]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
];
<treatmentCitation id="232E956A4B61FFADE4FEFA5EFB56F995" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[977,1181,1522,1547]" page="56" pageId="37" pageNumber="38" year="1996">
<bibRefCitation id="C61ECE8A4B61FFADE4FEFA5EFB56F995" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[977,1181,1522,1547]" pageId="37" pageNumber="38" pagination="37 - 63" refId="ref75202" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (1996) A new species of freshwater snapping shrimp, Alpheus cyanoteles (Decapoda: Caridea: Alpheidae) from Peninsular Malaysia and a redescription of Alpheus paludicola Kemp, 1915. Raffles Bulletin of Zoology, 44, 37 - 63." type="journal article" year="1996">Yeo &amp; Ng 1996: 56</bibRefCitation>
</treatmentCitation>
, fig. 8a, b [=
<taxonomicName id="658FC8F84B61FFADE208FA5EFABCF995" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1319,1399,1522,1546]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B61FFADE208FA5EFABCF995" box="[1319,1399,1522,1546]" italics="true" pageId="37" pageNumber="38">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B61FFADE251FA5FFF38F9B6" bold="true" pageId="37" pageNumber="38">
<taxonomicNameLabel id="8BC8D2124B61FFADE251FA5FFF38F9B6" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B61FFADE7B8F9FAFCA1F868" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="A230B37B4B61FFADE7B8F9FAFB71F90B" blockId="37.[151,1437,1622,2040]" pageId="37" pageNumber="38">
<emphasis id="90FB6F694B61FFADE7B8F9FAFE8BF9EF" bold="true" box="[151,320,1622,1649]" pageId="37" pageNumber="38">Type material</emphasis>
.
<materialsCitation id="12E7B9264B61FFADE67FF9FAFB71F90B" ID-GBIF-Occurrence="4104078451" collectingDate="1998-04-18" collectionCode="ZRC" collectorName="H. H. Tan" country="Indonesia" location="Kalimantan Barat" pageId="37" pageNumber="38" specimenCode="ZRC 2000.2193" specimenCount="1" stateProvince="Kalimantan Barat" typeStatus="neotype">
<typeStatus id="7D340DD94B61FFADE67FF9FAFE79F9EE" box="[336,434,1623,1649]" pageId="37" pageNumber="38" type="neotype">Neotype</typeStatus>
, male (cl
<quantity id="65771E9E4B61FFADE509F9FAFD45F9EE" box="[550,654,1623,1649]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="37" pageNumber="38" unit="mm" value="16.3">16.3 mm</quantity>
, tl 37.0 mm, chl 21.0 mm),
<specimenCode id="F2291B004B61FFADE4F9F9FAFB44F9EE" box="[982,1167,1623,1649]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="37" pageNumber="38">ZRC 2000.2193</specimenCode>
,
<collectingCountry id="DA98F3EB4B61FFADE3B2F9FAFAC6F9EE" box="[1181,1293,1623,1649]" name="Indonesia" pageId="37" pageNumber="38">Indonesia</collectingCountry>
,
<collectingRegion id="604B7D994B61FFADE235F9FAFF1DF90A" country="Indonesia" name="Kalimantan Barat" pageId="37" pageNumber="38">Kalimantan Barat</collectingRegion>
, Kabupaten Sambas, Sambas Pasar, fish market, leg.
<collectorName id="0F7AD6AD4B61FFADE403F9D6FC5EF90A" box="[812,917,1659,1685]" pageId="37" pageNumber="38">H.H. Tan</collectorName>
,
<date id="D63195BB4B61FFADE48FF9D7FBD5F90B" box="[928,1054,1658,1684]" pageId="37" pageNumber="38" value="1998-04-18">
<collectingDate id="C6756C534B61FFADE48FF9D7FBD5F90B" box="[928,1054,1658,1684]" pageId="37" pageNumber="38" value="1998-04-18">18.04.1998</collectingDate>
</date>
[THH-9833].
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B61FFADE7E8F930FD39F8D9" blockId="37.[151,1437,1622,2040]" pageId="37" pageNumber="38">
<emphasis id="90FB6F694B61FFADE7E8F930FE78F928" bold="true" box="[199,435,1693,1719]" pageId="37" pageNumber="38">Additional material</emphasis>
.
<materialsCitation id="12E7B9264B61FFADE6EEF933FCD6F927" ID-GBIF-Occurrence="4104078479" box="[449,797,1693,1720]" country="Thailand" location="Thailand" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<emphasis id="90FB6F694B61FFADE6EEF933FDEDF927" box="[449,550,1694,1720]" italics="true" pageId="37" pageNumber="38">
<collectingCountry id="DA98F3EB4B61FFADE6EEF933FDEDF927" box="[449,550,1694,1720]" name="Thailand" pageId="37" pageNumber="38">Thailand</collectingCountry>
</emphasis>
:
<specimenCount id="B48978F24B61FFADE51AF933FD4AF927" box="[565,641,1694,1720]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(cl 15.0 mm)
</materialsCitation>
,
<materialsCitation id="12E7B9264B61FFADE407F933FC78F944" ID-GBIF-Occurrence="4104078413" collectionCode="OUMNH, ZC" collectorName="P. K. L. Ng" country="Thailand" location="Bangkok" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Krung Thep Mahanakhon">
<specimenCount id="B48978F24B61FFADE407F933FC41F927" box="[808,906,1694,1720]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(cl
<quantity id="65771E9E4B61FFADE499F933FBD7F927" box="[950,1052,1694,1720]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="37" pageNumber="38" unit="mm" value="14.9">14.9 mm</quantity>
, in poor condition),
<collectionCode id="C49E2BBE4B61FFADE22EF933FABAF927" box="[1281,1393,1694,1720]" pageId="37" pageNumber="38">OUMNH</collectionCode>
.
<collectionCode id="C49E2BBE4B61FFADE25CF933FA53F927" box="[1395,1432,1694,1720]" country="Austria" name="Zoological Collection, University of Vienna" pageId="37" pageNumber="38" type="University or college">ZC</collectionCode>
. 2019.06.49,
<collectingRegion id="604B7D994B61FFADE60BF96CFE47F944" box="[292,396,1729,1755]" country="Thailand" name="Krung Thep Mahanakhon" pageId="37" pageNumber="38">Bangkok</collectingRegion>
, aquarium trade market, leg.
<collectorName id="0F7AD6AD4B61FFADE5FAF96FFC83F943" box="[725,840,1730,1756]" pageId="37" pageNumber="38">P.K.L. Ng</collectorName>
, 06.2001
</materialsCitation>
;
<materialsCitation id="12E7B9264B61FFADE491F96CFF3DF8BD" ID-GBIF-Occurrence="4104078415" collectorName="F. Fasquel" country="Thailand" location="Bangkok" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Krung Thep Mahanakhon">
<specimenCount id="B48978F24B61FFADE491F96CFBC1F944" box="[958,1034,1729,1755]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(cl 13.4, in poor condition),
<specimenCount id="B48978F24B61FFADE27FF96CFA57F944" box="[1360,1436,1729,1755]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(
<quantity id="65771E9E4B61FFADE7B0F949FEC2F960" box="[159,265,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="37" pageNumber="38" unit="mm" value="16.8">16.8 mm</quantity>
, tl 45.0 mm, chl 28.0 mm), MNHN-IU-2018-5739,
<collectingRegion id="604B7D994B61FFADE44BF949FC07F961" box="[868,972,1764,1790]" country="Thailand" name="Krung Thep Mahanakhon" pageId="37" pageNumber="38">Bangkok</collectingRegion>
, aquarium trade market, leg.
<collectorName id="0F7AD6AD4B61FFADE20FF948FA53F961" box="[1312,1432,1764,1791]" pageId="37" pageNumber="38">F. Fasquel</collectorName>
, 06.2001
</materialsCitation>
;
<materialsCitation id="12E7B9264B61FFADE62DF8A5FD39F8D9" ID-GBIF-Occurrence="4104078385" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" location="Bangkok" pageId="37" pageNumber="38" specimenCode="ZRC 2012.0399" specimenCount="3" specimenCount-male="3" stateProvince="Krung Thep Mahanakhon">
<specimenCount id="B48978F24B61FFADE62DF8A5FE91F8BD" box="[258,346,1800,1826]" pageId="37" pageNumber="38" type="male">3 males</specimenCount>
(cl 12.613.0 mm), 1 ov. female (
<quantity id="65771E9E4B61FFADE5CCF8A5FC86F8BD" box="[739,845,1800,1826]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="37" pageNumber="38" unit="mm" value="13.1">13.1 mm</quantity>
),
<specimenCode id="F2291B004B61FFADE44DF8A5FBD0F8BD" box="[866,1051,1800,1826]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="37" pageNumber="38">ZRC 2012.0399</specimenCode>
, between
<collectingRegion id="604B7D994B61FFADE3A2F8A5FB39F8BD" box="[1165,1266,1800,1826]" country="Thailand" name="Krung Thep Mahanakhon" pageId="37" pageNumber="38">Bangkok</collectingRegion>
and Chonburi, river mouth, local collectors, leg.
<collectorName id="0F7AD6AD4B61FFADE53EF881FD4FF8D9" box="[529,644,1836,1862]" pageId="37" pageNumber="38">P.K.L. Ng</collectorName>
, 04.2012.
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B61FFADE7E8F8FDFC5DF812" blockId="37.[151,1437,1622,2040]" pageId="37" pageNumber="38">
<materialsCitation id="12E7B9264B61FFADE7E8F8FDFC59F812" ID-GBIF-Occurrence="4104078476" country="Vietnam" location="Ho Chi Minh City" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Ho Chi Minh City">
<emphasis id="90FB6F694B61FFADE7E8F8FDFEE9F8F6" box="[199,290,1872,1897]" italics="true" pageId="37" pageNumber="38">
<collectingCountry id="DA98F3EB4B61FFADE7E8F8FDFEE9F8F6" box="[199,290,1872,1897]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
</emphasis>
:
<specimenCount id="B48978F24B61FFADE61BF8E2FE49F8F6" box="[308,386,1871,1897]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(cl
<quantity id="65771E9E4B61FFADE69AF8E2FDEBF8F6" box="[437,544,1871,1897]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="37" pageNumber="38" unit="mm" value="17.1">17.1 mm</quantity>
), MNHN-IU-2013-14789, near
<collectingRegion id="604B7D994B61FFADE4A0F8E2FBAEF8F6" box="[911,1125,1871,1897]" country="Vietnam" name="Ho Chi Minh City" pageId="37" pageNumber="38">Ho Chi Minh City</collectingRegion>
, mangrove, water brackish (“very slightly salty”), in burrow in mud, leg. N. Ngoc-Ho, 05.1993
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B61FFADE7E8F83BFAAFF82F" blockId="37.[151,1437,1622,2040]" box="[199,1380,1942,1968]" pageId="37" pageNumber="38">
<materialsCitation id="12E7B9264B61FFADE7E8F83BFC8CF82F" ID-GBIF-Occurrence="4104078475" box="[199,839,1942,1968]" collectionCode="ZRC" country="Malaysia" location="Malaysia" pageId="37" pageNumber="38" specimenCode="ZRC 2014.0670" specimenCount="2" specimenCount-female="2">
<emphasis id="90FB6F694B61FFADE7E8F83BFEE4F82F" box="[199,303,1942,1968]" italics="true" pageId="37" pageNumber="38">
<collectingCountry id="DA98F3EB4B61FFADE7E8F83BFEE4F82F" box="[199,303,1942,1968]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
</emphasis>
:
<specimenCount id="B48978F24B61FFADE611F83BFE61F82F" box="[318,426,1942,1968]" pageId="37" pageNumber="38" type="female">2 females</specimenCount>
(cl 14.3,
<quantity id="65771E9E4B61FFADE53AF83BFDB6F82F" box="[533,637,1942,1968]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="37" pageNumber="38" unit="mm" value="14.7">14.7 mm</quantity>
),
<specimenCode id="F2291B004B61FFADE5BFF83BFC8CF82F" box="[656,839,1942,1968]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="37" pageNumber="38">ZRC 2014.0670</specimenCode>
</materialsCitation>
,
<collectingCountry id="DA98F3EB4B61FFADE47DF83BFC7EF82F" box="[850,949,1942,1968]" name="Malaysia" pageId="37" pageNumber="38">Sarawak</collectingCountry>
, Kuching prawn farm, no further data.
</paragraph>
<paragraph id="A230B37B4B61FFADE7E8F814FCA1F868" blockId="37.[151,1437,1622,2040]" pageId="37" pageNumber="38">
<emphasis id="90FB6F694B61FFADE7E8F814FE14F84C" bold="true" box="[199,479,1977,2003]" pageId="37" pageNumber="38">Tentative identification</emphasis>
.
<taxonomicName id="658FC8F84B61FFADE6DFF814FCE9F84C" authorityName="sensu De Man" authorityYear="1897" box="[496,802,1977,2003]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B61FFADE6DFF814FD81F84C" box="[496,586,1977,2003]" italics="true" pageId="37" pageNumber="38">Alpheus</emphasis>
<materialsCitation id="12E7B9264B61FFADE57AF814FCE9F84C" ID-GBIF-Occurrence="4104078425" box="[597,802,1977,2003]" collectingDate="1923-01-07" collectionCode="USNM" collectorName="H. Smith" country="Thailand" location="Bang Pakong River" pageId="37" pageNumber="38" specimenCode="USNM 65477" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
cf.
<emphasis id="90FB6F694B61FFADE553F814FCE9F84C" box="[636,802,1977,2003]" italics="true" pageId="37" pageNumber="38">microrhynchus</emphasis>
</materialsCitation>
</taxonomicName>
.
<collectingCountry id="DA98F3EB4B61FFADE41CF814FC50F84C" box="[819,923,1977,2003]" name="Thailand" pageId="37" pageNumber="38">Thailand</collectingCountry>
:
<specimenCount id="B48978F24B61FFADE486F814FC3CF84C" box="[937,1015,1977,2003]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
(cl
<quantity id="65771E9E4B61FFADE306F814FB4DF84C" box="[1065,1158,1977,2003]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="37" pageNumber="38" unit="mm" value="7.4">7.4 mm</quantity>
),
<specimenCount id="B48978F24B61FFADE3B2F814FACAF84C" box="[1181,1281,1977,2003]" pageId="37" pageNumber="38" type="female">1 female</specimenCount>
(cl
<quantity id="65771E9E4B61FFADE21CF814FA5BF84C" box="[1331,1424,1977,2003]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="37" pageNumber="38" unit="mm" value="8.7">8.7 mm</quantity>
),
<specimenCode id="F2291B004B61FFADE7B8F870FEF0F867" box="[151,315,2013,2040]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="38" type="Museum">USNM 65477</specimenCode>
/1,
<location id="A750E5A04B61FFADE676F873FDFDF868" LSID="urn:lsid:plazi:treatment:2A26026D4B61FFA2E7B8FB7CFAA3F86C:A750E5A04B61FFADE676F873FDFDF868" box="[345,566,2013,2040]" country="Thailand" name="Bang Pakong River" pageId="37" pageNumber="38">Bang Pakong River</location>
, leg.
<collectorName id="0F7AD6AD4B61FFADE55EF873FD12F868" box="[625,729,2013,2040]" pageId="37" pageNumber="38">H. Smith</collectorName>
,
<date id="D63195BB4B61FFADE5CBF870FCAEF868" box="[740,869,2013,2039]" pageId="37" pageNumber="38" value="1923-01-07">
<collectingDate id="C6756C534B61FFADE5CBF870FCAEF868" box="[740,869,2013,2039]" pageId="37" pageNumber="38" value="1923-01-07">01.07.1923</collectingDate>
</date>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B62FFA1E7E8FF3BFB7AFD56" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="description">
<paragraph id="A230B37B4B62FFAEE7E8FF3BFCE7FE16" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">
<emphasis id="90FB6F694B62FFAEE7E8FF3BFEA6FF2F" bold="true" box="[199,365,150,176]" pageId="38" pageNumber="39">Redescription</emphasis>
. See
<figureCitation id="3AB4AFFE4B62FFAEE69BFF3AFDDFFF2E" box="[436,532,151,177]" captionStart="FIGURE 19" captionStartId="39.[151,250,1944,1968]" captionTargetBox="[159,1427,181,1920]" captionTargetId="figure-15@39.[159,1427,181,1920]" captionTargetPageId="39" captionText="FIGURE 19. Alpheus microrhynchus De Man, 1897: neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped, lateral view; D—same, mesial view; E—minor (left) cheliped, lateral view; F—same, mesial view. Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912352" httpUri="https://zenodo.org/record/7912352/files/figure.png" pageId="38" pageNumber="39">Figs. 19</figureCitation>
,
<figureCitation id="3AB4AFFE4B62FFAEE50BFF3AFD8BFF2E" box="[548,576,151,177]" captionStart="FIGURE 20" captionStartId="40.[151,249,1647,1671]" captionTargetBox="[164,1423,199,1623]" captionTargetId="figure-21@40.[164,1423,199,1623]" captionTargetPageId="40" captionText="FIGURE 20. Alpheus microrhynchus De Man, 1897: male (cl 17.1 mm) from near Ho Chi Minh City, Vietnam (MNHN-IU- 2013-14789) [AD]; neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193) [E]; A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped in situ, mesial view; D—minor (left) cheliped in situ, lateral view; E—minor (left) cheliped, lateral view. Refer to A for scale in BD, and to Fig. 19A for scale in E." figureDoi="http://doi.org/10.5281/zenodo.7912354" httpUri="https://zenodo.org/record/7912354/files/figure.png" pageId="38" pageNumber="39">20</figureCitation>
and
<bibRefCitation id="C61ECE8A4B62FFAEE550FF3AFC55FF2E" author="Banner, A. H. &amp; Banner, D. M." box="[639,926,151,177]" pageId="38" pageNumber="39" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 50). Medium-sized species of
<taxonomicName id="658FC8F84B62FFAEE26DFF3AFA57FF2E" authorityName="Fabricius" authorityYear="1798" box="[1346,1436,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B62FFAEE26DFF3AFA57FF2E" box="[1346,1436,151,177]" italics="true" pageId="38" pageNumber="39">Alpheus</emphasis>
</taxonomicName>
(maximal cl
<quantity id="65771E9E4B62FFAEE606FF16FE5AFF4A" box="[297,401,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="38" pageNumber="39" unit="mm" value="16.8">16.8 mm</quantity>
, tl ~45.0 mm). Carapace glabrous, minutely and sparsely pitted, without pubescence, with weak grooves. Rostrum very short, subtriangular, broader than long, narrowing towards apex, distally subacute, not reaching 0.3 length of first article of antennular peduncle, obliquely ascendant or pointing straight-forward (see remarks below); rostral carina present as faint broad elevation flattening posterior to orbital hoods; rostro-orbital furrows very shallow; anterior margin of orbital hoods very shallowly concave, almost perpendicular to rostrum. Pterygostomial angle broadly rounded; cardiac notch deep.
</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8FE3EFC4FFDFE" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">Pleon smooth, finely pitted, without pubescence. Telson broad, ovate-rectangular, gently tapering towards posterior margin, about 1.3 times as long as maximal width near proximal margin; lateral margins with weak convexity at about mid-length; dorsal surface finely pitted, without longitudinal median depression, with two pairs of stout spiniform setae inserted in deep pits at some distance from lateral margins, first pair at about telson mid-length, second pair at about 0.7 of telson length; posterior margin broadly rounded, with two pairs of minute spiniform setae at posterolateral angles, lateral shorter than mesial.</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8FDC6FCC1FCA7" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">Antennular peduncle with stylocerite slightly convex laterally, with acute tip, latter not overreaching distal margin of first article; ventromesial carina with broadly triangular tooth, latter longer than high and without acute point; second article about 1.81.9 times as long as wide. Antennal peduncle with distoventral margin of basicerite unarmed or with minute tooth; carpocerite reaching slightly beyond scaphocerite; scaphocerite with lateral margin nearly straight; blade broad, separated from broad strong distolateral tooth by deep cleft; anterior margin of blade rounded and reaching slightly beyond distolateral tooth.</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8FCEFFD66FC75" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">Third maxilliped with antepenultimate article not particularly broadened, rather slender; penultimate article almost 3.2 times as long as wide, with long setae on lateroventral and mesioventral margins, latter not surpassing ultimate article; ultimate article with dense transverse rows of short serrulate setae on mesial surface and some much longer setae on dorsal and lateral surfaces, especially near or on apex; coxal lateral plate bluntly protruding dorsally; exopod reaching end of antepenultimate article.</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8FC59FD51FA40" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">
Major cheliped of
<taxonomicName id="658FC8F84B62FFAEE6BCFC58FDD5FB91" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[403,542,1012,1039]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="90FB6F694B62FFAEE6BCFC58FDD5FB91" box="[403,542,1012,1039]" italics="true" pageId="38" pageNumber="39">A. edwardsii</emphasis>
</taxonomicName>
-
<typeStatus id="7D340DD94B62FFAEE509FC58FD90FB90" box="[550,603,1013,1039]" pageId="38" pageNumber="39">type</typeStatus>
. Merus very stout, about 1.8 as long as wide; ventromesial margin unarmed distally. Carpus very short, cup-shaped. Chela large, massive, with fingers about 0.60.7 length of palm. Mesial face of palm with feebly developed mesial transverse ridge in proximal half, extending from proximal edge of subtriangular mesial longitudinal groove to ventral surface of palm; surface smooth, without granules. Lateral face of palm smooth, without granulation; lateral longitudinal groove moderately deep, subrectangular; proximal half of palm without transverse ridge. Dorsal shoulder rounded, sloping gently with angle inferior to 45° into broad transverse groove, not overhanging. Ventral shoulder pronounced, slightly protruding in lateral view, rounded. Pollex with mesial face smooth over most of its surface, with prominent mesial subdistal ridge marked by low protuberance proximally and extending to subdistal angle of pollex; area ventral to mesial subdistal ridge noticeably depressed; distomesial angle distinctly superior to 90°, blunt; lateral face smooth; distolateral angle about 90°, rounded. Dactylus slightly longer than pollex, with moderately developed dorsal ridge, latter twisted mesially, rounded, not protruding; both mesial and lateral surfaces smooth; dactylar plunger moderately large, stout, clearly distinct from ventral dactylar margin; distal surface of plunger with obliquely flattened ventromesial area, without deep transverse groove. Adhesive discs small.
</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8FA45FB49F8D9" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">
<collectingRegion id="604B7D994B62FFAEE7E8FA45FECAF99D" box="[199,257,1512,1538]" country="Maldives" name="Maale" pageId="38" pageNumber="39">Male</collectingRegion>
minor cheliped with chela strongly balaeniceps.Merus somewhat slenderer than that of major cheliped,about 2.5 times as long as wide; ventral margin with distomesial angle unarmed. Chela moderately swollen, with fingers about 0.9 length of palm; all surfaces smooth, without granulation. Palm subcylindrical, slightly swollen; mesial face with subdistal swelling; mesial longitudinal and lateral longitudinal groove indistinct or poorly demarcated; dorsal shoulder absent; dorsal transverse groove noticeable as shallow sinus; ventral transverse groove reduced to shallow sinus; ventral shoulder poorly marked as low bump or indistinct. Pollex with row of balaeniceps setae on low crest extending from base to just before mid-length of pollex; lateral surface with similar row of balaeniceps setae as on mesial surface. Dactylus with moderate lateral expansion (balaeniceps expansion) in proximal half, about twice as long as maximal width; lateral and mesial dactylar ridges each with dense rows of balaeniceps setae. Cutting edges of both fingers shallowly excavated on each side of sharp, blade-like ridge.
</paragraph>
<paragraph id="A230B37B4B62FFAEE7E8F8FDFD8CF849" blockId="38.[151,1437,150,2006]" pageId="38" pageNumber="39">Female minor cheliped simple, not balaeniceps. Chela slender, not particularly swollen, with fingers of about same length as palm. Palm surface smooth, without granulation, without noticeable sculpture, i.e., without mesial and lateral longitudinal grooves, and without shoulders. Fingers simple, with sharp cutting edges, not gaping when closed; fingertips curved and crossing.</paragraph>
<caption id="F6F0E3F34B63FFAFE7B8F835FB04F867" ID-DOI="http://doi.org/10.5281/zenodo.7912352" ID-Zenodo-Dep="7912352" httpUri="https://zenodo.org/record/7912352/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1944,1968]" targetBox="[159,1427,181,1920]" targetPageId="39" targetType="figure">
<paragraph id="A230B37B4B63FFAFE7B8F835FB04F867" blockId="39.[151,1436,1944,2040]" pageId="39" pageNumber="40">
<emphasis id="90FB6F694B63FFAFE7B8F835FED2F82F" bold="true" box="[151,281,1944,1968]" pageId="39" pageNumber="40">FIGURE 19</emphasis>
.
<taxonomicName id="658FC8F84B63FFAFE609F835FD7AF82F" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[294,689,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B63FFAFE609F835FDDDF82F" box="[294,534,1944,1968]" italics="true" pageId="39" pageNumber="40">Alpheus microrhynchus</emphasis>
<bibRefCitation id="C61ECE8A4B63FFAFE533F835FD7AF82F" author="De Man, J. G." box="[540,689,1944,1968]" pageId="39" pageNumber="40" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped, lateral view; D—same, mesial view; E—minor (left) cheliped, lateral view; F—same, mesial view. Refer to A for scale.
</paragraph>
</caption>
<caption id="F6F0E3F34B6CFFA0E7B8F9C2FAFCF96D" ID-DOI="http://doi.org/10.5281/zenodo.7912354" ID-Zenodo-Dep="7912354" httpUri="https://zenodo.org/record/7912354/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,249,1647,1671]" targetBox="[164,1423,199,1623]" targetPageId="40" targetType="figure">
<paragraph id="A230B37B4B6CFFA0E7B8F9C2FAFCF96D" blockId="40.[151,1437,1647,1779]" pageId="40" pageNumber="41">
<emphasis id="90FB6F694B6CFFA0E7B8F9C2FED1F918" bold="true" box="[151,282,1647,1671]" pageId="40" pageNumber="41">FIGURE 20</emphasis>
.
<taxonomicName id="658FC8F84B6CFFA0E607F9C2FD7CF918" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[296,695,1647,1671]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6CFFA0E607F9C2FDDDF918" box="[296,534,1647,1671]" italics="true" pageId="40" pageNumber="41">Alpheus microrhynchus</emphasis>
<bibRefCitation id="C61ECE8A4B6CFFA0E530F9C2FD7CF918" author="De Man, J. G." box="[543,695,1647,1671]" pageId="40" pageNumber="41" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl 17.1 mm) from near Ho Chi Minh City, Vietnam (MNHN-IU- 2013-14789) [AD]; neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193) [E]; A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped
<emphasis id="90FB6F694B6CFFA0E4F6F915FBDDF950" box="[985,1046,1720,1743]" italics="true" pageId="40" pageNumber="41">in situ</emphasis>
, mesial view; D—minor (left) cheliped
<emphasis id="90FB6F694B6CFFA0E7B8F971FF1EF96C" box="[151,213,1756,1779]" italics="true" pageId="40" pageNumber="41">in situ</emphasis>
, lateral view; E—minor (left) cheliped, lateral view. Refer to A for scale in BD, and to Fig. 19A for scale in E.
</paragraph>
</caption>
<paragraph id="A230B37B4B6CFFA1E7E8F88CFECCFF66" blockId="40.[151,1436,1825,2032]" lastBlockId="41.[151,1437,151,2036]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">Second pereiopod with ratio of carpal subarticles ranging from 4: 1.7: 1: 1: 1.5 to 5: 1.8: 1: 1: 1.6. Third pereiopod moderately robust; ischium armed with small spiniform seta on lateroventral surface; merus about four times as long as maximal width, unarmed; propodus with about five spiniform setae on ventral margin and one pair of spiniform setae on distoventral margin adjacent to dactylus; dactylus close to 0.3 of propodus length, trigonal-conical, tapering, not broader than propodus in dorsal view. Fourth pereiopod generally similar to third pereiopod, slenderer; merus about 4.8 times as long as maximal width; dactylus about 0.3 length of propodus. Fifth pereiopod much slenderer than third and fourth pereiopods; ischium unarmed; propodus with at least two slender spiniform setae (or robust setae) and well-developed setal brush; dactylus trigonal-subspatulate, about 0.3 length of propodus.</paragraph>
<paragraph id="A230B37B4B6DFFA1E7E8FEAFFF3BFE17" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<collectingRegion id="604B7D994B6DFFA1E7E8FEAFFECAFE83" box="[199,257,258,284]" country="Maldives" name="Maale" pageId="41" pageNumber="42">Male</collectingRegion>
second pleopod with appendix masculina ~1.2 length of appendix interna. Uropod with each protopodal lobe ending in sharp tooth distally; exopod and endopod broad, ovate; diaeresis of exopod complete, almost straight, with two subacute teeth fringing small distolateral spiniform seta (occasionally two spiniform setae); lateral margin convex.
</paragraph>
<paragraph id="A230B37B4B6DFFA1E7E8FE3FFCC0FE4F" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
Eggs not numerous (
<quantity id="65771E9E4B6DFFA1E680FE3CFE22FE34" box="[431,489,401,427]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="41" pageNumber="42" unit="in" value="10.0">10 in</quantity>
one ovigerous female examined, ZRC 2012.0399, less than
<quantity id="65771E9E4B6DFFA1E3A0FE3CFB02FE34" box="[1167,1225,401,427]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="41" pageNumber="42" unit="in" value="40.0">40 in</quantity>
ovigerous females in
<figureCitation id="3AB4AFFE4B6DFFA1E79BFE18FEF6FE50" box="[180,317,437,463]" captionStart="FIGURE 21" captionStartId="43.[151,250,1724,1748]" captionTargetBox="[151,1435,196,1685]" captionTargetId="figure-15@43.[151,1435,196,1685]" captionTargetPageId="43" captionText="FIGURE 21. Alpheus microrhynchus De Man, 1897: ovigerous female (cl indet.) imported to Germany from Thailand [A]; ovigerous female (cl indet.) imported to Singapore from Thailand [BE]: A, B—habitus, lateral view; C—pleon and tail fan, dorsal view; D—pleon, ventrolateral view E—frontal region and chelipeds, dorsal view. Aquarium-kept specimens, probably not preserved / deposited. Note the low number and large size of developing embryos inA and D. Photographs courtesy of Chris Lukhaup (A) and Paul Ng (BE)." figureDoi="http://doi.org/10.5281/zenodo.7912358" httpUri="https://zenodo.org/record/7912358/files/figure.png" pageId="41" pageNumber="42">Fig. 21A, D</figureCitation>
), very large; egg diameter about
<quantity id="65771E9E4B6DFFA1E581FE18FCCCFE4F" box="[686,775,437,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<paragraph id="A230B37B4B6DFFA1E7E8FE75FB7AFD56" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FE75FEB4FE6C" bold="true" box="[199,383,472,499]" pageId="41" pageNumber="42">Colour pattern</emphasis>
. Carapace pale olive-brown, sometimes with purplish or bluish tinge; posterior margin of carapace and each pleonite with one narrow, transverse, dark grey-blue or dusky band; antennular peduncles and scaphocerite pale brown-green or greenish blue; antennular and antennal flagella bluish or greenish; mesial face of major chela olive-green to bluish green or bluish grey, with some olive-beige, pale orange or yellowish areas; fingers pinkish distally; minor chela dark olive-grey to brown or bluish green; walking legs reddish; telson greyish blue distally; uropods mostly greyish blue with some reddish tinge; exopod with conspicuous brick-red patch above distolateral spiniform seta; area posterior to transverse suture intense deep blue (
<figureCitation id="3AB4AFFE4B6DFFA1E334FD02FBB2FD56" box="[1051,1145,687,713]" captionStart="FIGURE 21" captionStartId="43.[151,250,1724,1748]" captionTargetBox="[151,1435,196,1685]" captionTargetId="figure-15@43.[151,1435,196,1685]" captionTargetPageId="43" captionText="FIGURE 21. Alpheus microrhynchus De Man, 1897: ovigerous female (cl indet.) imported to Germany from Thailand [A]; ovigerous female (cl indet.) imported to Singapore from Thailand [BE]: A, B—habitus, lateral view; C—pleon and tail fan, dorsal view; D—pleon, ventrolateral view E—frontal region and chelipeds, dorsal view. Aquarium-kept specimens, probably not preserved / deposited. Note the low number and large size of developing embryos inA and D. Photographs courtesy of Chris Lukhaup (A) and Paul Ng (BE)." figureDoi="http://doi.org/10.5281/zenodo.7912358" httpUri="https://zenodo.org/record/7912358/files/figure.png" pageId="41" pageNumber="42">Figs. 21</figureCitation>
,
<figureCitation id="3AB4AFFE4B6DFFA1E3AAFD02FB6EFD56" box="[1157,1189,687,713]" captionStart="FIGURE 22" captionStartId="44.[151,250,1907,1931]" captionTargetBox="[226,1361,193,1884]" captionTargetId="figure-21@44.[226,1361,181,1884]" captionTargetPageId="44" captionText="FIGURE 22. Alpheus microrhynchus De Man, 1897: female (cl indet.) imported to Singapore from Thailand; A, B—habitus, dorsal view (with one Singapore dollar coin, diameter 22.4 mm, for scale in B); C—detail of frontal margin; D—major (right) chela, mesial view; E—posterior pleonites and tail fan, d orsal view. Aquarium-kept specimen, probably not preserved / deposited. Photographs courtesy of Paul Ng." figureDoi="http://doi.org/10.5281/zenodo.7912362" httpUri="https://zenodo.org/record/7912362/files/figure.png" pageId="41" pageNumber="42">22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B6DFFA1E7E8FD7EFD97FC8E" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="A230B37B4B6DFFA1E7E8FD7EFD97FC8E" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FD7EFEABFD72" bold="true" box="[199,352,723,749]" pageId="41" pageNumber="42">Type locality</emphasis>
.
<materialsCitation id="12E7B9264B6DFFA1E641FD7EFCB7FD72" ID-GBIF-Occurrence="4104078407" box="[366,892,723,749]" country="Indonesia" location="Kabupanten Sambas" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="neotype">
Pontianak,
<collectingRegion id="604B7D994B6DFFA1E6C3FD7EFDA4FD72" box="[492,623,723,749]" country="Indonesia" name="Kalimantan Utara" pageId="41" pageNumber="42">Kalimantan</collectingRegion>
(west coast of Borneo).
</materialsCitation>
<materialsCitation id="12E7B9264B6DFFA1E4ACFD7EFD97FC8E" ID-GBIF-Occurrence="4104078554" country="Indonesia" location="Kabupanten Sambas" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="neotype">
<typeStatus id="7D340DD94B6DFFA1E4ACFD7EFC29FD72" box="[899,994,723,749]" pageId="41" pageNumber="42" type="neotype">Neotype</typeStatus>
from
<location id="A750E5A04B6DFFA1E309FD7EFAC5FD72" LSID="urn:lsid:plazi:treatment:2A26026D4B61FFA2E7B8FB7CFAA3F86C:A750E5A04B6DFFA1E309FD7EFAC5FD72" box="[1062,1294,723,749]" country="Indonesia" name="Kabupanten Sambas" pageId="41" pageNumber="42" stateProvince="Kalimantan Utara">Kabupanten Sambas</location>
,
<collectingRegion id="604B7D994B6DFFA1E235FD7EFF1FFC8E" country="Indonesia" name="Kalimantan Barat" pageId="41" pageNumber="42">Kalimantan Barat</collectingRegion>
(west coast of Borneo),
<collectingCountry id="DA98F3EB4B6DFFA1E6C8FD5AFD93FC8E" box="[487,600,759,785]" name="Indonesia" pageId="41" pageNumber="42">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B6DFFA1E7E8FCB7FE05FC00" pageId="41" pageNumber="42" type="distribution">
<paragraph id="A230B37B4B6DFFA1E7E8FCB7FE05FC00" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FCB7FE92FCAB" bold="true" box="[199,345,794,820]" pageId="41" pageNumber="42">Distribution</emphasis>
. South-East Asia (
<figureCitation id="3AB4AFFE4B6DFFA1E51AFCB7FD69FCAA" box="[565,674,794,821]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="41" pageNumber="42">Fig. 51D</figureCitation>
):
<collectingCountry id="DA98F3EB4B6DFFA1E596FCB7FCD6FCAB" box="[697,797,794,820]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
(vicinity of
<collectingRegion id="604B7D994B6DFFA1E49DFCB7FBDCFCAB" box="[946,1047,794,820]" country="Thailand" name="Krung Thep Mahanakhon" pageId="41" pageNumber="42">Bangkok</collectingRegion>
and possibly
<collectingRegion id="604B7D994B6DFFA1E392FCB7FAECFCAB" box="[1213,1319,794,820]" country="Thailand" name="Songkhla" pageId="41" pageNumber="42">Songkhla</collectingRegion>
Lake and Bang Pakong), Indonesian and Malaysian parts of Borneo (Kalimantan Barat,
<collectingCountry id="DA98F3EB4B6DFFA1E304FC93FB46FCC7" box="[1067,1165,830,856]" name="Malaysia" pageId="41" pageNumber="42">Sarawak</collectingCountry>
) (
<bibRefCitation id="C61ECE8A4B6DFFA1E388FC93FA85FCC7" author="De Man, J. G." box="[1191,1358,830,857]" pageId="41" pageNumber="42" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897</bibRefCitation>
,
<bibRefCitation id="C61ECE8A4B6DFFA1E273FC93FA5CFCC7" author="De Man, J. G." box="[1372,1431,830,856]" pageId="41" pageNumber="42" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">1898</bibRefCitation>
;
<bibRefCitation id="C61ECE8A4B6DFFA1E7B8FCCFFEE9FCE3" author="Nobili, G." box="[151,290,866,892]" pageId="41" pageNumber="42" pagination="473 - 523" refId="ref73484" refString="Nobili, G. (1900) Decapodi e stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale di Genova, 40, 473 - 523." type="journal article" year="1900">Nobili 1900</bibRefCitation>
; present study); records from eastern
<collectingRegion id="604B7D994B6DFFA1E5FFFCCFFCCFFCE3" box="[720,772,866,892]" country="Indonesia" name="Jawa Timur" pageId="41" pageNumber="42">Java</collectingRegion>
,
<collectingCountry id="DA98F3EB4B6DFFA1E43EFCCFFCB5FCE3" box="[785,894,866,892]" name="Indonesia" pageId="41" pageNumber="42">Indonesia</collectingCountry>
(
<bibRefCitation id="C61ECE8A4B6DFFA1E4A0FCCFFBE4FCE3" author="De Man, J. G." box="[911,1071,866,892]" pageId="41" pageNumber="42" pagination="133 - 465" refId="ref70992" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga-Expeditie, 39 a 1 (2), 133 - 465; (1915) Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article" year="1911">De Man 1911</bibRefCitation>
), and
<collectingCountry id="DA98F3EB4B6DFFA1E359FCCFFB23FCE3" box="[1142,1256,866,892]" name="Singapore" pageId="41" pageNumber="42">Singapore</collectingCountry>
(
<bibRefCitation id="C61ECE8A4B6DFFA1E3D8FCCFFA53FCE2" author="Johnson, D. S." box="[1271,1432,866,893]" pageId="41" pageNumber="42" pagination="7 - 11" refId="ref72065" refString="Johnson, D. S. (1965) A review of brackish water prawns of Malaya. Bulletin of the National Museum of Singapore, 33, 7 - 11." type="journal article" year="1965">Johnson 1965</bibRefCitation>
,
<bibRefCitation id="C61ECE8A4B6DFFA1E7B8FC28FF1BFC00" author="Johnson, D. S." box="[151,208,901,927]" pageId="41" pageNumber="42" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">1976</bibRefCitation>
) require confirmation.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B6DFFA1E7E8FC04FCC5FC5C" box="[199,782,937,964]" pageId="41" pageNumber="42" type="vernacular_names">
<paragraph id="A230B37B4B6DFFA1E7E8FC04FCC5FC5C" blockId="41.[151,1437,151,2036]" box="[199,782,937,964]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FC04FE38FC5C" bold="true" box="[199,499,937,963]" pageId="41" pageNumber="42">Common name proposed</emphasis>
. Klang snapping shrimp.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B6DFFA1E7E8FC60FAEAFB7E" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="A230B37B4B6DFFA1E7E8FC60FCA7FBE9" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FC60FE76FC78" bold="true" box="[199,445,973,999]" pageId="41" pageNumber="42">Ecology and biology</emphasis>
.
<taxonomicName id="658FC8F84B6DFFA1E6E2FC60FD1CFC78" authorityName="sensu De Man" authorityYear="1897" box="[461,727,973,999]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E6E2FC60FD1CFC78" box="[461,727,973,999]" italics="true" pageId="41" pageNumber="42">Alpheus microrhynchus</emphasis>
</taxonomicName>
can be qualified as an stenohaline snapping shrimp, confined to oligohaline habitats, such as rivers, creeks, lagoons and artificial canals (man-made for navigation or irrigation of rice fields) under slight tidal influence, living in deep burrows dug into muddy bottom or riverbanks. The small number of eggs and their large size suggest a highly abbreviated larval development in this species (H. Coutière in
<bibRefCitation id="C61ECE8A4B6DFFA1E7B8FBF0FEFEFBE9" author="De Man, J. G." box="[151,309,1116,1143]" pageId="41" pageNumber="42" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man 1898</bibRefCitation>
), although it remains to be studied in more details.
</paragraph>
<paragraph id="A230B37B4B6DFFA1E7E8FB2DFAEAFB7E" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<taxonomicName id="658FC8F84B6DFFA1E7E8FB2DFE05FB05" authorityName="sensu De Man" authorityYear="1897" box="[199,462,1152,1178]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E7E8FB2DFE05FB05" box="[199,462,1152,1178]" italics="true" pageId="41" pageNumber="42">Alpheus microrhynchus</emphasis>
</taxonomicName>
is occasionally sold on fish or pet markets, mainly in
<collectingCountry id="DA98F3EB4B6DFFA1E308FB2DFB40FB05" box="[1063,1163,1152,1178]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
(e.g., Chatuchak market in
<collectingRegion id="604B7D994B6DFFA1E799FB0EFEEBFB22" box="[182,288,1187,1213]" country="Thailand" name="Krung Thep Mahanakhon" pageId="41" pageNumber="42">Bangkok</collectingRegion>
). According to observations of F. Fasquel (pers. comm.), the species needs a low percentage of salt added to freshwater since all shrimps kept in pure freshwater unexpectedly died after only a few weeks.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B6DFFA2E7E8FB46FAA3F86C" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="discussion">
<paragraph id="A230B37B4B6DFFA1E7E8FB46FD38F966" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
<emphasis id="90FB6F694B6DFFA1E7E8FB46FE7DFA9A" bold="true" box="[199,438,1259,1285]" pageId="41" pageNumber="42">Taxonomic remarks</emphasis>
. The
<typeStatus id="7D340DD94B6DFFA1E6DAFB46FD85FA9A" box="[501,590,1259,1285]" pageId="41" pageNumber="42" type="neotype">neotype</typeStatus>
of
<taxonomicName id="658FC8F84B6DFFA1E55CFB46FCF3FA9A" authorityName="sensu De Man" authorityYear="1897" box="[627,824,1259,1285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E55CFB46FCF3FA9A" box="[627,824,1259,1285]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
from western Borneo corresponds to
<bibRefCitation id="C61ECE8A4B6DFFA1E3F2FB46FA57FA9A" author="De Man, J. G." box="[1245,1436,1259,1285]" pageId="41" pageNumber="42" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Mans (1897)</bibRefCitation>
description in all but a few minor details. The first small discrepancy is the direction of the rostrum, which was described as pointing obliquely upwards (= “schräg nach oben gerichtetes Spitzchen”). In most examined specimens of
<taxonomicName id="658FC8F84B6DFFA1E798FAFBFEB5FAEF" authorityName="sensu De Man" authorityYear="1897" box="[183,382,1366,1392]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E798FAFBFEB5FAEF" box="[183,382,1366,1392]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
, including the
<typeStatus id="7D340DD94B6DFFA1E504FAFAFD4CFAEE" box="[555,647,1367,1393]" pageId="41" pageNumber="42" type="neotype">neotype</typeStatus>
, the rostrum is directed straight forwards (
<figureCitation id="3AB4AFFE4B6DFFA1E343FAFBFB1DFAEF" box="[1132,1238,1366,1392]" captionStart="FIGURE 20" captionStartId="40.[151,249,1647,1671]" captionTargetBox="[164,1423,199,1623]" captionTargetId="figure-21@40.[164,1423,199,1623]" captionTargetPageId="40" captionText="FIGURE 20. Alpheus microrhynchus De Man, 1897: male (cl 17.1 mm) from near Ho Chi Minh City, Vietnam (MNHN-IU- 2013-14789) [AD]; neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193) [E]; A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped in situ, mesial view; D—minor (left) cheliped in situ, lateral view; E—minor (left) cheliped, lateral view. Refer to A for scale in BD, and to Fig. 19A for scale in E." figureDoi="http://doi.org/10.5281/zenodo.7912354" httpUri="https://zenodo.org/record/7912354/files/figure.png" pageId="41" pageNumber="42">Fig. 20A</figureCitation>
), although in
<specimenCount id="B48978F24B6DFFA1E25BFAFAFF06FA0B" pageId="41" pageNumber="42" type="male">one male</specimenCount>
specimen from
<collectingCountry id="DA98F3EB4B6DFFA1E6A5FAD7FE25FA0B" box="[394,494,1402,1428]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
(ZRC 2012.0399), it is directed slightly obliquely upwards. In addition, a clearly obliquely upwards-pointing rostrum can be seen in the illustrated (but not preserved) specimen from
<collectingCountry id="DA98F3EB4B6DFFA1E216FA30FA56FA28" box="[1337,1437,1437,1463]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
(
<figureCitation id="3AB4AFFE4B6DFFA1E7B0FA6CFEC2FA44" box="[159,265,1473,1499]" captionStart="FIGURE 21" captionStartId="43.[151,250,1724,1748]" captionTargetBox="[151,1435,196,1685]" captionTargetId="figure-15@43.[151,1435,196,1685]" captionTargetPageId="43" captionText="FIGURE 21. Alpheus microrhynchus De Man, 1897: ovigerous female (cl indet.) imported to Germany from Thailand [A]; ovigerous female (cl indet.) imported to Singapore from Thailand [BE]: A, B—habitus, lateral view; C—pleon and tail fan, dorsal view; D—pleon, ventrolateral view E—frontal region and chelipeds, dorsal view. Aquarium-kept specimens, probably not preserved / deposited. Note the low number and large size of developing embryos inA and D. Photographs courtesy of Chris Lukhaup (A) and Paul Ng (BE)." figureDoi="http://doi.org/10.5281/zenodo.7912358" httpUri="https://zenodo.org/record/7912358/files/figure.png" pageId="41" pageNumber="42">Fig. 21A</figureCitation>
). Some variation in the direction of the rostrum is also present in the closely related
<taxonomicName id="658FC8F84B6DFFA1E3FFFA6FFAACFA44" box="[1232,1383,1473,1500]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cyanoteles">
<emphasis id="90FB6F694B6DFFA1E3FFFA6FFAACFA44" box="[1232,1383,1473,1500]" italics="true" pageId="41" pageNumber="42">A. cyanoteles</emphasis>
</taxonomicName>
(see below). The second difference is the armature of the antennal basicerite, which according to
<bibRefCitation id="C61ECE8A4B6DFFA1E391FA4BFAA5FA60" author="De Man, J. G." box="[1214,1390,1509,1536]" pageId="41" pageNumber="42" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897)</bibRefCitation>
has a rudimentary, hardly 1.0 mm long tooth (= “rudimentäres, kaum mm langes Stachelchen”). In the
<typeStatus id="7D340DD94B6DFFA1E20DF9A4FAB0F9BC" box="[1314,1403,1545,1571]" pageId="41" pageNumber="42" type="neotype">neotype</typeStatus>
of
<taxonomicName id="658FC8F84B6DFFA1E7B8F980FEABF9D9" authorityName="sensu De Man" authorityYear="1897" box="[151,352,1580,1607]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E7B8F980FEABF9D9" box="[151,352,1580,1607]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
from Kalimantan and all specimens from
<collectingCountry id="DA98F3EB4B6DFFA1E479F981FC77F9D9" box="[854,956,1580,1606]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
, the distoventral margin of the basicerite is smoothly rounded, i.e., has no tooth, exactly as in the material from
<collectingCountry id="DA98F3EB4B6DFFA1E4EBF9FDFBE3F9F5" box="[964,1064,1616,1642]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
reported by
<bibRefCitation id="C61ECE8A4B6DFFA1E394F9FCFAA0F9F5" author="De Man, J. G." box="[1211,1387,1616,1643]" pageId="41" pageNumber="42" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man (1898)</bibRefCitation>
and
<bibRefCitation id="C61ECE8A4B6DFFA1E7B8F9D8FE78F911" author="Banner, A. H. &amp; Banner, D. M." box="[151,435,1652,1679]" pageId="41" pageNumber="42" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
. Because of its small size, this tooth may break off or get blunter after each moult, to the point of eventually becoming obsolete in full-grown adults. However, the development of the tooth on the basicerite may be simply variable in
<taxonomicName id="658FC8F84B6DFFA1E6FFF911FD53F94A" authorityName="sensu De Man" authorityYear="1897" box="[464,664,1723,1750]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E6FFF911FD53F94A" box="[464,664,1723,1750]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
. Noteworthy here is that a similar variation in the armature of the basicerite was observed in
<taxonomicName id="658FC8F84B6DFFA1E6EAF94DFDAFF966" authorityName="sensu De Man" authorityYear="1897" box="[453,612,1759,1786]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6DFFA1E6EAF94DFDAFF966" box="[453,612,1759,1786]" italics="true" pageId="41" pageNumber="42">A. euphrosyne</emphasis>
</taxonomicName>
(see above).
</paragraph>
<paragraph id="A230B37B4B6DFFA2E7E8F8AEFED0FE51" blockId="41.[151,1437,151,2036]" lastBlockId="42.[151,1437,151,2036]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
Two further characters of
<taxonomicName id="658FC8F84B6DFFA1E6CEF8AEFD6EF882" authorityName="sensu De Man" authorityYear="1897" box="[481,677,1795,1821]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E6CEF8AEFD6EF882" box="[481,677,1795,1821]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
, viz. the proportions of the carpal subarticles of the second pereiopod (P2) and the shape of the dactyli of the third to fifth pereiopods need to be discussed in more detail. In the original description of
<taxonomicName id="658FC8F84B6DFFA1E619F8E6FE32F8FB" authorityName="sensu De Man" authorityYear="1897" box="[310,505,1866,1893]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6DFFA1E619F8E6FE32F8FB" box="[310,505,1866,1893]" italics="true" pageId="41" pageNumber="42">A. microrhynchus</emphasis>
</taxonomicName>
(material from Pontianak, as
<taxonomicName id="658FC8F84B6DFFA1E410F8E7FC0BF8FB" box="[831,960,1866,1892]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B6DFFA1E410F8E7FC52F8FB" box="[831,921,1866,1892]" italics="true" pageId="41" pageNumber="42">Alpheus</emphasis>
sp.
</taxonomicName>
),
<bibRefCitation id="C61ECE8A4B6DFFA1E4FBF8E6FBB6F8FB" author="De Man, J. G." box="[980,1149,1866,1893]" pageId="41" pageNumber="42" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897)</bibRefCitation>
described the proportions of the carpal subarticles of the second pereiopod as 3.5: 1.3: 0.6: 0.6: 0.96, which is roughly similar to his observations (
<bibRefCitation id="C61ECE8A4B6DFFA1E61CF83FFE05F833" author="De Man, J. G." box="[307,462,1938,1964]" pageId="41" pageNumber="42" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man 1898</bibRefCitation>
) of the material from
<collectingRegion id="604B7D994B6DFFA1E5EEF83FFCE2F833" box="[705,809,1938,1964]" country="Thailand" name="Krung Thep Mahanakhon" pageId="41" pageNumber="42">Bangkok</collectingRegion>
, 4.3: 1.6: 0.8: 0.72: 1.04 (right P2), and 4.0: 1.5: 0.8: 0.72: 1.1 (left P2). These numbers show that the ratio of the first to second subarticle is proportionally higher in the material from
<collectingCountry id="DA98F3EB4B6DFFA1E619F874FE57F86C" box="[310,412,2009,2035]" name="Thailand" pageId="41" pageNumber="42">Thailand</collectingCountry>
, 2.73.0, as illustrated by
<bibRefCitation id="C61ECE8A4B6DFFA1E5EFF877FC1BF86C" author="Banner, A. H. &amp; Banner, D. M." box="[704,976,2009,2036]" pageId="41" pageNumber="42" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 50F), compared to the material from Borneo, where this ratio is at most 2.4. Regarding the condition of the dactylus of the ambulatory pereiopods,
<bibRefCitation id="C61ECE8A4B6EFFA2E253FF3AFE9EFF4A" author="De Man, J. G." pageId="42" pageNumber="43" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897: 754)</bibRefCitation>
stated that the third to fifth pereiopods are somewhat similar to those of
<taxonomicName id="658FC8F84B6EFFA2E3ADFF16FAEAFF4B" authorityName="sensu De Man" authorityYear="1897" box="[1154,1313,186,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E3ADFF16FAEAFF4B" box="[1154,1313,186,213]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
(“gleichen ungefähr denen von
<emphasis id="90FB6F694B6EFFA2E6A8FF73FDF1FF67" box="[391,570,222,248]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="658FC8F84B6EFFA2E6A8FF73FDFBFF67" authorityName="sensu De Man" authorityYear="1897" box="[391,560,222,248]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
</emphasis>
). No specific details on the dactylus were provided in the complementary description, where
<bibRefCitation id="C61ECE8A4B6EFFA2E642FEAFFD99FE83" author="De Man, J. G." box="[365,594,258,284]" pageId="42" pageNumber="43" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man (1898: 321)</bibRefCitation>
only noted that the ambulatory pereiopods agree with those of the material from Pontianak. However,
<bibRefCitation id="C61ECE8A4B6EFFA2E6E1FE8BFD26FEA0" author="Banner, A. H. &amp; Banner, D. M." box="[462,749,293,320]" pageId="42" pageNumber="43" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
noted that the dactylus of
<taxonomicName id="658FC8F84B6EFFA2E308FE8BFB25FEA0" authorityName="sensu De Man" authorityYear="1897" box="[1063,1262,293,320]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E308FE8BFB25FEA0" box="[1063,1262,293,320]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
, at least in the material from
<collectingCountry id="DA98F3EB4B6EFFA2E611FEE4FE6FFEFC" box="[318,420,329,355]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
, is not spatulate, as in
<taxonomicName id="658FC8F84B6EFFA2E598FEE7FC91FEFC" authorityName="sensu De Man" authorityYear="1897" box="[695,858,329,356]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E598FEE7FC91FEFC" box="[695,858,329,356]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
, but rather “conical, tapering”.
<bibRefCitation id="C61ECE8A4B6EFFA2E3E1FEE7FA57FEFC" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[1230,1436,329,356]" pageId="42" pageNumber="43" pagination="37 - 63" refId="ref75202" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (1996) A new species of freshwater snapping shrimp, Alpheus cyanoteles (Decapoda: Caridea: Alpheidae) from Peninsular Malaysia and a redescription of Alpheus paludicola Kemp, 1915. Raffles Bulletin of Zoology, 44, 37 - 63." type="journal article" year="1996">Yeo &amp; Ng (1996)</bibRefCitation>
examined several specimens identified as
<taxonomicName id="658FC8F84B6EFFA2E55AFEC0FCF7FE18" authorityName="sensu De Man" authorityYear="1897" box="[629,828,365,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E55AFEC0FCF7FE18" box="[629,828,365,391]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
in the ZRC holdings and concluded that the dactylus in this species is “subconical to subspatulate”, which is essentially the same as trigonal-subspatulate in the above description.
</paragraph>
<paragraph id="A230B37B4B6EFFA2E7E8FE75FB8FFBE9" blockId="42.[151,1437,151,2036]" pageId="42" pageNumber="43">
<taxonomicName id="658FC8F84B6EFFA2E7E8FE75FE06FE6D" authorityName="sensu De Man" authorityYear="1897" box="[199,461,472,498]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E7E8FE75FE06FE6D" box="[199,461,472,498]" italics="true" pageId="42" pageNumber="43">Alpheus microrhynchus</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="658FC8F84B6EFFA2E5E4FE74FCA3FE6D" authorityName="sensu De Man" authorityYear="1897" box="[715,872,472,499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E5E4FE74FCA3FE6D" box="[715,872,472,499]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
,
<taxonomicName id="658FC8F84B6EFFA2E45AFE74FBE9FE6D" authorityName="De Man" authorityYear="1920" box="[885,1058,472,499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E45AFE74FBE9FE6D" box="[885,1058,472,499]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E37AFE74FB3EFE6D" authorityName="Yaldwyn" authorityYear="1971" box="[1109,1269,472,499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E37AFE74FB3EFE6D" box="[1109,1269,472,499]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
by the ischium of the third and fourth pereiopods unarmed (
<emphasis id="90FB6F694B6EFFA2E5B3FE50FD78FD89" box="[668,691,509,534]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
. usually armed with a spiniform seta in the other three species); the much larger size of eggs, with the diameter around
<quantity id="65771E9E4B6EFFA2E40CFD8DFCB6FDA5" box="[803,893,544,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="42" pageNumber="43" unit="mm" value="1.4">1.4 mm</quantity>
(
<emphasis id="90FB6F694B6EFFA2E4BFFD8DFC64FDA6" box="[912,943,544,569]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
equal to or less than
<quantity id="65771E9E4B6EFFA2E39DFD8DFAC0FDA5" box="[1202,1291,544,570]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="43" unit="mm" value="0.7">0.7 mm</quantity>
in the other species); the less expanded, trigonal-conical (at most weakly subspatulate) dactyli of the third to fifth pereiopods (
<emphasis id="90FB6F694B6EFFA2E78EFDC5FF73FD1E" box="[161,184,616,641]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
. spatulate, much more expanded, in the other species; see also below); the perfectly smooth mesial surface of the major chela (
<emphasis id="90FB6F694B6EFFA2E673FD21FEB8FD3A" box="[348,371,652,677]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
. with a small distal field of granules in
<taxonomicName id="658FC8F84B6EFFA2E468FD26FC20FD3A" authorityName="Yaldwyn" authorityYear="1971" box="[839,1003,651,677]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E468FD26FC20FD3A" box="[839,1003,651,677]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
and with large granulated areas in
<taxonomicName id="658FC8F84B6EFFA2E2ABFD26FEDCFD56" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E2ABFD26FEDCFD56" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E660FD02FE36FD56" authorityName="De Man" authorityYear="1920" box="[335,509,687,713]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E660FD02FE36FD56" box="[335,509,687,713]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
); and the absence of a median longitudinal depression on the dorsal surface of the telson (present in the other species, especially well demarcated in
<taxonomicName id="658FC8F84B6EFFA2E451FD7EFBD6FD73" authorityName="sensu De Man" authorityYear="1897" box="[894,1053,722,749]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E451FD7EFBD6FD73" box="[894,1053,722,749]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E37AFD7EFAC8FD73" authorityName="De Man" authorityYear="1920" box="[1109,1283,722,749]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E37AFD7EFAC8FD73" box="[1109,1283,722,749]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
). In addition,
<taxonomicName id="658FC8F84B6EFFA2E7B8FD5BFE97FC8F" authorityName="sensu De Man" authorityYear="1897" box="[151,348,758,784]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E7B8FD5BFE97FC8F" box="[151,348,758,784]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="658FC8F84B6EFFA2E6C1FD5BFD57FC8F" authorityName="De Man" authorityYear="1920" box="[494,668,758,784]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E6C1FD5BFD57FC8F" box="[494,668,758,784]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E5FEFD5BFCB9FC8F" authorityName="Yaldwyn" authorityYear="1971" box="[721,882,758,784]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E5FEFD5BFCB9FC8F" box="[721,882,758,784]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
by the much shorter rostrum; from
<taxonomicName id="658FC8F84B6EFFA2E3D1FD5BFA57FC8F" authorityName="sensu De Man" authorityYear="1897" box="[1278,1436,758,784]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E3D1FD5BFA57FC8F" box="[1278,1436,758,784]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E7E7FCB7FEBCFCAB" authorityName="De Man" authorityYear="1920" box="[200,375,794,820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E7E7FCB7FEBCFCAB" box="[200,375,794,820]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
by the absence of dorsal and ventral transverse grooves on the palm of the male minor chela (well developed in
<taxonomicName id="658FC8F84B6EFFA2E65DFC93FDD9FCC8" authorityName="sensu De Man" authorityYear="1897" box="[370,530,829,856]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E65DFC93FDD9FCC8" box="[370,530,829,856]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E565FC93FD32FCC8" authorityName="De Man" authorityYear="1920" box="[586,761,829,856]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E565FC93FD32FCC8" box="[586,761,829,856]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
); specifically from
<taxonomicName id="658FC8F84B6EFFA2E4F5FC93FBB1FCC8" authorityName="sensu De Man" authorityYear="1897" box="[986,1146,829,856]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E4F5FC93FBB1FCC8" box="[986,1146,829,856]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
by the dorsal shoulder of the major chela smoothly sloping into the adjacent transverse groove (
<emphasis id="90FB6F694B6EFFA2E48EFCCFFC0BFCE4" box="[929,960,866,891]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
protruding and overhanging the transverse groove in
<taxonomicName id="658FC8F84B6EFFA2E628FC28FE6EFC00" authorityName="sensu De Man" authorityYear="1897" box="[263,421,901,927]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E628FC28FE6EFC00" box="[263,421,901,927]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
); and specifically from
<taxonomicName id="658FC8F84B6EFFA2E581FC28FC85FC00" authorityName="Yaldwyn" authorityYear="1971" box="[686,846,901,927]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E581FC28FC85FC00" box="[686,846,901,927]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
by the much stronger distomesial ridge on the mesial face of the major chela pollex (which is feebly developed in
<taxonomicName id="658FC8F84B6EFFA2E47FFC04FC38FC5C" authorityName="Yaldwyn" authorityYear="1971" box="[848,1011,937,963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E47FFC04FC38FC5C" box="[848,1011,937,963]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
). In life,
<taxonomicName id="658FC8F84B6EFFA2E34FFC04FAECFC5C" authorityName="sensu De Man" authorityYear="1897" box="[1120,1319,937,963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E34FFC04FAECFC5C" box="[1120,1319,937,963]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B6EFFA2E217FC04FA5CFC5C" box="[1336,1431,937,963]" captionStart="FIGURE 21" captionStartId="43.[151,250,1724,1748]" captionTargetBox="[151,1435,196,1685]" captionTargetId="figure-15@43.[151,1435,196,1685]" captionTargetPageId="43" captionText="FIGURE 21. Alpheus microrhynchus De Man, 1897: ovigerous female (cl indet.) imported to Germany from Thailand [A]; ovigerous female (cl indet.) imported to Singapore from Thailand [BE]: A, B—habitus, lateral view; C—pleon and tail fan, dorsal view; D—pleon, ventrolateral view E—frontal region and chelipeds, dorsal view. Aquarium-kept specimens, probably not preserved / deposited. Note the low number and large size of developing embryos inA and D. Photographs courtesy of Chris Lukhaup (A) and Paul Ng (BE)." figureDoi="http://doi.org/10.5281/zenodo.7912358" httpUri="https://zenodo.org/record/7912358/files/figure.png" pageId="42" pageNumber="43">Figs. 21</figureCitation>
,
<figureCitation id="3AB4AFFE4B6EFFA2E7B8FC60FF7FFC78" box="[151,180,973,999]" captionStart="FIGURE 22" captionStartId="44.[151,250,1907,1931]" captionTargetBox="[226,1361,193,1884]" captionTargetId="figure-21@44.[226,1361,181,1884]" captionTargetPageId="44" captionText="FIGURE 22. Alpheus microrhynchus De Man, 1897: female (cl indet.) imported to Singapore from Thailand; A, B—habitus, dorsal view (with one Singapore dollar coin, diameter 22.4 mm, for scale in B); C—detail of frontal margin; D—major (right) chela, mesial view; E—posterior pleonites and tail fan, d orsal view. Aquarium-kept specimen, probably not preserved / deposited. Photographs courtesy of Paul Ng." figureDoi="http://doi.org/10.5281/zenodo.7912362" httpUri="https://zenodo.org/record/7912362/files/figure.png" pageId="42" pageNumber="43">22</figureCitation>
) can be distinguished from
<taxonomicName id="658FC8F84B6EFFA2E6D4FC60FD56FC79" authorityName="sensu De Man" authorityYear="1897" box="[507,669,972,999]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E6D4FC60FD56FC79" box="[507,669,972,999]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B6EFFA2E580FC61FCD4FC78" box="[687,799,972,999]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="19.[151,250,1902,1926]" captionStartId-1="20.[151,250,1875,1899]" captionStartId-2="21.[151,250,1866,1890]" captionTargetBox-0="[151,1435,181,1877]" captionTargetBox-1="[151,1435,181,1851]" captionTargetBox-2="[159,1429,181,1842]" captionTargetId-0="figure-15@19.[151,1435,181,1878]" captionTargetId-1="figure-21@20.[151,1435,181,1851]" captionTargetId-2="figure-15@21.[159,1429,181,1842]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." captionText-1="FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae." captionText-2="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7912312" figureDoi-1="http://doi.org/10.5281/zenodo.7912318" figureDoi-2="http://doi.org/10.5281/zenodo.7912320" httpUri-0="https://zenodo.org/record/7912312/files/figure.png" httpUri-1="https://zenodo.org/record/7912318/files/figure.png" httpUri-2="https://zenodo.org/record/7912320/files/figure.png" pageId="42" pageNumber="43">Fig. 810</figureCitation>
) and
<taxonomicName id="658FC8F84B6EFFA2E44CFC60FBCCFC79" authorityName="Yaldwyn" authorityYear="1971" box="[867,1031,972,999]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E44CFC60FBCCFC79" box="[867,1031,972,999]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B6EFFA2E336FC61FBA5FC78" box="[1049,1134,972,999]" captionStart="FIGURE 17" captionStartId="35.[151,250,1930,1954]" captionTargetBox="[151,1435,237,1907]" captionTargetId="figure-46@35.[151,1435,237,1907]" captionTargetPageId="35" captionText="FIGURE 17. Alpheus richardsoni Yaldwyn, 1971: male (cl 8.3 mm) from Loders Creek, Moreton Bay, Queensland, Australia (FLMNH UF 18694) [A]; male (cl 9.3 mm) from Cleveland, Queensland, Australia (OUMNH.ZC 2009-09-0015); A, B— habitus, dorsal view; C—same, lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.7912344" httpUri="https://zenodo.org/record/7912344/files/figure.png" pageId="42" pageNumber="43">Fig. 17</figureCitation>
,
<figureCitation id="3AB4AFFE4B6EFFA2E352FC60FB51FC78" box="[1149,1178,973,999]" captionStart="FIGURE 18" captionStartId="36.[151,250,1909,1933]" captionTargetBox="[221,1366,181,1885]" captionTargetId="figure-21@36.[221,1366,181,1885]" captionTargetPageId="36" captionText="FIGURE 18. Alpheus richardsoni Yaldwyn, 1971: male (cl indet.) from Brunswisk River, New South Wales, Australia [A, B]; male from Parengarenga (Paua), New Zealand [C]; male from Whangarei, New Zealand [D]; A, C—habitus, dorsal view; B—same, lateral view; D—tail fan, dorsal view. Specimens not deposited / deposition unknown. Photographs courtesy of Denis Riek (A, B) and Richard B. Taylor (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7912350" httpUri="https://zenodo.org/record/7912350/files/figure.png" pageId="42" pageNumber="43">18</figureCitation>
) by its colour pattern, which is more similar to that of
<taxonomicName id="658FC8F84B6EFFA2E52DFC5CFD7AFB95" authorityName="De Man" authorityYear="1920" box="[514,689,1008,1035]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E52DFC5CFD7AFB95" box="[514,689,1008,1035]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B6EFFA2E5EEFC5DFCDEFB95" box="[705,789,1008,1034]" captionStart="FIGURE 16" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[152,1435,181,1885]" captionTargetId="figure-15@31.[152,1435,181,1885]" captionTargetPageId="31" captionText="FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (AC) and Sukkrit Nimitkul (D)." figureDoi="http://doi.org/10.5281/zenodo.7912340" httpUri="https://zenodo.org/record/7912340/files/figure.png" pageId="42" pageNumber="43">Fig. 16</figureCitation>
). There also seem to be an ecological separation between
<taxonomicName id="658FC8F84B6EFFA2E7B8FBB9FE96FBB1" authorityName="sensu De Man" authorityYear="1897" box="[151,349,1044,1070]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E7B8FBB9FE96FBB1" box="[151,349,1044,1070]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
,
<taxonomicName id="658FC8F84B6EFFA2E644FBB9FDC0FBB1" authorityName="sensu De Man" authorityYear="1897" box="[363,523,1044,1070]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B6EFFA2E644FBB9FDC0FBB1" box="[363,523,1044,1070]" italics="true" pageId="42" pageNumber="43">A. euphrosyne</emphasis>
</taxonomicName>
,
<taxonomicName id="658FC8F84B6EFFA2E536FBB9FD0CFBB1" authorityName="De Man" authorityYear="1920" box="[537,711,1044,1070]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E536FBB9FD0CFBB1" box="[537,711,1044,1070]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="658FC8F84B6EFFA2E5D1FBB9FC6BFBB1" authorityName="Yaldwyn" authorityYear="1971" box="[766,928,1044,1070]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="richardsoni">
<emphasis id="90FB6F694B6EFFA2E5D1FBB9FC6BFBB1" box="[766,928,1044,1070]" italics="true" pageId="42" pageNumber="43">A. richardsoni</emphasis>
</taxonomicName>
, with only the former species being confined to oligohaline habitats (sections of rivers, lagoons and canals under weak tidal influence). For separation of
<taxonomicName id="658FC8F84B6EFFA2E2ABFB95FEF6FBEA" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E2ABFB95FEF6FBEA" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
from other species treated in this study see below or refer to
<tableCitation id="EF0D86C04B6EFFA2E4C5FBF6FBF5FBE9" box="[1002,1086,1115,1142]" captionStart="TABLE 1" captionStartId="93.[145,233,465,491]" captionTargetPageId="93" captionText="TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897—A. micro-" httpUri="http://table.plazi.org/id/F6F0E3F34B19FFD5E7BEFE7CF8F8FD90" pageId="42" pageNumber="43" tableUuid="F6F0E3F34B19FFD5E7BEFE7CF8F8FD90">Table 1</tableCitation>
.
</paragraph>
<paragraph id="A230B37B4B6EFFA2E7E8FBD2FAFDFAEF" blockId="42.[151,1437,151,2036]" pageId="42" pageNumber="43">
<bibRefCitation id="C61ECE8A4B6EFFA2E7E8FBD2FEA7FB06" author="Kemp, S." box="[199,364,1151,1177]" pageId="42" pageNumber="43" pagination="81 - 102" refId="ref72298" refString="Kemp, S. (1918) Crustacea Decapoda of the Inle Lake Basin. Records of the Indian Museum, 14, 81 - 102, pls. 24 + 25. https: // doi. org / 10.26515 / rzsi / v 14 / i 4 / 1918 / 163580" type="journal article" year="1918">Kemps (1918)</bibRefCitation>
record of
<taxonomicName id="658FC8F84B6EFFA2E6F5FB2DFDA7FB06" authorityName="Kemp" authorityYear="1915" box="[474,620,1151,1178]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="90FB6F694B6EFFA2E6F5FB2DFDA7FB06" box="[474,620,1151,1178]" italics="true" pageId="42" pageNumber="43">A. paludicola</emphasis>
</taxonomicName>
from Thale Sap (
<collectingRegion id="604B7D994B6EFFA2E407FBD2FC5AFB06" box="[808,913,1151,1177]" country="Thailand" name="Songkhla" pageId="42" pageNumber="43">Songkhla</collectingRegion>
Lake) in southern
<collectingCountry id="DA98F3EB4B6EFFA2E373FBD2FB75FB06" box="[1116,1214,1151,1177]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
most likely refers to
<taxonomicName id="658FC8F84B6EFFA2E7B8FB0EFE93FB22" authorityName="sensu De Man" authorityYear="1897" box="[151,344,1187,1213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E7B8FB0EFE93FB22" box="[151,344,1187,1213]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
. Kemp noted that the Thale Sap specimens had a distinctly longer rostrum, “extending considerably beyond the end of the orbital hoods”. They also had a different colour pattern, described by the collector, Dr. N. Annandale, as “translucent, without definite markings, but tinged ... with reddish brown, ... palm and fingers of both chelae were deeply tinged with blue, especially on the dorsal surface ... eggs were pale green”. All females had very large eggs,
<quantity id="65771E9E4B6EFFA2E63EFA9FFE58FAD3" box="[273,403,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="42" pageNumber="43" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
in diameter. This combination of characters agrees more with the diagnosis of
<taxonomicName id="658FC8F84B6EFFA2E3F2FA9FFA57FAD3" authorityName="sensu De Man" authorityYear="1897" box="[1245,1436,1330,1356]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E3F2FA9FFA57FAD3" box="[1245,1436,1330,1356]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
than with that of
<emphasis id="90FB6F694B6EFFA2E67CFAFBFE26FAF0" box="[339,493,1365,1392]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="658FC8F84B6EFFA2E67CFAFBFE22FAF0" authorityName="Kemp" authorityYear="1915" box="[339,489,1365,1392]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="paludicola">A. paludicola</taxonomicName>
,
</emphasis>
although in the latter species, females bear equally large eggs (
<bibRefCitation id="C61ECE8A4B6EFFA2E38CFAFBFAE1FAEF" author="Kemp, S." box="[1187,1322,1365,1392]" pageId="42" pageNumber="43" pagination="201 - 325" refId="ref72246" refString="Kemp, S. (1915) Fauna of the Chilka Lake. No. 3. Crustacea Decapoda. Memoirs of the Indian Museum, 5, 201 - 325, pls. 12 - 13. https: // doi. org / 10.5962 / bhl. title. 10414" type="journal article" year="1915">Kemp 1915</bibRefCitation>
).
</paragraph>
<paragraph id="A230B37B4B6EFFA2E7E8FAD4FBCEF9D9" blockId="42.[151,1437,151,2036]" pageId="42" pageNumber="43">
<bibRefCitation id="C61ECE8A4B6EFFA2E7E8FAD4FEB4FA0B" author="Johnson, D. S." box="[199,383,1401,1428]" pageId="42" pageNumber="43" pagination="7 - 11" refId="ref72065" refString="Johnson, D. S. (1965) A review of brackish water prawns of Malaya. Bulletin of the National Museum of Singapore, 33, 7 - 11." type="journal article" year="1965">Johnsons (1965</bibRefCitation>
,
<bibRefCitation id="C61ECE8A4B6EFFA2E6A4FAD4FE0FFA0C" author="Johnson, D. S." box="[395,452,1401,1427]" pageId="42" pageNumber="43" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">1976</bibRefCitation>
) records of
<taxonomicName id="658FC8F84B6EFFA2E566FAD7FCC6FA0C" authorityName="sensu De Man" authorityYear="1897" box="[585,781,1401,1428]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E566FAD7FCC6FA0C" box="[585,781,1401,1428]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
from
<collectingCountry id="DA98F3EB4B6EFFA2E47EFAD4FC08FA0C" box="[849,963,1401,1427]" name="Singapore" pageId="42" pageNumber="43">Singapore</collectingCountry>
are probably based on material containing at least two species. All large-sized specimens (tl&gt;
<quantity id="65771E9E4B6EFFA2E5F1FA30FCFBFA28" box="[734,816,1437,1464]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="50.0">50 mm</quantity>
) identified as
<taxonomicName id="658FC8F84B6EFFA2E4FCFA30FB51FA28" authorityName="sensu De Man" authorityYear="1897" box="[979,1178,1437,1463]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E4FCFA30FB51FA28" box="[979,1178,1437,1463]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
by D.S. Johnson were herein reidentified as either
<taxonomicName id="658FC8F84B6EFFA2E6F0FA6CFD5BFA44" authorityName="De Man" authorityYear="1920" box="[479,656,1473,1499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E6F0FA6CFD5BFA44" box="[479,656,1473,1499]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
or
<taxonomicName id="658FC8F84B6EFFA2E593FA6CFCDCFA44" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[700,791,1473,1499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B6EFFA2E593FA6CFCDCFA44" box="[700,791,1473,1499]" italics="true" pageId="42" pageNumber="43">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B6EFFA2E40EFA6CFCB5FA44" bold="true" box="[801,894,1473,1499]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="8BC8D2124B6EFFA2E40EFA6CFCB5FA44" box="[801,894,1473,1499]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see below), whereas two smaller problematic females (tl&lt;
<quantity id="65771E9E4B6EFFA2E633FA49FEA7FA60" box="[284,364,1508,1535]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="mm" value="30.0">30 mm</quantity>
) were included under
<taxonomicName id="658FC8F84B6EFFA2E54EFA48FCFAFA61" authorityName="De Man" authorityYear="1920" box="[609,817,1508,1535]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E54EFA48FDB1FA60" box="[609,634,1509,1535]" italics="true" pageId="42" pageNumber="43">A.</emphasis>
cf.
<emphasis id="90FB6F694B6EFFA2E58DFA49FCFAFA61" box="[674,817,1508,1534]" italics="true" pageId="42" pageNumber="43">eurydactylus</emphasis>
</taxonomicName>
B. (see above). Although the historical occurrence of
<taxonomicName id="658FC8F84B6EFFA2E2ABFA48FEF6F9BD" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E2ABFA48FEF6F9BD" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
in
<collectingCountry id="DA98F3EB4B6EFFA2E64DF9A5FE1FF9BD" box="[354,468,1544,1570]" name="Singapore" pageId="42" pageNumber="43">Singapore</collectingCountry>
cannot be excluded, it is rather unlikely that the species occurs there today, after most of transitional freshwater-to-brackish habitats were lost to urban development.
</paragraph>
<paragraph id="A230B37B4B6EFFA2E7E8F9FDFAA3F86C" blockId="42.[151,1437,151,2036]" pageId="42" pageNumber="43">
Two specimens from Bang Pakong River east of
<collectingRegion id="604B7D994B6EFFA2E435F9FDFC49F9F5" box="[794,898,1616,1642]" country="Thailand" name="Krung Thep Mahanakhon" pageId="42" pageNumber="43">Bangkok</collectingRegion>
,
<collectingCountry id="DA98F3EB4B6EFFA2E4BCF9FDFC3CF9F5" box="[915,1015,1616,1642]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
(USNM 65477/1), were tentatively identified as
<taxonomicName id="658FC8F84B6EFFA2E604F9D9FDDCF912" authorityName="sensu De Man" authorityYear="1897" box="[299,535,1651,1678]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E604F9D9FE8FF911" box="[299,324,1652,1678]" italics="true" pageId="42" pageNumber="43">A.</emphasis>
cf.
<emphasis id="90FB6F694B6EFFA2E65EF9DEFDDCF912" box="[369,535,1651,1677]" italics="true" pageId="42" pageNumber="43">microrhynchus</emphasis>
</taxonomicName>
, because they deviate even more from the present redescription of the species. The third pereiopod merus of these specimens is somewhat broader, reaching the proportions of its homolog in
<taxonomicName id="658FC8F84B6EFFA2E2ABF935FEEDF94A" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E2ABF935FEEDF94A" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
. Most importantly, the minor chela of the male specimen has a more broadened, balaeniceps-shaped dactylus, almost as broad as in
<taxonomicName id="658FC8F84B6EFFA2E6C4F972FD53F966" authorityName="De Man" authorityYear="1920" box="[491,664,1759,1785]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E6C4F972FD53F966" box="[491,664,1759,1785]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
. On the other hand, the major chela has no trace of granulation (which is present in
<taxonomicName id="658FC8F84B6EFFA2E609F8AEFE1EF883" authorityName="De Man" authorityYear="1920" box="[294,469,1794,1821]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E609F8AEFE1EF883" box="[294,469,1794,1821]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
) and its dorsal shoulder is sloping very gently into the adjacent transverse groove (vs. sloping with a stronger angle in
<taxonomicName id="658FC8F84B6EFFA2E528F88BFD7DF8DF" authorityName="De Man" authorityYear="1920" box="[519,694,1830,1856]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B6EFFA2E528F88BFD7DF8DF" box="[519,694,1830,1856]" italics="true" pageId="42" pageNumber="43">A. eurydactylus</emphasis>
</taxonomicName>
). The antennal basicerite of these specimens is unarmed, as in
<emphasis id="90FB6F694B6EFFA2E2ABF88BFEE6F8FB" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="658FC8F84B6EFFA2E2ABF88BFEE2F8FB" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="eurydactylus">A. eurydactylus</taxonomicName>
,
</emphasis>
which contradicts
<bibRefCitation id="C61ECE8A4B6EFFA2E52CF8E7FD08F8FB" author="De Man, J. G." box="[515,707,1866,1892]" pageId="42" pageNumber="43" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Mans (1897)</bibRefCitation>
original diagnosis of
<taxonomicName id="658FC8F84B6EFFA2E496F8E7FBB5F8FB" authorityName="sensu De Man" authorityYear="1897" box="[953,1150,1866,1892]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E496F8E7FBB5F8FB" box="[953,1150,1866,1892]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
(see above) and does not allow assigning them to
<taxonomicName id="658FC8F84B6EFFA2E683F8C3FD89F818" box="[428,578,1901,1928]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cyanoteles">
<emphasis id="90FB6F694B6EFFA2E683F8C3FD89F818" box="[428,578,1901,1928]" italics="true" pageId="42" pageNumber="43">A. cyanoteles</emphasis>
</taxonomicName>
, the latter with a well-developed tooth (see below). Furthermore, it remains unknown where in the river (i.e., at the mouth or more upstream) these specimens were collected. More fresh material from Bang Pakong River and other parts of
<collectingCountry id="DA98F3EB4B6EFFA2E5C8F818FC86F850" box="[743,845,1973,1999]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
, as well as other localities across South-East Asia, is needed for a molecular analysis of various populations of
<taxonomicName id="658FC8F84B6EFFA2E40FF874FC2DF86C" authorityName="sensu De Man" authorityYear="1897" box="[800,998,2009,2035]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B6EFFA2E40FF874FC2DF86C" box="[800,998,2009,2035]" italics="true" pageId="42" pageNumber="43">A. microrhynchus</emphasis>
</taxonomicName>
and confirmation of their identity.
</paragraph>
</subSubSection>
</treatment>
</document>