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<mods:title id="C6A449AACBFF90E53CCEA49B2FAC7B1B">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<taxonomicName id="658FC8F84B53FF9FE7B8FEE7FD9BFEFB" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[151,592,330,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurydactylus">
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<emphasis id="90FB6F694B53FF9FE7B8FEE7FE50FEFA" bold="true" box="[151,411,330,357]" italics="true" pageId="23" pageNumber="24">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE68DFEE6FD9BFEFB" author="De Man, J. G." box="[418,592,330,357]" pageId="23" pageNumber="24" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
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<paragraph id="A230B37B4B53FF9FE7B8FEC2FE7AFE16" blockId="23.[151,652,330,429]" box="[151,433,366,393]" pageId="23" pageNumber="24">
(
<figureCitation id="3AB4AFFE4B53FF9FE7B0FEC2FED1FE16" box="[159,282,367,393]" captionStart="FIGURE 3" captionStartId="8.[151,250,1711,1735]" captionTargetBox="[164,1423,188,1674]" captionTargetId="figure-21@8.[164,1423,177,1691]" captionTargetPageId="8" captionText="FIGURE 3. Three species of Alpheus described by Kemp (1915), De Man (1920) and Schmitt (1926). Alpheus paludicola Kemp, 1915: syntype, male (cl indet., tl 22.0 mm) from Chilika Lake, India [A–G]; A—frontal region, dorsal view; B—scaphocerite of left antenna, dorsal view; C—(left) major cheliped, mesial view; D—same, lateral view; E—minor (right) cheliped, distal portion, mesial view; F—second pereiopod, lateral view; G—telson, dorsal view (after Kemp 1915). Alpheus eurydactylus De Man, 1920: syntypes, male (cl 13.8 mm) [H] and ovigerous female (cl 15.0 mm) [I] from Java, Indonesia (see text for lectotype / paralectotype designation); H—male minor (right) chela, distal portion of palm and fingers, dorsal view; I—female (right) minor chela, lateral view (after De Man 1924). Alpheus langi (Schmitt, 1926), originally described as Crangon langi, presenly junior synonym of Alpheus pontederiae de Rochebrune, 1883: holotype, male (cl 11.0 mm, tl 34.0 mm) from Congo [J–M]: J—frontal region, dorsal view; K—major (left) chela, lateral view; L—minor (right) chela, lateral view; M—second pereiopod, lateral view (after Schmitt 1926). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7912300" httpUri="https://zenodo.org/record/7912300/files/figure.png" pageId="23" pageNumber="24">Figs. 3H, I</figureCitation>
, 
<figureCitation id="3AB4AFFE4B53FF9FE609FEC2FEA5FE16" box="[294,366,367,393]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStartId-0="25.[151,250,1892,1916]" captionStartId-1="26.[151,250,1819,1843]" captionStartId-2="27.[151,250,885,909]" captionStartId-3="28.[151,250,1839,1863]" captionStartId-4="29.[151,250,1821,1845]" captionStartId-5="31.[151,250,1908,1932]" captionTargetBox-0="[158,1429,181,1868]" captionTargetBox-1="[151,1435,181,1795]" captionTargetBox-2="[151,1435,181,861]" captionTargetBox-3="[174,1408,280,1804]" captionTargetBox-4="[167,1423,194,1791]" captionTargetBox-5="[152,1435,181,1885]" captionTargetId-0="figure-15@25.[158,1429,181,1868]" captionTargetId-1="figure-21@26.[151,1435,181,1795]" captionTargetId-2="figure-15@27.[151,1435,181,861]" captionTargetId-3="figure-62@28.[164,1423,273,1815]" captionTargetId-4="figure-15@29.[151,1436,181,1798]" captionTargetId-5="figure-15@31.[152,1435,181,1885]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="29" captionTargetPageId-5="31" captionText-0="FIGURE 11. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—cephalothorax and chelipeds, dorsal view; B—frontal region, dorsal view; C, D— major (left) cheliped, mesial view, with slightly different angles and illuminations (slightly more ventral and enhancing mesial subdistal ridge on pollex in D). Refer to A for scale." captionText-1="FIGURE 12. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) [A–E] and paralectotype, ovigerous female (cl 15.0 mm) [F, G], both from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—major (left) cheliped, lateral view; B—same, chela, latreral view; C—same, fingers open, mesial view (see also Fig. 10C); D—minor chela in situ, dorsal view (cf. Fig. 3H); E—same, lateral view; F—frontal region, lateral view; G—frontal region and chelipeds in situ, dorsal view. Refer to Fig. 11A for scale in A–E." captionText-2="FIGURE 13. Alpheus eurydactylus De Man, 1920: female (cl 21.3 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49664): A—habitus, lateral view; B—minor (left) chela in situ, mesial view; C—major (right) cheliped in situ, mesial view; D—same, lateral view. Refer to A for scale." captionText-3="FIGURE 14. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–H, J–N]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [I]; A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—same, ischium, lateral view; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—same, basis and ischium, lateral view; M—second pleopod, appendices masculina and interna, anterior view; N—right uropod, dorsal view." captionText-4="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." captionText-5="FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D)." figureDoi-0="http://doi.org/10.5281/zenodo.7912324" figureDoi-1="http://doi.org/10.5281/zenodo.7912328" figureDoi-2="http://doi.org/10.5281/zenodo.7912332" figureDoi-3="http://doi.org/10.5281/zenodo.7912334" figureDoi-4="http://doi.org/10.5281/zenodo.7912338" figureDoi-5="http://doi.org/10.5281/zenodo.7912340" httpUri-0="https://zenodo.org/record/7912324/files/figure.png" httpUri-1="https://zenodo.org/record/7912328/files/figure.png" httpUri-2="https://zenodo.org/record/7912332/files/figure.png" httpUri-3="https://zenodo.org/record/7912334/files/figure.png" httpUri-4="https://zenodo.org/record/7912338/files/figure.png" httpUri-5="https://zenodo.org/record/7912340/files/figure.png" pageId="23" pageNumber="24">11–16</figureCitation>
, 
<figureCitation id="3AB4AFFE4B53FF9FE656FEC3FE62FE16" box="[377,425,366,393]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="23" pageNumber="24">51B</figureCitation>
)
</paragraph>
<paragraph id="A230B37B4B53FF9FE7B8FE3EFD47FE32" blockId="23.[151,652,330,429]" box="[151,652,402,429]" pageId="23" pageNumber="24">
[see also 
<figureCitation id="3AB4AFFE4B53FF9FE7D0FE3EFE49FE32" box="[255,386,402,429]" captionStart="FIGURE 53" captionStartId="107.[151,250,1859,1883]" captionTargetBox="[239,1349,192,1835]" captionTargetId="figure-15@107.[239,1349,181,1835]" captionTargetPageId="107" captionText="FIGURE 53. Alpheus euphrosyne De Man, 1897 complex in the northern and western Indian Ocean: A—Alpheus cf. mangalis sp. nov. (upper row) and A. euphrosyne (lower row) from an estuary near Kochi, India; B, C—Alpheus cf. eurydactylus De Man, 1920, ovigerious female (cl indet.) from Kochi, India (RMNH.Crus.D.49749), lateral view (B) and pleon and tail fan, dorsal view; D, E—Alpheus sp. euphrosyne complex (?), female photographed in situ in a mangrove at Watamu, Kenya. Photographs courtesy of Mahadevan Harikrishnan (A), Juan C. Miquel (B, C) and Dawn Goebbels (D, E)." figureDoi="http://doi.org/10.5281/zenodo.7912459" httpUri="https://zenodo.org/record/7912459/files/figure.png" pageId="23" pageNumber="24">Fig. 53B, C</figureCitation>
for 
<taxonomicName id="658FC8F84B53FF9FE69EFE3EFD48FE32" authorityName="De Man" authorityYear="1920" box="[433,643,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B53FF9FE69EFE3EFE02FE32" box="[433,457,403,429]" italics="true" pageId="23" pageNumber="24">A.</emphasis>
cf. 
<emphasis id="90FB6F694B53FF9FE6DBFE3EFD48FE32" box="[500,643,403,429]" italics="true" pageId="23" pageNumber="24">eurydactylus</emphasis>
</taxonomicName>
]
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<treatmentCitation id="232E956A4B53FF9FE7B8FE7AFDF2FE70" author="De Man, J. G." box="[151,569,471,495]" page="109" pageId="23" pageNumber="24" year="1920">
<taxonomicName id="658FC8F84B53FF9FE7B8FE7AFDF2FE70" ID-CoL="85LHH" authority="De Man 1920: 109" authorityName="De Man" authorityPageNumber="109" authorityYear="1920" box="[151,569,471,495]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B53FF9FE7B8FE7AFEBBFE70" box="[151,368,471,495]" italics="true" pageId="23" pageNumber="24">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE658FE75FDF2FE70" author="De Man, J. G." box="[375,569,471,495]" pageId="23" pageNumber="24" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man 1920: 109</bibRefCitation>
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; 
<treatmentCitation id="232E956A4B53FF9FE56CFE75FD33FE70" author="De Man, J. G." box="[579,760,471,495]" page="48" pageId="23" pageNumber="24" year="1924">
<bibRefCitation id="C61ECE8A4B53FF9FE56CFE75FD33FE70" author="De Man, J. G." box="[579,760,471,495]" pageId="23" pageNumber="24" pagination="1 - 60" refId="ref71119" refString="De Man, J. G. (1924) On a collection of macrurous decapod Crustacea chiefly Penaeidae and Alpheidae from the Indian Archipelago. Archiv fu ¨ r Naturgeschichte, Berlin, Series A, 90, 1 - 60." type="journal article" year="1924">De Man 1924: 48</bibRefCitation>
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, fig. 17.
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<emphasis id="90FB6F694B53FF9FE7B8FE5BFE48FD91" box="[151,387,502,526]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="658FC8F84B53FF9FE7B8FE5BFEAEFD91" ID-CoL="84W7P" authorityName="sensu De Man" authorityYear="1897" box="[151,357,502,526]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">Alpheus euphrosyne</taxonomicName>
.—
</emphasis>
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<bibRefCitation id="C61ECE8A4B53FF9FE6ACFE5BFD65FD91" author="Banner, A. H. &amp; Banner, D. M." box="[387,686,502,526]" pageId="23" pageNumber="24" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966: 130</bibRefCitation>
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(part.?), fig. 49; 
<treatmentCitation id="232E956A4B53FF9FE448FE5BFBEBFD91" author="Johnson, D. S." box="[871,1056,502,526]" page="37" pageId="23" pageNumber="24" year="1976">
<bibRefCitation id="C61ECE8A4B53FF9FE448FE5BFBEBFD91" author="Johnson, D. S." box="[871,1056,502,526]" pageId="23" pageNumber="24" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">Johnson 1976: 37</bibRefCitation>
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(part.) [not 
<taxonomicName id="658FC8F84B53FF9FE38BFE5BFF34FDB2" ID-CoL="84W7P" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B53FF9FE38BFE5BFAFCFD91" box="[1188,1335,502,526]" italics="true" pageId="23" pageNumber="24">A. euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE26FFE5BFF34FDB2" author="De Man, J. G." pageId="23" pageNumber="24" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B53FF9FE7B8FD9EFE8FFD16" blockId="23.[151,1437,471,710]" pageId="23" pageNumber="24">
(?) 
<emphasis id="90FB6F694B53FF9FE793FD9EFDE8FDD4" box="[188,547,563,587]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="658FC8F84B53FF9FE793FD9EFDCEFDD4" ID-CoL="84W7P" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[188,517,563,587]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">Alpheus euphrosyne euphrosyne</taxonomicName>
.—
</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE50CFD99FC8BFDD4" author="Banner, D. M. &amp; Banner, A. H." box="[547,832,563,587]" pageId="23" pageNumber="24" pagination="1 - 99" refId="ref69349" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 23</bibRefCitation>
, 92 (part.?); 
<bibRefCitation id="C61ECE8A4B53FF9FE4E7FD99FB39FDD4" author="Banner, D. M. &amp; Banner, A. H." box="[968,1266,563,587]" pageId="23" pageNumber="24" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982: 232</bibRefCitation>
, fig. 73 (part.?); 
<bibRefCitation id="C61ECE8A4B53FF9FE7E8FDFFFE14FDF5" author="Banner, A. H. &amp; Banner, D. M." box="[199,479,594,618]" pageId="23" pageNumber="24" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983: 30</bibRefCitation>
(part.?); 
<bibRefCitation id="C61ECE8A4B53FF9FE56FFDFFFC93FDF5" author="Banner, D. M. &amp; Banner, A. H." box="[576,856,594,618]" pageId="23" pageNumber="24" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. &amp; Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II. Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner &amp; Banner 1985: 16</bibRefCitation>
(part.?); 
<bibRefCitation id="C61ECE8A4B53FF9FE496FDFFFB90FDF5" author="Chace, F. A. Jr." box="[953,1115,594,618]" pageId="23" pageNumber="24" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988: 27</bibRefCitation>
(part.?) [not 
<taxonomicName id="658FC8F84B53FF9FE3CAFDFFFEF1FD16" ID-CoL="84W7P" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B53FF9FE3CAFDFFFABCFDF5" box="[1253,1399,594,618]" italics="true" pageId="23" pageNumber="24">A. euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE250FDFFFEF1FD16" author="De Man, J. G." pageId="23" pageNumber="24" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="A230B37B4B53FF9FE7B8FD22FBCCFD38" blockId="23.[151,1437,471,710]" box="[151,1031,655,679]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="829F94554B53FF9FE7B8FD22FBCCFD38" box="[151,1031,655,679]" pageId="23" pageNumber="24">
<taxonomicName id="658FC8F84B53FF9FE7B8FD22FE4DFD38" ID-CoL="84W7P" authorityName="sensu De Man" authorityYear="1897" box="[151,390,655,679]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B53FF9FE7B8FD22FF22FD38" box="[151,233,655,679]" italics="true" pageId="23" pageNumber="24">Alpheus</emphasis>
cf. 
<emphasis id="90FB6F694B53FF9FE620FD22FE4DFD38" box="[271,390,655,679]" italics="true" pageId="23" pageNumber="24">euphrosyne</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B53FF9FE6A8FD22FE40FD38" box="[391,395,655,679]" italics="true" pageId="23" pageNumber="24">.</emphasis>
— 
<treatmentCitation id="232E956A4B53FF9FE68CFD22FD84FD38" author="Marin, I. N." box="[419,591,655,679]" page="381" pageId="23" pageNumber="24" year="2021">
<bibRefCitation id="C61ECE8A4B53FF9FE68CFD22FD84FD38" author="Marin, I. N." box="[419,591,655,679]" pageId="23" pageNumber="24" pagination="369 - 385" refId="ref72908" refString="Marin, I. N. (2021) A new infaunal species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) and a new crustacean association from anoxic mangrove habitats in southern Vietnam. Arthropoda Selecta, 30, 369 - 385. https: // doi. org / 10.15298 / arthsel. 30.3.10" type="journal article" year="2021">Marin 2021: 381</bibRefCitation>
</treatmentCitation>
, fig. 1a [not 
<taxonomicName id="658FC8F84B53FF9FE5FFFD22FC36FD38" ID-CoL="84W7P" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[720,1021,655,679]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B53FF9FE5FFFD22FCABFD38" box="[720,864,655,679]" italics="true" pageId="23" pageNumber="24">A. euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE449FD3DFC36FD38" author="De Man, J. G." box="[870,1021,655,679]" pageId="23" pageNumber="24" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B53FF9FE7B8FD03FBECFD59" blockId="23.[151,1437,471,710]" box="[151,1063,686,710]" pageId="23" pageNumber="24">
(?) 
<taxonomicName id="658FC8F84B53FF9FE795FD03FEE4FD5A" box="[186,303,686,710]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B53FF9FE795FD03FEC7FD59" box="[186,268,686,710]" italics="true" pageId="23" pageNumber="24">Alpheus</emphasis>
sp.
</taxonomicName>
<taxonomicName id="658FC8F84B53FF9FE61AFD03FE64FD59" ID-CoL="85LF6" authorityName="Coutiere" authorityYear="1905" box="[309,431,686,710]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="audouini">
cf. 
<emphasis id="90FB6F694B53FF9FE67AFD03FE64FD59" box="[341,431,686,710]" italics="true" pageId="23" pageNumber="24">audouini</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B53FF9FE680FD03FE07FD59" box="[431,460,686,710]" italics="true" pageId="23" pageNumber="24">.—</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE6E3FD03FDB5FD59" author="Johnson, D. S." box="[460,638,686,710]" pageId="23" pageNumber="24" pagination="44 - 79" refId="ref72009" refString="Johnson, D. S. (1962) A synopsis of the Decapoda Caridea and Stenopodidea of Singapore, with notes on their distribution and a key to the genera of Caridea occuring in Malayan waters. Bulletin of the National Museum of Singapore, 30, 44 - 79, pl. 2." type="journal article" year="1962">Johnson 1962: 53</bibRefCitation>
(part.?) [not 
<taxonomicName id="658FC8F84B53FF9FE42BFD03FBD6FD59" ID-CoL="85LF6" authority="Coutiere, 1905" authorityName="Coutiere" authorityYear="1905" box="[772,1053,686,710]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="audouini">
<emphasis id="90FB6F694B53FF9FE42BFD03FCB0FD59" box="[772,891,686,710]" italics="true" pageId="23" pageNumber="24">A. audouini</emphasis>
<bibRefCitation id="C61ECE8A4B53FF9FE4AEFD03FBD6FD59" author="Coutiere, H." box="[897,1053,686,710]" pageId="23" pageNumber="24" pagination="852 - 921" refId="ref70260" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière, 1905</bibRefCitation>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B53FF90E7B8FD59FE6CFB22" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="A230B37B4B53FF9FE7B8FD59FEBEFCC9" blockId="23.[151,1437,756,2043]" pageId="23" pageNumber="24">
<emphasis id="90FB6F694B53FF9FE7B8FD59FEF4FC91" bold="true" box="[151,319,756,782]" pageId="23" pageNumber="24">Type material</emphasis>
. 
<materialsCitation id="12E7B9264B53FF9FE661FD58FC14FC91" ID-GBIF-Occurrence="4104078416" box="[334,991,756,783]" collectionCode="RMNH, ZMA" collectorName="Crus. D. &amp; P. Buitendijk" country="Indonesia" location="Surabaya" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Jawa Timur" typeStatus="lectotype">
<typeStatus id="7D340DD94B53FF9FE661FD58FE0AFC90" box="[334,449,757,783]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
, male (cl 
<quantity id="65771E9E4B53FF9FE51EFD59FD53FC90" box="[561,664,756,783]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="23" pageNumber="24" unit="mm" value="13.8">13.8 mm</quantity>
, tl ~40.0 mm, chl 21.0 mm),
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE4C8FD59FEBEFCC9" ID-GBIF-Occurrence="4104078531" collectionCode="RMNH, ZMA" collectorName="Crus. D. &amp; P. Buitendijk" country="Indonesia" location="Surabaya" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Jawa Timur" typeStatus="paralectotype">
<typeStatus id="7D340DD94B53FF9FE4C8FD59FB4AFC91" box="[999,1153,756,782]" pageId="23" pageNumber="24" type="paralectotype">paralectotype</typeStatus>
, ov. female (cl 15.0 mm, tl ~44.0 mm, chl 
<quantity id="65771E9E4B53FF9FE646FCB5FE1DFCAD" box="[361,470,792,819]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="23" pageNumber="24" unit="mm" value="21.5">21.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B53FF9FE6C2FCB4FD8AFCAC" box="[493,577,793,819]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49670 = 
<collectionCode id="C49E2BBE4B53FF9FE43FFCB5FC86FCAD" box="[784,845,792,818]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="23" pageNumber="24">ZMA</collectionCode>
Crus.D.102536, 
<collectingCountry id="DA98F3EB4B53FF9FE339FCB5FB4DFCAD" box="[1046,1158,792,818]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
, 
<collectingRegion id="604B7D994B53FF9FE3BAFCB4FB02FCAC" box="[1173,1225,793,819]" country="Indonesia" name="Jawa Timur" pageId="23" pageNumber="24">Java</collectingRegion>
, 
<location id="A750E5A04B53FF9FE3F6FCB5FA8FFCAD" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE3F6FCB5FA8FFCAD" box="[1241,1348,792,818]" country="Indonesia" name="Surabaya" pageId="23" pageNumber="24" stateProvince="Jawa Timur">Surabaya</location>
, leg. 
<collectorName id="0F7AD6AD4B53FF9FE2A6FCB4FEDBFCC9" pageId="23" pageNumber="24">P. Buitendijk</collectorName>
, no date.
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B53FF9FE7E8FCCDFF1CFAD1" blockId="23.[151,1437,756,2043]" pageId="23" pageNumber="24">
<emphasis id="90FB6F694B53FF9FE7E8FCCDFDFDFCE5" bold="true" box="[199,566,864,890]" pageId="23" pageNumber="24">Additional material examined</emphasis>
. 
<materialsCitation id="12E7B9264B53FF9FE567FCCDFE28FC00" ID-GBIF-Occurrence="4104078544" collectorName="Kamora &amp; A. Darmawan" country="Indonesia" location="Papua" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat">
<emphasis id="90FB6F694B53FF9FE567FCCDFD7DFCE5" box="[584,694,864,890]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="DA98F3EB4B53FF9FE567FCCDFD7DFCE5" box="[584,694,864,890]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B53FF9FE5E5FCCDFCD1FCE5" box="[714,794,864,890]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE460FCCDFC70FCE5" box="[847,955,864,890]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="23" pageNumber="24" unit="mm" value="19.4">19.4 mm</quantity>
), MNHN-IU-2018-5738, Papua, Kamora, mangrove, leg. A. Darmawan
</materialsCitation>
<emphasis id="90FB6F694B53FF9FE6C4FC28FDD6FC01" box="[491,541,900,926]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 
<date id="D63195BB4B53FF9FE51CFC29FD7EFC01" box="[563,693,900,926]" pageId="23" pageNumber="24" value="2012-07-08">07.08.2012</date>
; 
<materialsCitation id="12E7B9264B53FF9FE5EFFC29FCA5FC5C" ID-GBIF-Occurrence="4104078515" collectionCode="OUMNH, ZC" collectorName="A. Darmawan" country="Indonesia" location="Ajkwa Island" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat">
<specimenCount id="B48978F24B53FF9FE5EFFC29FCE9FC01" box="[704,802,900,926]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE460FC29FC73FC01" box="[847,952,900,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="23" pageNumber="24" unit="mm" value="12.6">12.6 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B53FF9FE3ABFC29FB26FC01" box="[1156,1261,900,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="23" pageNumber="24" unit="mm" value="17.3">17.3 mm</quantity>
), 
<collectionCode id="C49E2BBE4B53FF9FE22EFC29FABAFC01" box="[1281,1393,900,926]" pageId="23" pageNumber="24">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B53FF9FE25CFC29FA53FC01" box="[1395,1432,900,926]" country="Austria" name="Zoological Collection, University of Vienna" pageId="23" pageNumber="24" type="University or college">ZC</collectionCode>
. 2019.06.44, Papua, 
<location id="A750E5A04B53FF9FE657FC05FDD9FC5D" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE657FC05FDD9FC5D" box="[376,530,936,962]" country="Indonesia" name="Ajkwa Island" pageId="23" pageNumber="24" stateProvince="Papua Barat">Ajkwa Island</location>
, mangrove, leg. A. Darmawan
</materialsCitation>
<emphasis id="90FB6F694B53FF9FE458FC04FC61FC5D" box="[887,938,936,962]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 
<date id="D63195BB4B53FF9FE4EEFC05FB88FC5D" box="[961,1091,936,962]" pageId="23" pageNumber="24" value="2012-07-20">20.07.2012</date>
; 
<materialsCitation id="12E7B9264B53FF9FE360FC05FDB1FB95" ID-GBIF-Occurrence="4104078497" collectionCode="ZMA, RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Madura Island" municipality="Siboga Expedition" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B53FF9FE360FC05FB79FC5D" box="[1103,1202,936,962]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE3CEFC05FA80FC5D" box="[1249,1355,936,963]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="23" pageNumber="24" unit="mm" value="21.5">21.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B53FF9FE24FFC05FA56FC5D" box="[1376,1437,936,962]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="23" pageNumber="24">ZMA</collectionCode>
Crus.D.202791, 
<collectingMunicipality id="425429014B53FF9FE660FC61FDD5FC79" box="[335,542,972,998]" pageId="23" pageNumber="24">Siboga Expedition</collectingMunicipality>
sta. 4, “Strand” (beach), no further details; 1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE229FC60FA91FC78" box="[1286,1370,973,999]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus. D.49671, 
<location id="A750E5A04B53FF9FE629FC5DFE60FB95" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE629FC5DFE60FB95" box="[262,427,1008,1034]" country="Indonesia" municipality="Siboga Expedition" name="Madura Island" pageId="23" pageNumber="24">Madura Island</location>
, leg. P. Buitendijk
</materialsCitation>
, 03.1930; 
<materialsCitation id="12E7B9264B53FF9FE5C0FC5DFD70FBB1" ID-GBIF-Occurrence="4104078512" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Java" pageId="23" pageNumber="24" specimenCount="2" specimenCount-male="2" stateProvince="Jawa Timur">
<specimenCount id="B48978F24B53FF9FE5C0FC5DFC83FB95" box="[751,840,1008,1034]" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
, 1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE371FC5CFB79FB94" box="[1118,1202,1009,1035]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49660, Java, Tanjong Priok (near Jakarta), leg. P. Buitendijk
</materialsCitation>
, 1907; 
<materialsCitation id="12E7B9264B53FF9FE43BFBB9FC9BFBCD" ID-GBIF-Occurrence="4104078517" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Java" pageId="23" pageNumber="24" specimenCount="4" specimenCount-male="4" stateProvince="Jawa Timur">
<specimenCount id="B48978F24B53FF9FE43BFBB9FCBBFBB1" box="[788,880,1044,1070]" pageId="23" pageNumber="24" type="male">4 males</specimenCount>
, 1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE3B5FBB8FB25FBB0" box="[1178,1262,1045,1071]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49666, Java, Tanjong Priok (near Jakarta), leg. P. Buitendijk, 07.1927;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE478FB95FE39FBE9" ID-GBIF-Occurrence="4104078506" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Java" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Jawa Timur">
1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE370FB94FB78FBCC" box="[1119,1203,1081,1107]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49667, Java, Probolinggo, leg. P. Buitendijk
</materialsCitation>
, 04.1928; 
<materialsCitation id="12E7B9264B53FF9FE548FBF1FE97FB05" ID-GBIF-Occurrence="4104078526" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Sumatra" pageId="23" pageNumber="24" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sumatera Utara">
<specimenCount id="B48978F24B53FF9FE548FBF1FD7FFBE9" box="[615,692,1116,1142]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
, 
<specimenCount id="B48978F24B53FF9FE590FBF1FCE0FBE9" box="[703,811,1116,1142]" pageId="23" pageNumber="24" type="female">2 females</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE483FBF0FBCBFBE8" box="[940,1024,1117,1143]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49661, Sumatra, Pulau Weh, leg. P. Buitendijk
</materialsCitation>
, 1910; 
<materialsCitation id="12E7B9264B53FF9FE69FFB2DFA9BFB05" ID-GBIF-Occurrence="4104078508" box="[432,1360,1152,1179]" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="East Java" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur">
<specimenCount id="B48978F24B53FF9FE69FFB2DFE37FB05" box="[432,508,1152,1178]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE5AFFB2CFD1FFB04" box="[640,724,1153,1179]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49668, East Java, Passeroean, leg. P. Buitendijk
</materialsCitation>
, 1929; 
<materialsCitation id="12E7B9264B53FF9FE7B8FB09FB8FFB21" ID-GBIF-Occurrence="4104078369" box="[151,1092,1188,1215]" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="East Java" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur">
<specimenCount id="B48978F24B53FF9FE7B8FB09FF2FFB21" box="[151,228,1188,1214]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE645FB08FE75FB20" box="[362,446,1189,1215]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49665, East Java, Panaroekan, leg. P. Buitendijk
</materialsCitation>
, 08.1927; 
<materialsCitation id="12E7B9264B53FF9FE393FB09FD63FA99" ID-GBIF-Occurrence="4104078422" collectionCode="RMNH" collectorName="Crus. D." country="Indonesia" latitude="-5.9333334" location="Pulau Pamudjan Besar" longLatPrecision="1305" longitude="106.2" municipality="Baai" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat">
<specimenCount id="B48978F24B53FF9FE393FB09FAD4FB21" box="[1212,1311,1188,1214]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B53FF9FE7B8FB64FF20FB7C" box="[151,235,1225,1251]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49408, West Java, Banten 
<collectingMunicipality id="425429014B53FF9FE5A6FB65FD74FB7D" box="[649,703,1224,1250]" pageId="23" pageNumber="24">Baai</collectingMunicipality>
, 
<location id="A750E5A04B53FF9FE5E0FB65FC1CFB7C" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE5E0FB65FC1CFB7C" box="[719,983,1224,1251]" country="Indonesia" latitude="-5.9333334" longLatPrecision="1305" longitude="106.2" municipality="Baai" name="Pulau Pamudjan Besar" pageId="23" pageNumber="24" stateProvince="Jawa Barat">Pulau Pamudjan Besar</location>
, 
<geoCoordinate id="C7BBD5BC4B53FF9FE4C7FB65FBFDFB7C" box="[1000,1078,1224,1251]" degrees="5" direction="south" minutes="56" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-5.9333334">5°56’S</geoCoordinate>
<geoCoordinate id="C7BBD5BC4B53FF9FE36DFB65FB7BFB7D" box="[1090,1200,1224,1250]" degrees="106" direction="east" minutes="12" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="106.2">106°12’E</geoCoordinate>
, “laboratorium voor Onderzoek der zee, Batavia”, no further details
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE59CFB41FE5CFAB5" ID-GBIF-Occurrence="4104078398" collectionCode="RMNH" collectorName="P. Buitendijk" country="Indonesia" location="Java" municipality="Bay" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur">
<specimenCount id="B48978F24B53FF9FE59CFB41FCD8FA99" box="[691,787,1260,1286]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE411FB41FC6DFA99" box="[830,934,1260,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="23" pageNumber="24" unit="mm" value="21.3">21.3 mm</quantity>
), 
<collectionCode id="C49E2BBE4B53FF9FE497FB40FBC7FA98" box="[952,1036,1261,1287]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49664, Java, Surabaya, leg. P. Buitendijk, 02.1927;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE6B1FABDFA53FAB5" ID-GBIF-Occurrence="4104078463" box="[414,1432,1296,1323]" collectionCode="RMNH" collectorName="Crus. D. &amp; P. Buitendijk" country="Indonesia" location="Alkmaar Island" municipality="Bay" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Jawa Timur">
3 chelipeds, 
<collectionCode id="C49E2BBE4B53FF9FE505FABCFDB5FAB4" box="[554,638,1297,1323]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
Crus.D.49672, Java, 
<collectingMunicipality id="425429014B53FF9FE441FABCFC50FAB4" box="[878,923,1297,1323]" pageId="23" pageNumber="24">Bay</collectingMunicipality>
of Jakarta, 
<location id="A750E5A04B53FF9FE334FABDFB01FAB5" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE334FABDFB01FAB5" box="[1051,1226,1296,1322]" country="Indonesia" municipality="Bay" name="Alkmaar Island" pageId="23" pageNumber="24" stateProvince="Jawa Timur">Alkmaar Island</location>
, leg. P. Buitendijk
</materialsCitation>
, 1906.
</paragraph>
<paragraph id="A230B37B4B53FF9FE7E8FAF5FB7DF9F1" blockId="23.[151,1437,756,2043]" pageId="23" pageNumber="24">
<materialsCitation id="12E7B9264B53FF9FE7E8FAF5FD71FA09" ID-GBIF-Occurrence="4104078486" collectingDate="2001-11-22" collectionCode="OUMNH, ZC" collectorName="Nibong Tebal &amp; Z. Jaafar" country="Malaysia" location="Penang" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Pulau Pinang">
<emphasis id="90FB6F694B53FF9FE7E8FAF5FEE4FAED" box="[199,303,1368,1394]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="DA98F3EB4B53FF9FE7E8FAF5FEE4FAED" box="[199,303,1368,1394]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B53FF9FE611FAF5FE42FAED" box="[318,393,1368,1394]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE69AFAF5FDD7FAED" box="[437,540,1368,1394]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="23" pageNumber="24" unit="mm" value="11.2">11.2 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B53FF9FE5CBFAF5FC86FAED" box="[740,845,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="23" pageNumber="24" unit="mm" value="17.5">17.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B53FF9FE44FFAF5FC1BFAED" box="[864,976,1368,1394]" pageId="23" pageNumber="24">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B53FF9FE4FDFAF5FC3CFAED" box="[978,1015,1368,1394]" country="Austria" name="Zoological Collection, University of Vienna" pageId="23" pageNumber="24" type="University or college">ZC</collectionCode>
. 2019.06.45, 
<collectingRegion id="604B7D994B53FF9FE3A1FAF4FB2BFAEC" box="[1166,1248,1369,1395]" country="Malaysia" name="Pulau Pinang" pageId="23" pageNumber="24">Penang</collectingRegion>
(mainland part), 
<collectorName id="0F7AD6AD4B53FF9FE7B8FAD1FEFAFA09" box="[151,305,1404,1430]" pageId="23" pageNumber="24">Nibong Tebal</collectorName>
, mudflat, leg. 
<collectorName id="0F7AD6AD4B53FF9FE6E2FAD0FDFBFA09" box="[461,560,1404,1431]" pageId="23" pageNumber="24">Z. Jaafar</collectorName>
, 
<date id="D63195BB4B53FF9FE516FAD1FD71FA09" box="[569,698,1404,1430]" pageId="23" pageNumber="24" value="2001-11-22">
<collectingDate id="C6756C534B53FF9FE516FAD1FD71FA09" box="[569,698,1404,1430]" pageId="23" pageNumber="24" value="2001-11-22">22.11.2001</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="B48978F24B53FF9FE5ECFAD1FCD3FA09" box="[707,792,1404,1430]" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
(cl 11.5, 
<quantity id="65771E9E4B53FF9FE452FAD1FC2FFA09" box="[893,996,1404,1431]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="23" pageNumber="24" unit="mm" value="15.2">15.2 mm</quantity>
), 
<materialsCitation id="12E7B9264B53FF9FE4DAFAD1FC99FA25" ID-GBIF-Occurrence="4104078384" collectingDate="2001-11-22" collectionCode="OUMNH, ZC" collectorName="Nibong Tebal &amp; Z. Jaafar" country="Malaysia" location="Penang" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" stateProvince="Pulau Pinang">
<specimenCount id="B48978F24B53FF9FE4DAFAD1FBABFA09" box="[1013,1120,1404,1430]" pageId="23" pageNumber="24" type="female">2 females</specimenCount>
(cl 10.3, 
<quantity id="65771E9E4B53FF9FE3E9FAD1FAE7FA09" box="[1222,1324,1404,1430]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="23" pageNumber="24" unit="mm" value="11.7">11.7 mm</quantity>
), MNHN-IU-2018-5680, same collection data as for previous specimens
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE472FA0DFC4AFA41" ID-GBIF-Occurrence="4104078543" collectingDate="2009-10-02" collectorName="N. K. Ng &amp; J. C. Y. Lai" country="Malaysia" location="Muar" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Johor">
<specimenCount id="B48978F24B53FF9FE472FA0DFC6CFA25" box="[861,935,1440,1466]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE4FDFA0DFBF2FA25" box="[978,1081,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="23" pageNumber="24" unit="mm" value="15.1">15.1 mm</quantity>
), MNHN-IU-2018-5599, 
<collectingRegion id="604B7D994B53FF9FE277FA0DFA53FA25" box="[1368,1432,1440,1466]" country="Malaysia" name="Johor" pageId="23" pageNumber="24">Johor</collectingRegion>
, 
<location id="A750E5A04B53FF9FE7B8FA68FF1EFA40" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE7B8FA68FF1EFA40" box="[151,213,1477,1503]" country="Malaysia" name="Muar" pageId="23" pageNumber="24" stateProvince="Johor">Muar</location>
, mudflat, soggy mud, leg. 
<collectorName id="0F7AD6AD4B53FF9FE6D0FA68FD95FA40" box="[511,606,1477,1503]" pageId="23" pageNumber="24">N.K. Ng</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B53FF9FE5ADFA69FD3FFA41" box="[642,756,1476,1502]" pageId="23" pageNumber="24">J.C.Y. Lai</collectorName>
, 
<date id="D63195BB4B53FF9FE5D0FA69FC4AFA41" box="[767,897,1476,1502]" pageId="23" pageNumber="24" value="2009-10-02">
<collectingDate id="C6756C534B53FF9FE5D0FA69FC4AFA41" box="[767,897,1476,1502]" pageId="23" pageNumber="24" value="2009-10-02">10.02.2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE4A3FA69FD6FF99D" ID-GBIF-Occurrence="4104078485" collectingDate="2009-10-02" collectorName="N. K. Ng &amp; J. C. Y. Lai" country="Malaysia" location="Muar" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Johor">1 ov. female (cl indet.), MNHN-IU-2018-5598, same collection data as for previous specimen</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE59EFA45FC2CF9B8" ID-GBIF-Occurrence="4104078459" collectingDate="2002-05-11" collectionCode="ZRC" collectorName="Z. Jaafar" country="Malaysia" location="Telok Tebrau" pageId="23" pageNumber="24" specimenCode="ZRC 2009.0306" specimenCount="1" specimenCount-male="1" stateProvince="Terengganu">
<specimenCount id="B48978F24B53FF9FE59EFA45FD36F99D" box="[689,765,1512,1538]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE404FA45FC5EF99D" box="[811,917,1512,1539]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="23" pageNumber="24" unit="mm" value="15.1">15.1 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B53FF9FE34DFA45FB00F99D" box="[1122,1227,1512,1538]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="23" pageNumber="24" unit="mm" value="13.8">13.8 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE3F0FA45FA53F99D" box="[1247,1432,1512,1538]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 2009.0306</specimenCode>
, 
<collectingRegion id="604B7D994B53FF9FE7B8F9A0FED5F9B8" box="[151,286,1549,1575]" country="Malaysia" name="Terengganu" pageId="23" pageNumber="24">Terengganu</collectingRegion>
, 
<location id="A750E5A04B53FF9FE607F9A1FE75F9B9" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE607F9A1FE75F9B9" box="[296,446,1548,1574]" country="Malaysia" name="Telok Tebrau" pageId="23" pageNumber="24" stateProvince="Terengganu">Telok Tebrau</location>
, mudflats, with gobies, leg. 
<collectorName id="0F7AD6AD4B53FF9FE5D8F9A0FC90F9B9" box="[759,859,1548,1575]" pageId="23" pageNumber="24">Z. Jaafar</collectorName>
, 
<date id="D63195BB4B53FF9FE449F9A1FC2CF9B8" box="[870,999,1548,1575]" pageId="23" pageNumber="24" value="2002-05-11">
<collectingDate id="C6756C534B53FF9FE449F9A1FC2CF9B8" box="[870,999,1548,1575]" pageId="23" pageNumber="24" value="2002-05-11">05.11.2002</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE4DEF9A1FBDAF9D5" ID-GBIF-Occurrence="4104078564" collectingDate="1968-08-27" collectionCode="ZRC" collectorName="S. Kumar" country="Malaysia" location="Port Swettenham" municipality="Port Klang" pageId="23" pageNumber="24" specimenCode="ZRC 1979.4" specimenCount="1" specimenCount-male="1" stateProvince="Selangor">
<specimenCount id="B48978F24B53FF9FE4DEF9A1FBF7F9B9" box="[1009,1084,1548,1574]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE348F9A1FB04F9B9" box="[1127,1231,1548,1574]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="23" pageNumber="24" unit="mm" value="13.9">13.9 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE3CEF9A1FABBF9B9" box="[1249,1392,1548,1574]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1979.4</specimenCode>
.3.7, 
<collectingRegion id="604B7D994B53FF9FE7B8F99DFF37F9D5" box="[151,252,1584,1610]" country="Malaysia" name="Selangor" pageId="23" pageNumber="24">Selangor</collectingRegion>
, 
<collectingMunicipality id="425429014B53FF9FE626F99CFE4FF9D5" box="[265,388,1584,1611]" pageId="23" pageNumber="24">Port Klang</collectingMunicipality>
(
<location id="A750E5A04B53FF9FE6BBF99DFD96F9D5" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE6BBF99DFD96F9D5" box="[404,605,1584,1610]" country="Malaysia" municipality="Port Klang" name="Port Swettenham" pageId="23" pageNumber="24" stateProvince="Selangor">Port Swettenham</location>
), mudflats, leg. 
<collectorName id="0F7AD6AD4B53FF9FE43BF99DFC48F9D4" box="[788,899,1584,1611]" pageId="23" pageNumber="24">S. Kumar</collectorName>
, 
<date id="D63195BB4B53FF9FE4A0F99DFBDAF9D5" box="[911,1041,1584,1610]" pageId="23" pageNumber="24" value="1968-08-27">
<collectingDate id="C6756C534B53FF9FE4A0F99DFBDAF9D5" box="[911,1041,1584,1610]" pageId="23" pageNumber="24" value="1968-08-27">27.08.1968</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE332F99DFB79F9F1" ID-GBIF-Occurrence="4104078537" collectingDate="2009-10-02" collectorName="N. K. Ng &amp; J. C. Y. Lai" country="Malaysia" location="Taman Sri Sementa" municipality="Port Klang" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Selangor">
<specimenCount id="B48978F24B53FF9FE332F99DFBA1F9D5" box="[1053,1130,1584,1610]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE3B6F99DFAC9F9D5" box="[1177,1282,1584,1610]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="23" pageNumber="24" unit="mm" value="14.7">14.7 mm</quantity>
), MNHN-IU-2018-5595, 
<collectingRegion id="604B7D994B53FF9FE630F9F9FE4FF9F1" box="[287,388,1620,1646]" country="Malaysia" name="Selangor" pageId="23" pageNumber="24">Selangor</collectingRegion>
, 
<collectingMunicipality id="425429014B53FF9FE6A0F9F8FDC7F9F1" box="[399,524,1620,1647]" pageId="23" pageNumber="24">Port Klang</collectingMunicipality>
, 
<location id="A750E5A04B53FF9FE538F9F8FD3FF9F1" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE538F9F8FD3FF9F1" box="[535,756,1620,1647]" country="Malaysia" municipality="Port Klang" name="Taman Sri Sementa" pageId="23" pageNumber="24" stateProvince="Selangor">Taman Sri Sementa</location>
, leg. 
<collectorName id="0F7AD6AD4B53FF9FE41FF9F8FC44F9F0" box="[816,911,1621,1647]" pageId="23" pageNumber="24">N.K. Ng</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B53FF9FE49CF9F9FBEEF9F1" box="[947,1061,1620,1646]" pageId="23" pageNumber="24">J.C.Y. Lai</collectorName>
, 
<date id="D63195BB4B53FF9FE31EF9F9FB79F9F1" box="[1073,1202,1620,1646]" pageId="23" pageNumber="24" value="2009-10-02">
<collectingDate id="C6756C534B53FF9FE31EF9F9FB79F9F1" box="[1073,1202,1620,1646]" pageId="23" pageNumber="24" value="2009-10-02">10.02.2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B53FF9FE7E8F9D5FD76F8F5" blockId="23.[151,1437,756,2043]" pageId="23" pageNumber="24">
<materialsCitation id="12E7B9264B53FF9FE7E8F9D5FAFAF90D" ID-GBIF-Occurrence="4104078316" box="[199,1329,1656,1683]" collectingDate="1955-03-03" collectingDateMax="1996-04-18" collectingDateMin="1955-03-03" collectionCode="R, T" collectorName="P. K. L. Ng" country="Singapore" county="Geylang River" determinerName="D. S. Johnson" elevation="2" location="East Coast" municipality="Zoology Class" pageId="23" pageNumber="24" specimenCode="ZRC 2000.1220, ZRC 1996.0463, ZRC 1996.47, ZRC 2000.2145, ZRC 1979.4, ZRC 1991.10140" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<emphasis id="90FB6F694B53FF9FE7E8F9D5FEF2F90D" box="[199,313,1656,1682]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="DA98F3EB4B53FF9FE7E8F9D5FEF2F90D" box="[199,313,1656,1682]" name="Singapore" pageId="23" pageNumber="24">Singapore</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B53FF9FE665F9D5FE5DF90D" box="[330,406,1656,1682]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 18.0 mm), 
<specimenCode id="F2291B004B53FF9FE56CF9D5FD36F90D" box="[579,765,1656,1682]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 2000.1220</specimenCode>
, 
<location id="A750E5A04B53FF9FE426F9D4FC4EF90D" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE426F9D4FC4EF90D" box="[777,901,1656,1683]" country="Singapore" county="Geylang River" municipality="Zoology Class" name="East Coast" pageId="23" pageNumber="24">East Coast</location>
, mudflat, leg. 
<collectorName id="0F7AD6AD4B53FF9FE305F9D4FB55F90C" box="[1066,1182,1657,1683]" pageId="23" pageNumber="24">P.K.L. Ng</collectorName>
, 
<date id="D63195BB4B53FF9FE384F9D5FAE6F90D" box="[1195,1325,1656,1682]" pageId="23" pageNumber="24" value="1996-04-18">
<collectingDate id="C6756C534B53FF9FE384F9D5FAE6F90D" box="[1195,1325,1656,1682]" pageId="23" pageNumber="24" value="1996-04-18">18.04.1996</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE216F9D5FB53F928" ID-GBIF-Occurrence="4104078395" collectingDate="1955-03-03" collectingDateMax="1996-04-18" collectingDateMin="1955-03-03" collectionCode="R, T" collectorName="P. K. L. Ng" country="Singapore" county="Geylang River" determinerName="D. S. Johnson" elevation="2" location="East Coast" municipality="Zoology Class" pageId="23" pageNumber="24" specimenCode="ZRC 2000.1220, ZRC 1996.0463, ZRC 1996.47, ZRC 2000.2145, ZRC 1979.4, ZRC 1991.10140" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="B48978F24B53FF9FE216F9D5FA57F90D" box="[1337,1436,1656,1682]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE794F931FEE8F929" box="[187,291,1692,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="23" pageNumber="24" unit="mm" value="10.6">10.6 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE61AF931FE27F929" box="[309,492,1692,1718]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1996.0463</specimenCode>
, 
<location id="A750E5A04B53FF9FE6D8F931FD60F929" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE6D8F931FD60F929" box="[503,683,1692,1718]" country="Singapore" county="Geylang River" municipality="Zoology Class" name="Sungei Punggol" pageId="23" pageNumber="24">Sungei Punggol</location>
, sta. D1, mudflat, leg. 
<collectorName id="0F7AD6AD4B53FF9FE489F931FBC8F929" box="[934,1027,1692,1718]" pageId="23" pageNumber="24">R.E.S.T</collectorName>
., 
<date id="D63195BB4B53FF9FE33DF931FB5FF928" box="[1042,1172,1692,1719]" pageId="23" pageNumber="24" value="1995-06-28">
<collectingDate id="C6756C534B53FF9FE33DF931FB5FF928" box="[1042,1172,1692,1719]" pageId="23" pageNumber="24" value="1995-06-28">28.06.1995</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE3B0F931FB91F944" ID-GBIF-Occurrence="4104078338" collectingDate="1955-03-03" collectingDateMax="1996-04-18" collectingDateMin="1955-03-03" collectionCode="R, T" collectorName="P. K. L. Ng" country="Singapore" county="Geylang River" determinerName="D. S. Johnson" elevation="2" location="East Coast" municipality="Zoology Class" pageId="23" pageNumber="24" specimenCode="ZRC 2000.1220, ZRC 1996.0463, ZRC 1996.47, ZRC 2000.2145, ZRC 1979.4, ZRC 1991.10140" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="B48978F24B53FF9FE3B0F931FB34F929" box="[1183,1279,1692,1718]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 11.0 mm), 
<specimenCode id="F2291B004B53FF9FE7B8F96DFEFDF945" box="[151,310,1728,1754]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1996.47</specimenCode>
, 
<location id="A750E5A04B53FF9FE66BF96DFE57F945" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE66BF96DFE57F945" box="[324,412,1728,1754]" country="Singapore" county="Geylang River" municipality="Zoology Class" name="Mandai" pageId="23" pageNumber="24">Mandai</location>
, mangrove, burrow in soft mud, leg. 
<collectorName id="0F7AD6AD4B53FF9FE47FF96CFC0DF944" box="[848,966,1729,1755]" pageId="23" pageNumber="24">P.K.L. Ng</collectorName>
, 
<date id="D63195BB4B53FF9FE4FAF96DFB9CF944" box="[981,1111,1728,1755]" pageId="23" pageNumber="24" value="1984-05-18">
<collectingDate id="C6756C534B53FF9FE4FAF96DFB9CF944" box="[981,1111,1728,1755]" pageId="23" pageNumber="24" value="1984-05-18">18.05.1984</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE34AF96DFB88F961" ID-GBIF-Occurrence="4104078454" collectingDate="1955-03-03" collectingDateMax="1996-04-18" collectingDateMin="1955-03-03" collectionCode="R, T" collectorName="P. K. L. Ng" country="Singapore" county="Geylang River" determinerName="D. S. Johnson" elevation="2" location="East Coast" municipality="Zoology Class" pageId="23" pageNumber="24" specimenCode="ZRC 2000.1220, ZRC 1996.0463, ZRC 1996.47, ZRC 2000.2145, ZRC 1979.4, ZRC 1991.10140" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="B48978F24B53FF9FE34AF96DFB78F945" box="[1125,1203,1728,1754]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE3CAF96DFA9BF945" box="[1253,1360,1728,1754]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="23" pageNumber="24" unit="mm" value="10.8">10.8 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE249F96DFEDAF960" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 2000.2145</specimenCode>
, 
<location id="A750E5A04B53FF9FE630F949FE75F961" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE630F949FE75F961" box="[287,446,1764,1790]" country="Singapore" county="Geylang River" municipality="Zoology Class" name="Sungei Buloh" pageId="23" pageNumber="24">Sungei Buloh</location>
, mangrove, mud, leg. NUS 
<collectingMunicipality id="425429014B53FF9FE429F949FC65F961" box="[774,942,1764,1790]" pageId="23" pageNumber="24">Zoology Class</collectingMunicipality>
, 
<date id="D63195BB4B53FF9FE492F949FBF4F961" box="[957,1087,1764,1790]" pageId="23" pageNumber="24" value="1992-07-28">
<collectingDate id="C6756C534B53FF9FE492F949FBF4F961" box="[957,1087,1764,1790]" pageId="23" pageNumber="24" value="1992-07-28">28.07.1992</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B53FF9FE362F949FA53F8D9" ID-GBIF-Occurrence="4104078374" collectingDate="1955-03-03" collectingDateMax="1996-04-18" collectingDateMin="1955-03-03" collectionCode="R, T" collectorName="P. K. L. Ng" country="Singapore" county="Geylang River" determinerName="D. S. Johnson" elevation="2" location="East Coast" municipality="Zoology Class" pageId="23" pageNumber="24" specimenCode="ZRC 2000.1220, ZRC 1996.0463, ZRC 1996.47, ZRC 2000.2145, ZRC 1979.4, ZRC 1991.10140" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="B48978F24B53FF9FE362F949FB50F961" box="[1101,1179,1764,1790]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
(cl 15.0 mm, missing minor cheliped), 
<specimenCode id="F2291B004B53FF9FE676F8A5FE2CF8BD" box="[345,487,1800,1826]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1979.4</specimenCode>
.4.14, 
<collectingCounty id="4B51CBF74B53FF9FE504F8A5FD19F8BD" box="[555,722,1800,1826]" pageId="23" pageNumber="24">Geylang River</collectingCounty>
, sta. B71, mud bottom, depth: 
<quantity id="65771E9E4B53FF9FE300F8A5FB96F8BC" box="[1071,1117,1800,1827]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="m" value="2.0">
<elevation id="29A254484B53FF9FE300F8A5FB96F8BC" box="[1071,1117,1800,1827]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="m" value="2.0">2 m</elevation>
</quantity>
, leg. 
<collectorName id="0F7AD6AD4B53FF9FE3B4F8A5FADAF8BD" box="[1179,1297,1800,1826]" pageId="23" pageNumber="24">S.R.F.R.S.</collectorName>
, 
<date id="D63195BB4B53FF9FE231F8A5FA57F8BC" box="[1310,1436,1800,1827]" pageId="23" pageNumber="24" value="1955-03-03">
<collectingDate id="C6756C534B53FF9FE231F8A5FA57F8BC" box="[1310,1436,1800,1827]" pageId="23" pageNumber="24" value="1955-03-03">03.03.1955</collectingDate>
</date>
[J6851, det. 
<determinerName id="57EC5FBF4B53FF9FE60CF881FE73F8D9" box="[291,440,1836,1862]" pageId="23" pageNumber="24">D.S. Johnson</determinerName>
as 
<taxonomicName id="658FC8F84B53FF9FE6F1F881FD6FF8D9" authorityName="sensu De Man" authorityYear="1897" box="[478,676,1836,1862]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B53FF9FE6F1F881FD6FF8D9" box="[478,676,1836,1862]" italics="true" pageId="23" pageNumber="24">A. microrhynchus</emphasis>
</taxonomicName>
]; 
<specimenCount id="B48978F24B53FF9FE592F881FCD5F8D9" box="[701,798,1836,1862]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE464F881FC7AF8D8" box="[843,945,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="23" pageNumber="24" unit="mm" value="10.7">10.7 mm</quantity>
, identification tentative), 
<specimenCode id="F2291B004B53FF9FE3FDF881FA53F8D9" box="[1234,1432,1836,1862]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1991.10140</specimenCode>
</materialsCitation>
, 
<materialsCitation id="12E7B9264B53FF9FE7B8F8FDFD73F8F5" ID-GBIF-Occurrence="4104078510" box="[151,696,1872,1898]" collectingDate="1988-03-10" collectionCode="R, T" country="Singapore" location="River" pageId="23" pageNumber="24" specimenCount="1">
<collectingCountry id="DA98F3EB4B53FF9FE7B8F8FDFEC2F8F5" box="[151,265,1872,1898]" name="Singapore" pageId="23" pageNumber="24">Singapore</collectingCountry>
<location id="A750E5A04B53FF9FE63FF8FDFE9BF8F5" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE63FF8FDFE9BF8F5" box="[272,336,1872,1898]" country="Singapore" name="River" pageId="23" pageNumber="24">River</location>
, mud, leg. 
<collectorName id="0F7AD6AD4B53FF9FE6E4F8FDFDE0F8F5" box="[459,555,1872,1898]" pageId="23" pageNumber="24">R.E.S.T.</collectorName>
, 
<date id="D63195BB4B53FF9FE518F8FDFD73F8F5" box="[567,696,1872,1898]" pageId="23" pageNumber="24" value="1988-03-10">
<collectingDate id="C6756C534B53FF9FE518F8FDFD73F8F5" box="[567,696,1872,1898]" pageId="23" pageNumber="24" value="1988-03-10">03.10.1988</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B53FF90E7E8F8D9FEA8FF4B" blockId="23.[151,1437,756,2043]" lastBlockId="24.[151,1437,151,2035]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<materialsCitation id="12E7B9264B53FF9FE7E8F8D9FD40F811" ID-GBIF-Occurrence="4104078563" box="[199,651,1908,1935]" country="Philippines" location="Philippines" pageId="23" pageNumber="24" specimenCount="2" specimenCount-male="2">
<emphasis id="90FB6F694B53FF9FE7E8F8D9FE8CF811" box="[199,327,1908,1934]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="DA98F3EB4B53FF9FE7E8F8D9FE8CF811" box="[199,327,1908,1934]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B53FF9FE679F8D9FE66F811" box="[342,429,1908,1934]" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
(cl 13.8, 
<quantity id="65771E9E4B53FF9FE535F8D9FD48F811" box="[538,643,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="23" pageNumber="24" unit="mm" value="14.2">14.2 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="12E7B9264B53FF9FE5B8F8D9FE47F82D" ID-GBIF-Occurrence="4104078303" collectingDate="1996-03-26" collectionCode="ZRC" collectorName="S. H. Tan" country="Philippines" location="Manila" pageId="23" pageNumber="24" specimenCode="ZRC 1999.0235" specimenCount="1" specimenCount-female="1" stateProvince="Manila">
<specimenCount id="B48978F24B53FF9FE5B8F8D9FD32F811" box="[663,761,1908,1934]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE409F8D9FC44F811" box="[806,911,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="23" pageNumber="24" unit="mm" value="18.3">18.3 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE48CF8D9FB90F810" box="[931,1115,1908,1935]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 1999.0235</specimenCode>
, 
<collectingRegion id="604B7D994B53FF9FE348F8D9FB7DF811" box="[1127,1206,1908,1934]" country="Philippines" name="Manila" pageId="23" pageNumber="24">Manila</collectingRegion>
(?) fish market, leg. 
<collectorName id="0F7AD6AD4B53FF9FE7B8F835FF36F82C" box="[151,253,1944,1971]" pageId="23" pageNumber="24">S.H. Tan</collectorName>
, 
<date id="D63195BB4B53FF9FE625F835FE47F82D" box="[266,396,1944,1970]" pageId="23" pageNumber="24" value="1996-03-26">
<collectingDate id="C6756C534B53FF9FE625F835FE47F82D" box="[266,396,1944,1970]" pageId="23" pageNumber="24" value="1996-03-26">26.03.1996</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE6B6F835FACCF82D" ID-GBIF-Occurrence="4104078446" box="[409,1287,1944,1971]" collectingDate="1979-06-19" collectionCode="RMNH" collectorName="J. C. Miquel" country="Philippines" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B53FF9FE6B6F835FE34F82D" box="[409,511,1944,1970]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
, 
<collectionCode id="C49E2BBE4B53FF9FE524F834FD94F82C" box="[523,607,1945,1971]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24" type="Museum">RMNH</collectionCode>
<collectorName id="0F7AD6AD4B53FF9FE547F835FD77F82C" box="[616,700,1944,1971]" pageId="23" pageNumber="24">Crus.D.</collectorName>
49750, 
<collectorName id="0F7AD6AD4B53FF9FE43AF834FC90F82C" box="[789,859,1945,1971]" pageId="23" pageNumber="24">Panay</collectorName>
, Ilo Ilo, leg. 
<collectorName id="0F7AD6AD4B53FF9FE4DFF835FBB3F82D" box="[1008,1144,1944,1970]" pageId="23" pageNumber="24">J.C. Miquel</collectorName>
, 
<date id="D63195BB4B53FF9FE3AAF835FACCF82D" box="[1157,1287,1944,1970]" pageId="23" pageNumber="24" value="1979-06-19">
<collectingDate id="C6756C534B53FF9FE3AAF835FACCF82D" box="[1157,1287,1944,1970]" pageId="23" pageNumber="24" value="1979-06-19">19.06.1979</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF9FE23CF835FA4BF848" ID-GBIF-Occurrence="4104078304" collectingDate="2005-01-22" collectionCode="ZRC" collectorName="N. K. Ng &amp; J. C. Y. Lai" country="Philippines" pageId="23" pageNumber="24" specimenCode="ZRC 2014.0725" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B53FF9FE23CF835FABDF82D" box="[1299,1398,1944,1970]" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B53FF9FE7B8F811FEC8F849" box="[151,259,1980,2006]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="23" pageNumber="24" unit="mm" value="16.9">16.9 mm</quantity>
), 
<specimenCode id="F2291B004B53FF9FE636F811FE1FF848" box="[281,468,1980,2007]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="23" pageNumber="24">ZRC 2014.0725</specimenCode>
, 
<collectorName id="0F7AD6AD4B53FF9FE6CDF810FDE3F848" box="[482,552,1981,2007]" pageId="23" pageNumber="24">Panay</collectorName>
, Ilo Ilo, market near fishing port, leg. 
<collectorName id="0F7AD6AD4B53FF9FE4C3F810FB84F848" box="[1004,1103,1981,2007]" pageId="23" pageNumber="24">N.K. Ng</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B53FF9FE355F811FB24F849" box="[1146,1263,1980,2006]" pageId="23" pageNumber="24">J.C.Y. Lai</collectorName>
, 
<date id="D63195BB4B53FF9FE3D1F811FA4BF848" box="[1278,1408,1980,2007]" pageId="23" pageNumber="24" value="2005-01-22">
<collectingDate id="C6756C534B53FF9FE3D1F811FA4BF848" box="[1278,1408,1980,2007]" pageId="23" pageNumber="24" value="2005-01-22">22.01.2005</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B53FF90E2A1F811FED2FF2E" ID-GBIF-Occurrence="4104078571" collectingDate="1908-12-26" collectionCode="USNM" country="Philippines" lastPageId="24" lastPageNumber="25" location="Malampaya River" pageId="23" pageNumber="24" specimenCode="USNM 252028" specimenCount="1" specimenCount-female="1" stateProvince="Palawan">
<specimenCount id="B48978F24B53FF9FE2A1F811FF2DF865" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
? (cl 
<quantity id="65771E9E4B53FF9FE60FF84DFE43F864" box="[288,392,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="23" pageNumber="24" unit="mm" value="21.4">21.4 mm</quantity>
, pleopods damaged, missing minor cheliped), 
<specimenCode id="F2291B004B53FF9FE4B1F84DFB84F864" box="[926,1103,2016,2043]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24" type="Museum">USNM 252028</specimenCode>
, 
<collectingRegion id="604B7D994B53FF9FE373F84DFB75F865" box="[1116,1214,2016,2042]" country="Philippines" name="Palawan" pageId="23" pageNumber="24">Palawan</collectingRegion>
, 
<location id="A750E5A04B53FF9FE3E4F84DFA53F865" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B53FF9FE3E4F84DFA53F865" box="[1227,1432,2016,2042]" country="Philippines" name="Malampaya River" pageId="23" pageNumber="24" stateProvince="Palawan">Malampaya River</location>
, 
<date id="D63195BB4B5CFF90E7B8FF3AFED2FF2E" box="[151,281,151,177]" pageId="24" pageNumber="25" value="1908-12-26">
<collectingDate id="C6756C534B5CFF90E7B8FF3AFED2FF2E" box="[151,281,151,177]" pageId="24" pageNumber="25" value="1908-12-26">26.12.1908</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E609FF3AFE94FF4B" ID-GBIF-Occurrence="4104078386" collectingDate="1927-02-09" collectionCode="USNM" country="Philippines" county="Luzon" location="Malolos" municipality="Bay" pageId="24" pageNumber="25" specimenCode="USNM 1199121" specimenCount="1" specimenCount-male="1" stateProvince="Manila">
<specimenCount id="B48978F24B5CFF90E609FF3AFEBFFF2E" box="[294,372,151,177]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E68AFF3AFDC4FF2E" box="[421,527,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="24" pageNumber="25" unit="mm" value="18.4">18.4 mm</quantity>
), 
<specimenCode id="F2291B004B5CFF90E50BFF3AFD28FF2E" box="[548,739,151,177]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">USNM 1199121</specimenCode>
, 
<collectingCounty id="4B51CBF74B5CFF90E5DFFF3AFCF1FF2E" box="[752,826,151,177]" pageId="24" pageNumber="25">Luzon</collectingCounty>
, 
<collectingRegion id="604B7D994B5CFF90E467FF3AFC5CFF2E" box="[840,919,151,177]" country="Philippines" name="Manila" pageId="24" pageNumber="25">Manila</collectingRegion>
<collectingMunicipality id="425429014B5CFF90E48EFF3AFC04FF2E" box="[929,975,151,177]" pageId="24" pageNumber="25">Bay</collectingMunicipality>
, 
<location id="A750E5A04B5CFF90E4F2FF3AFBF7FF2E" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E4F2FF3AFBF7FF2E" box="[989,1084,151,177]" country="Philippines" county="Luzon" municipality="Bay" name="Malolos" pageId="24" pageNumber="25" stateProvince="Manila">Malolos</location>
, 
<location id="A750E5A04B5CFF90E366FF3AFB33FF2E" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E366FF3AFB33FF2E" box="[1097,1272,151,177]" country="Philippines" county="Luzon" municipality="Bay" name="Guzman Island" pageId="24" pageNumber="25" stateProvince="Manila">Guzman Island</location>
, Bulacan, fish pond, 
<date id="D63195BB4B5CFF90E7F2FF17FE94FF4B" box="[221,351,186,212]" pageId="24" pageNumber="25" value="1927-02-09">
<collectingDate id="C6756C534B5CFF90E7F2FF17FE94FF4B" box="[221,351,186,212]" pageId="24" pageNumber="25" value="1927-02-09">02.09.1927</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FF73FAC1FE6D" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<materialsCitation id="12E7B9264B5CFF90E7E8FF73FED2FE83" ID-GBIF-Occurrence="4104078550" collectingDate="2013-08-17" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" location="Lake" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Songkhla">
<emphasis id="90FB6F694B5CFF90E7E8FF73FEE7FF67" box="[199,300,222,248]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FF73FEE7FF67" box="[199,300,222,248]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B5CFF90E610FF73FE45FF67" box="[319,398,222,248]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E6EEFF73FDE6FF67" box="[449,557,222,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="24" pageNumber="25" unit="mm" value="20.3">20.3 mm</quantity>
), 
<collectionCode id="C49E2BBE4B5CFF90E56CFF73FD78FF67" box="[579,691,222,248]" pageId="24" pageNumber="25">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B5CFF90E599FF73FD10FF67" box="[694,731,222,248]" country="Austria" name="Zoological Collection, University of Vienna" pageId="24" pageNumber="25" type="University or college">ZC</collectionCode>
. 2019.06.46, 
<collectingRegion id="604B7D994B5CFF90E455FF73FC2FFF67" box="[890,996,222,248]" country="Thailand" name="Songkhla" pageId="24" pageNumber="25">Songkhla</collectingRegion>
<location id="A750E5A04B5CFF90E4C1FF73FBEDFF67" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E4C1FF73FBEDFF67" box="[1006,1062,222,248]" country="Thailand" name="Lake" pageId="24" pageNumber="25" stateProvince="Songkhla">Lake</location>
(Thale Sap), leg. 
<collectorName id="0F7AD6AD4B5CFF90E3D1FF73FA53FF67" box="[1278,1432,222,248]" pageId="24" pageNumber="25">S. Hajisamae</collectorName>
, 
<date id="D63195BB4B5CFF90E7B8FEAFFED2FE83" box="[151,281,258,284]" pageId="24" pageNumber="25" value="2013-08-17">
<collectingDate id="C6756C534B5CFF90E7B8FEAFFED2FE83" box="[151,281,258,284]" pageId="24" pageNumber="25" value="2013-08-17">17.08.2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E60BFEAFFEA7FEA0" ID-GBIF-Occurrence="4104078376" collectingDate="2013-08-17" collectionCode="ZRC" collectorName="S. Hajisamae" country="Thailand" location="Lake" pageId="24" pageNumber="25" specimenCode="ZRC 2014.0675" specimenCount="2" specimenCount-female="2" stateProvince="Songkhla">
<specimenCount id="B48978F24B5CFF90E60BFEAFFEB1FE83" box="[292,378,258,284]" pageId="24" pageNumber="25" type="male">2 males</specimenCount>
(cl 13.3, 
<quantity id="65771E9E4B5CFF90E6CBFEAFFD87FE83" box="[484,588,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="24" pageNumber="25" unit="mm" value="15.6">15.6 mm</quantity>
), 
<specimenCount id="B48978F24B5CFF90E570FEAFFD00FE83" box="[607,715,258,284]" pageId="24" pageNumber="25" type="female">2 females</specimenCount>
(cl 10.1, 
<quantity id="65771E9E4B5CFF90E41AFEAFFC56FE83" box="[821,925,258,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="24" pageNumber="25" unit="mm" value="13.2">13.2 mm</quantity>
), 
<specimenCode id="F2291B004B5CFF90E49FFEAFFBACFE82" box="[944,1127,258,285]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 2014.0675</specimenCode>
, same collection data as for previous specimen
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E658FE8BFA8DFEDF" ID-GBIF-Occurrence="4104078521" box="[375,1350,293,320]" collectionCode="ZRC" collectorName="S. Hajisamae" country="Thailand" location="Bay" pageId="24" pageNumber="25" specimenCode="ZRC 2014.0672" specimenCount="1" specimenCount-male="1" stateProvince="Pattani">
<specimenCount id="B48978F24B5CFF90E658FE8BFE09FEA0" box="[375,450,293,320]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E6C1FE88FD9DFEDF" box="[494,598,293,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="24" pageNumber="25" unit="mm" value="16.8">16.8 mm</quantity>
), 
<specimenCode id="F2291B004B5CFF90E546FE8BFCEBFEA0" box="[617,800,293,320]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 2014.0672</specimenCode>
, 
<collectingRegion id="604B7D994B5CFF90E404FE88FCB2FEA0" box="[811,889,293,319]" country="Thailand" name="Pattani" pageId="24" pageNumber="25">Pattani</collectingRegion>
<location id="A750E5A04B5CFF90E4AFFE8BFC65FEDF" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E4AFFE8BFC65FEDF" box="[896,942,294,320]" country="Thailand" name="Bay" pageId="24" pageNumber="25" stateProvince="Pattani">Bay</location>
, Pattani, leg. 
<collectorName id="0F7AD6AD4B5CFF90E36AFE8BFB17FEA0" box="[1093,1244,293,320]" pageId="24" pageNumber="25">S. Hajisamae</collectorName>
, 01.2013
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E27EFE8BFB4AFEFC" ID-GBIF-Occurrence="4104078333" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" location="Bay" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Pattani">
<specimenCount id="B48978F24B5CFF90E27EFE8BFA57FEA0" box="[1361,1436,293,320]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 16.0 mm), 
<collectionCode id="C49E2BBE4B5CFF90E612FEE4FE66FEFC" box="[317,429,329,355]" pageId="24" pageNumber="25">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B5CFF90E680FEE4FE1FFEFC" box="[431,468,329,355]" country="Austria" name="Zoological Collection, University of Vienna" pageId="24" pageNumber="25" type="University or college">ZC</collectionCode>
. 2019.06.47, same collection data as for previous specimen
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E3A0FEE4FCB8FE18" ID-GBIF-Occurrence="4104078359" collectingDate="1923-10-30" collectionCode="USNM" collectorName="H. M. Smith" country="Thailand" location="Tachin River" pageId="24" pageNumber="25" specimenCode="USNM 65478" specimenCount="4" specimenCount-female="4">
<specimenCount id="B48978F24B5CFF90E3A0FEE4FB36FEFC" box="[1167,1277,329,355]" pageId="24" pageNumber="25" type="female">4 females</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E203FEE4FF0CFE18" metricMagnitude="-2" metricUnit="m" metricValue="1.88" metricValueMax="2.08" metricValueMin="1.68" pageId="24" pageNumber="25" unit="mm" value="18.8" valueMax="20.8" valueMin="16.8">16.8–20.8 mm</quantity>
), 
<specimenCode id="F2291B004B5CFF90E7F4FEC0FEB5FE17" box="[219,382,365,392]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">USNM 65478</specimenCode>
, 
<location id="A750E5A04B5CFF90E6A6FEC0FDD6FE18" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E6A6FEC0FDD6FE18" box="[393,541,365,391]" country="Thailand" name="Tachin River" pageId="24" pageNumber="25">Tachin River</location>
, leg. 
<collectorName id="0F7AD6AD4B5CFF90E574FEC3FD2EFE18" box="[603,741,365,392]" pageId="24" pageNumber="25">H.M. Smith</collectorName>
, 
<date id="D63195BB4B5CFF90E5DEFEC0FCB8FE18" box="[753,883,365,391]" pageId="24" pageNumber="25" value="1923-10-30">
<collectingDate id="C6756C534B5CFF90E5DEFEC0FCB8FE18" box="[753,883,365,391]" pageId="24" pageNumber="25" value="1923-10-30">30.10.1923</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E450FEC0FC9BFE33" ID-GBIF-Occurrence="4104078482" collectingDate="1925-07-21" collectionCode="USNM" collectorName="H. Smith" country="Thailand" location="Menam Chao Phraya" municipality="Paknam" pageId="24" pageNumber="25" specimenCode="USNM 65557" specimenCount="1" specimenCount-female="1" stateProvince="Krung Thep Mahanakhon">
<specimenCount id="B48978F24B5CFF90E450FEC0FC2AFE18" box="[895,993,365,391]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 20.0 mm), 
<specimenCode id="F2291B004B5CFF90E3A2FEC0FAE4FE17" box="[1165,1327,365,392]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">USNM 65557</specimenCode>
, 
<collectingMunicipality id="425429014B5CFF90E214FEC0FA53FE18" box="[1339,1432,365,391]" pageId="24" pageNumber="25">Paknam</collectingMunicipality>
, 
<location id="A750E5A04B5CFF90E7B8FE3CFE4AFE34" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E7B8FE3CFE4AFE34" box="[151,385,401,427]" country="Thailand" municipality="Paknam" name="Menam Chao Phraya" pageId="24" pageNumber="25" stateProvince="Krung Thep Mahanakhon">Menam Chao Phraya</location>
near 
<collectingRegion id="604B7D994B5CFF90E692FE3CFDEEFE34" box="[445,549,401,427]" country="Thailand" name="Krung Thep Mahanakhon" pageId="24" pageNumber="25">Bangkok</collectingRegion>
, leg. 
<collectorName id="0F7AD6AD4B5CFF90E571FE3CFD0EFE34" box="[606,709,401,427]" pageId="24" pageNumber="25">H. Smith</collectorName>
, 
<date id="D63195BB4B5CFF90E5E1FE3CFC9BFE33" box="[718,848,401,428]" pageId="24" pageNumber="25" value="1925-07-21">
<collectingDate id="C6756C534B5CFF90E5E1FE3CFC9BFE33" box="[718,848,401,428]" pageId="24" pageNumber="25" value="1925-07-21">21.07.1925</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E475FE3CFCC2FE50" ID-GBIF-Occurrence="4104078570" country="Thailand" determinerName="D. M. Banner" location="Phuket" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Phuket">
<specimenCount id="B48978F24B5CFF90E475FE3CFC71FE34" box="[858,954,401,427]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 22.0 mm), NHM 1976.559, 
<collectingRegion id="604B7D994B5CFF90E237FE3CFAA2FE34" box="[1304,1385,401,427]" country="Thailand" name="Phuket" pageId="24" pageNumber="25">Phuket</collectingRegion>
, leg. N. Ngoc-Ho, 1973 [det. 
<determinerName id="57EC5FBF4B5CFF90E687FE18FDF0FE50" box="[424,571,437,463]" pageId="24" pageNumber="25">D.M. Banner</determinerName>
as 
<taxonomicName id="658FC8F84B5CFF90E54FFE18FD34FE51" authorityName="sensu De Man" authorityYear="1897" box="[608,767,436,463]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5CFF90E54FFE18FD34FE51" box="[608,767,436,463]" italics="true" pageId="24" pageNumber="25">A. euphrosyne</emphasis>
</taxonomicName>
]
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E438FE18FACEFE6D" ID-GBIF-Occurrence="4104078542" country="Thailand" determinerName="A. Anker" location="Phuket" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="2" stateProvince="Phuket">
<specimenCount id="B48978F24B5CFF90E438FE18FCA6FE51" box="[791,877,436,463]" pageId="24" pageNumber="25" type="male">2 males</specimenCount>
(cl 21.6, 24.0 mm), 
<specimenCount id="B48978F24B5CFF90E37DFE18FB75FE51" box="[1106,1214,436,463]" pageId="24" pageNumber="25" type="female">2 females</specimenCount>
(cl 22.5, 
<quantity id="65771E9E4B5CFF90E207FE19FA5BFE50" box="[1320,1424,436,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="24" pageNumber="25" unit="mm" value="24.5">24.5 mm</quantity>
), NHM 1976.560, 
<collectingRegion id="604B7D994B5CFF90E678FE75FE63FE6D" box="[343,424,472,498]" country="Thailand" name="Phuket" pageId="24" pageNumber="25">Phuket</collectingRegion>
, leg. N. Ngoc-Ho, no date (possibly 1973) [det. 
<determinerName id="57EC5FBF4B5CFF90E4EDFE75FBE3FE6D" box="[962,1064,472,498]" pageId="24" pageNumber="25">A. Anker</determinerName>
as 
<taxonomicName id="658FC8F84B5CFF90E361FE74FB26FE6D" authorityName="sensu De Man" authorityYear="1897" box="[1102,1261,472,499]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5CFF90E361FE74FB26FE6D" box="[1102,1261,472,499]" italics="true" pageId="24" pageNumber="25">A. euphrosyne</emphasis>
</taxonomicName>
?]
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FE51FE57FDC2" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<emphasis id="90FB6F694B5CFF90E7E8FE51FEF7FD89" box="[199,316,508,534]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FE51FEF7FD89" box="[199,316,508,534]" name="Cambodia" pageId="24" pageNumber="25">Cambodia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B5CFF90E667FE51FE5BFD89" box="[328,400,508,534]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E698FE51FDD6FD89" box="[439,541,508,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="24" pageNumber="25" unit="mm" value="18.4">18.4 mm</quantity>
), 3 ov. females (cl 16.0– 
<quantity id="65771E9E4B5CFF90E40EFE51FC4CFD89" box="[801,903,508,534]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="24" pageNumber="25" unit="mm" value="20.7">20.7 mm</quantity>
), MNHN-IU-2014-10287, locality not specified, “Sac 200 / 7C”, leg. M. Fily; 
<specimenCount id="B48978F24B5CFF90E6CCFD8DFDF6FDA5" box="[483,573,544,570]" pageId="24" pageNumber="25" type="male">2 males</specimenCount>
, 3 ov. females (cl indet.), MNHN-IU-2018-5727, locality not specified, “Sac 200 / 7B”, leg. M. Fily.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FDC5FCA2FC8E" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<materialsCitation id="12E7B9264B5CFF90E7E8FDC5FEDEFD39" ID-GBIF-Occurrence="4104078354" collectingDate="1995-12-09" collectionCode="OUMNH, ZC" collectorName="K. Wada" country="Vietnam" location="Cam River" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Hai Phong">
<emphasis id="90FB6F694B5CFF90E7E8FDC5FEE9FD1E" box="[199,290,616,641]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FDC5FEE9FD1E" box="[199,290,616,641]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B5CFF90E61EFDCAFE59FD1E" box="[305,402,615,641]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E691FDCAFDEEFD1E" box="[446,549,615,641]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="24" pageNumber="25" unit="mm" value="11.9">11.9 mm</quantity>
), 
<collectionCode id="C49E2BBE4B5CFF90E517FDCAFD63FD1E" box="[568,680,615,641]" pageId="24" pageNumber="25">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B5CFF90E584FDCAFD1BFD1E" box="[683,720,615,641]" country="Austria" name="Zoological Collection, University of Vienna" pageId="24" pageNumber="25" type="University or college">ZC</collectionCode>
. 2019.06.48, 
<collectingRegion id="604B7D994B5CFF90E448FDCAFC13FD1E" box="[871,984,615,641]" country="Vietnam" name="Hai Phong" pageId="24" pageNumber="25">Haiphong</collectingRegion>
, 
<location id="A750E5A04B5CFF90E4CBFDCAFB94FD1E" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E4CBFDCAFB94FD1E" box="[996,1119,615,641]" country="Vietnam" name="Cam River" pageId="24" pageNumber="25" stateProvince="Hai Phong">Cam River</location>
, back swamp, leg. 
<collectorName id="0F7AD6AD4B5CFF90E21BFDC5FA53FD1E" box="[1332,1432,615,642]" pageId="24" pageNumber="25">K. Wada</collectorName>
, 
<date id="D63195BB4B5CFF90E7B8FD26FEDEFD39" box="[151,277,651,678]" pageId="24" pageNumber="25" value="1995-12-09">
<collectingDate id="C6756C534B5CFF90E7B8FD26FEDEFD39" box="[151,277,651,678]" pageId="24" pageNumber="25" value="1995-12-09">12.09.1995</collectingDate>
</date>
</materialsCitation>
[YMP-1557]; 
<materialsCitation id="12E7B9264B5CFF90E692FD26FE43FD56" ID-GBIF-Occurrence="4104078535" collectingDate="1995-12-09" collectorName="K. Wada" country="Vietnam" location="Cam River" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Hai Phong">
<specimenCount id="B48978F24B5CFF90E692FD26FDD5FD3A" box="[445,542,651,677]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E564FD26FD78FD3A" box="[587,691,651,677]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="24" pageNumber="25" unit="mm" value="17.8">17.8 mm</quantity>
), MNHN-IU-2018-5737, 
<collectingRegion id="604B7D994B5CFF90E4FBFD26FB8EFD3A" box="[980,1093,651,677]" country="Vietnam" name="Hai Phong" pageId="24" pageNumber="25">Haiphong</collectingRegion>
, 
<location id="A750E5A04B5CFF90E37EFD26FB06FD3A" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E37EFD26FB06FD3A" box="[1105,1229,651,677]" country="Vietnam" name="Cam River" pageId="24" pageNumber="25" stateProvince="Hai Phong">Cam River</location>
, back swamp, leg. 
<collectorName id="0F7AD6AD4B5CFF90E7B8FD02FF36FD56" box="[151,253,687,713]" pageId="24" pageNumber="25">K. Wada</collectorName>
, 
<date id="D63195BB4B5CFF90E625FD03FE43FD56" box="[266,392,686,713]" pageId="24" pageNumber="25" value="1995-12-09">
<collectingDate id="C6756C534B5CFF90E625FD03FE43FD56" box="[266,392,686,713]" pageId="24" pageNumber="25" value="1995-12-09">12.09.1995</collectingDate>
</date>
</materialsCitation>
[YMP-1558]; 
<materialsCitation id="12E7B9264B5CFF90E51CFD02FC4FFD73" ID-GBIF-Occurrence="4104078339" collectorName="H. Muhllk &amp; J. Bourgeois" country="Vietnam" location="Tonkin" pageId="24" pageNumber="25" specimenCount="3" specimenCount-male="3">
<specimenCount id="B48978F24B5CFF90E51CFD02FD40FD56" box="[563,651,687,713]" pageId="24" pageNumber="25" type="male">3 males</specimenCount>
(cl 13.0– 
<quantity id="65771E9E4B5CFF90E5D4FD02FCA8FD56" box="[763,867,687,713]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="24" pageNumber="25" unit="mm" value="14.2">14.2 mm</quantity>
, pinned dried specimens), MNHN-IU-2000-1124, 
<collectingCountry id="DA98F3EB4B5CFF90E7B8FD7FFF22FD73" box="[151,233,722,748]" name="Vietnam" pageId="24" pageNumber="25">Tonkin</collectingCountry>
, locality not specified, leg. 
<collectorName id="0F7AD6AD4B5CFF90E507FD7EFD54FD73" box="[552,671,722,749]" pageId="24" pageNumber="25">H. Muhllk</collectorName>
/ coll. 
<collectorName id="0F7AD6AD4B5CFF90E5DBFD7EFC4FFD73" box="[756,900,722,749]" pageId="24" pageNumber="25">J. Bourgeois</collectorName>
</materialsCitation>
, 
<materialsCitation id="12E7B9264B5CFF90E4BEFD7FFCAEFC8E" ID-GBIF-Occurrence="4104078453" collectingDate="1995-08-21" collectorName="H. Duy" country="Tonkin" elevation="25" location="Sau" municipality="Ha Tien" pageId="24" pageNumber="25" specimenCount="1">
1911; 1 ov. female (cl 
<quantity id="65771E9E4B5CFF90E3B8FD7FFACAFD73" box="[1175,1281,722,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="24" pageNumber="25" unit="mm" value="18.7">18.7 mm</quantity>
), MNHN-IU-2018-5729, 
<collectingMunicipality id="425429014B5CFF90E630FD5AFEB0FC8F" box="[287,379,758,785]" pageId="24" pageNumber="25">Ha Tien</collectingMunicipality>
, 
<location id="A750E5A04B5CFF90E6A9FD5BFE78FC8F" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E6A9FD5BFE78FC8F" box="[390,435,758,784]" country="Tonkin" municipality="Ha Tien" name="Sau" pageId="24" pageNumber="25">Sau</location>
, 
<quantity id="65771E9E4B5CFF90E691FD5BFDEAFC8E" box="[446,545,758,785]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
<elevation id="29A254484B5CFF90E691FD5BFDEAFC8E" box="[446,545,758,785]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20–30 m</elevation>
</quantity>
(?), leg. 
<collectorName id="0F7AD6AD4B5CFF90E5AAFD5AFD13FC8E" box="[645,728,759,785]" pageId="24" pageNumber="25">H. Duy</collectorName>
, 
<date id="D63195BB4B5CFF90E5CBFD5BFCAEFC8E" box="[740,869,758,785]" pageId="24" pageNumber="25" value="1995-08-21">
<collectingDate id="C6756C534B5CFF90E5CBFD5BFCAEFC8E" box="[740,869,758,785]" pageId="24" pageNumber="25" value="1995-08-21">21.08.1995</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FCB7FEC6FCC8" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<materialsCitation id="12E7B9264B5CFF90E7E8FCB7FEC2FCC8" ID-GBIF-Occurrence="4104078326" collectionCode="USNM" collectorName="S. F. Light" country="China" location="China" pageId="24" pageNumber="25" specimenCode="USNM 62108" specimenCount="2" specimenCount-female="2">
<emphasis id="90FB6F694B5CFF90E7E8FCB7FEC0FCAB" box="[199,267,794,820]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FCB7FEC0FCAB" box="[199,267,794,820]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B5CFF90E633FCB7FE42FCAB" box="[284,393,794,820]" pageId="24" pageNumber="25" type="female">2 females</specimenCount>
(cl 18.0, mm, 
<quantity id="65771E9E4B5CFF90E51CFCB7FD57FCAB" box="[563,668,794,821]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="24" pageNumber="25" unit="mm" value="18.5">18.5 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B5CFF90E444FCB7FC1FFCAB" box="[875,980,794,820]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="24" pageNumber="25" unit="mm" value="13.4">13.4 mm</quantity>
), 
<specimenCode id="F2291B004B5CFF90E4C6FCB7FB40FCAB" box="[1001,1163,794,820]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">USNM 62108</specimenCode>
, locality unknown, don. 
<collectorName id="0F7AD6AD4B5CFF90E7B8FC93FEC2FCC8" box="[151,265,829,856]" pageId="24" pageNumber="25">S.F. Light</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FCCCFAAEFC00" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<materialsCitation id="12E7B9264B5CFF90E7E8FCCCFAAAFC00" ID-GBIF-Occurrence="4104078301" collectingDate="1963-02-11" collectionCode="AM" collectorName="J. C. Yaldwyn" country="Australia" latitude="-16.916666" location="Barron River" longLatPrecision="1280" longitude="145.76666" municipality="Cairns" pageId="24" pageNumber="25" specimenCode="P 28130" specimenCount="1" stateProvince="Queensland">
<emphasis id="90FB6F694B5CFF90E7E8FCCCFEE4FCE4" box="[199,303,865,891]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FCCCFEE4FCE4" box="[199,303,865,891]" name="Australia" pageId="24" pageNumber="25">Australia</collectingCountry>
</emphasis>
: 1 ov. female (cl 
<quantity id="65771E9E4B5CFF90E6DDFCCCFD9CFCE3" box="[498,599,865,892]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="24" pageNumber="25" unit="mm" value="16.2">16.2 mm</quantity>
, missing minor cheliped), 
<collectionCode id="C49E2BBE4B5CFF90E451FCCFFC60FCE3" box="[894,939,866,892]" country="Australia" name="Australian Museum" pageId="24" pageNumber="25" type="Museum">AM</collectionCode>
<specimenCode id="F2291B004B5CFF90E49DFCCFFBDBFCE4" box="[946,1040,865,892]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="24" pageNumber="25" type="Museum">P 28130</specimenCode>
, 
<collectingRegion id="604B7D994B5CFF90E334FCCCFB69FCE4" box="[1051,1186,865,891]" country="Australia" name="Queensland" pageId="24" pageNumber="25">Queensland</collectingRegion>
, 
<collectingMunicipality id="425429014B5CFF90E382FCCCFB32FCE4" box="[1197,1273,865,891]" pageId="24" pageNumber="25">Cairns</collectingMunicipality>
, 
<location id="A750E5A04B5CFF90E22BFCCFFA53FCE4" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E22BFCCFFA53FCE4" box="[1284,1432,865,892]" country="Australia" latitude="-16.916666" longLatPrecision="1280" longitude="145.76666" municipality="Cairns" name="Barron River" pageId="24" pageNumber="25" stateProvince="Queensland">Barron River</location>
, 
<geoCoordinate id="C7BBD5BC4B5CFF90E7B8FC28FF3FFC3F" box="[151,244,901,928]" degrees="16" direction="south" minutes="55" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-16.916666">16°55’S</geoCoordinate>
<geoCoordinate id="C7BBD5BC4B5CFF90E7D5FC28FEA2FC3F" box="[250,361,901,928]" degrees="145" direction="east" minutes="46" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="145.76666">145°46’E</geoCoordinate>
, among crab burrows (
<taxonomicName id="658FC8F84B5CFF90E545FC28FD38FC00" box="[618,755,901,927]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B5CFF90E545FC28FD01FC00" box="[618,714,901,927]" italics="true" pageId="24" pageNumber="25">Sesarma</emphasis>
sp.
</taxonomicName>
), deep within riverbank, leg. 
<collectorName id="0F7AD6AD4B5CFF90E312FC28FB1EFC00" box="[1085,1237,901,927]" pageId="24" pageNumber="25">J.C. Yaldwyn</collectorName>
, 
<date id="D63195BB4B5CFF90E3CFFC28FAAAFC00" box="[1248,1377,901,927]" pageId="24" pageNumber="25" value="1963-02-11">
<collectingDate id="C6756C534B5CFF90E3CFFC28FAAAFC00" box="[1248,1377,901,927]" pageId="24" pageNumber="25" value="1963-02-11">02.11.1963</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FC05FED6FBE9" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<emphasis id="90FB6F694B5CFF90E7E8FC05FE2AFC5D" bold="true" box="[199,481,936,962]" pageId="24" pageNumber="25">Tentative identification</emphasis>
. 
<taxonomicName id="658FC8F84B5CFF90E6DBFC04FCD9FC5C" authorityName="De Man" authorityYear="1920" box="[500,786,937,963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5CFF90E6DBFC04FD85FC5C" box="[500,590,937,963]" italics="true" pageId="24" pageNumber="25">Alpheus</emphasis>
cf. 
<emphasis id="90FB6F694B5CFF90E5ACFC04FCD9FC5C" box="[643,786,937,963]" italics="true" pageId="24" pageNumber="25">eurydactylus</emphasis>
</taxonomicName>
. 
<emphasis id="90FB6F694B5CFF90E40AFC04FC5CFC5C" box="[805,919,937,963]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E40AFC04FC5CFC5C" box="[805,919,937,963]" name="Singapore" pageId="24" pageNumber="25">Singapore</collectingCountry>
</emphasis>
: 
<materialsCitation id="12E7B9264B5CFF90E484FC04FAE6FC78" ID-GBIF-Occurrence="4104078401" collectingDate="1989-04-04" collectionCode="ZRC" collectorName="Lim Chu Kang" pageId="24" pageNumber="25" specimenCode="ZRC 2014.0724" specimenCount="1" specimenCount-male="1">
<specimenCount id="B48978F24B5CFF90E484FC04FC37FC5C" box="[939,1020,937,963]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E31DFC04FB56FC5C" box="[1074,1181,937,964]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="24" pageNumber="25" unit="mm" value="10.5">10.5 mm</quantity>
, tl 28.0 mm, rostrum damaged, missing minor cheliped), 
<specimenCode id="F2291B004B5CFF90E500FC60FD23FC78" box="[559,744,973,999]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 2014.0724</specimenCode>
, 
<collectorName id="0F7AD6AD4B5CFF90E5DAFC61FC69FC78" box="[757,930,972,999]" pageId="24" pageNumber="25">Lim Chu Kang</collectorName>
, mangrove, sta. Y595, 
<date id="D63195BB4B5CFF90E384FC60FAE6FC78" box="[1195,1325,973,999]" pageId="24" pageNumber="25" value="1989-04-04">
<collectingDate id="C6756C534B5CFF90E384FC60FAE6FC78" box="[1195,1325,973,999]" pageId="24" pageNumber="25" value="1989-04-04">04.04.1989</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E216FC60FACCFB95" ID-GBIF-Occurrence="4104078547" collectionCode="ZRC" collectorName="Jurong" pageId="24" pageNumber="25" specimenCode="ZRC 2014.0674" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B5CFF90E216FC60FA57FC79" box="[1337,1436,972,999]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E792FC5DFEEEFB94" box="[189,293,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="24" pageNumber="25" unit="mm" value="15.7">15.7 mm</quantity>
, missing both chelipeds), 
<specimenCode id="F2291B004B5CFF90E561FC5DFCCCFB95" box="[590,775,1008,1034]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 2014.0674</specimenCode>
, 
<collectorName id="0F7AD6AD4B5CFF90E43CFC5CFC94FB94" box="[787,863,1009,1035]" pageId="24" pageNumber="25">Jurong</collectorName>
prawn pond, no further data [J8048]
</materialsCitation>
; 
<materialsCitation id="12E7B9264B5CFF90E23BFC5DFB69FBB0" ID-GBIF-Occurrence="4104078379" collectingDate="1954-06-02" collectionCode="R" location="Jurong" pageId="24" pageNumber="25" specimenCode="ZRC 1979.4" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B5CFF90E23BFC5DFABDFB95" box="[1300,1398,1008,1034]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E7B8FBB9FF39FBB0" box="[151,242,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="24" pageNumber="25" unit="mm" value="9.6">9.6 mm</quantity>
, tl 26.0 mm), 
<specimenCode id="F2291B004B5CFF90E6B7FBB9FDE2FBB1" box="[408,553,1044,1070]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 1979.4</specimenCode>
.4.6, 
<location id="A750E5A04B5CFF90E570FBB8FD65FBB0" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E570FBB8FD65FBB0" box="[607,686,1045,1071]" name="Jurong" pageId="24" pageNumber="25">Jurong</location>
, prawn ponds, leg. S. 
<collectionCode id="C49E2BBE4B5CFF90E48AFBB9FC77FBB1" box="[933,956,1044,1070]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="25">R</collectionCode>
.F. 
<collectionCode id="C49E2BBE4B5CFF90E4F7FBB9FC24FBB1" box="[984,1007,1044,1070]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="25">R</collectionCode>
.S., 1– 
<date id="D63195BB4B5CFF90E302FBB9FB69FBB0" box="[1069,1186,1044,1071]" pageId="24" pageNumber="25" value="1954-06-02">
<collectingDate id="C6756C534B5CFF90E302FBB9FB69FBB0" box="[1069,1186,1044,1071]" pageId="24" pageNumber="25" value="1954-06-02">2.06.1954</collectingDate>
</date>
</materialsCitation>
[J8096]; 
<materialsCitation id="12E7B9264B5CFF90E23DFBB9FED3FBE9" ID-GBIF-Occurrence="4104078448" collectingDate="1991-03-04" collectionCode="ZRC" collectorName="P. K. L. Ng" location="Sungei Buloh" pageId="24" pageNumber="25" specimenCode="ZRC 1992.8064" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B5CFF90E23DFBB9FABEFBB1" box="[1298,1397,1044,1070]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B5CFF90E7B8FB9AFF35FBCD" box="[151,254,1079,1106]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="24" pageNumber="25" unit="mm" value="12.5">12.5 mm</quantity>
, tl 35.0 mm, missing major cheliped), 
<specimenCode id="F2291B004B5CFF90E59DFB95FCA1FBCD" box="[690,874,1080,1106]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="25">ZRC 1992.8064</specimenCode>
, 
<location id="A750E5A04B5CFF90E459FB95FBD9FBCD" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5CFF90E459FB95FBD9FBCD" box="[886,1042,1080,1106]" name="Sungei Buloh" pageId="24" pageNumber="25">Sungei Buloh</location>
, mangrove swamp, leg. 
<collectorName id="0F7AD6AD4B5CFF90E20AFB95FA53FBCD" box="[1317,1432,1080,1106]" pageId="24" pageNumber="25">P.K.L. Ng</collectorName>
, 
<date id="D63195BB4B5CFF90E7B8FBF1FED3FBE9" box="[151,280,1116,1142]" pageId="24" pageNumber="25" value="1991-03-04">
<collectingDate id="C6756C534B5CFF90E7B8FBF1FED3FBE9" box="[151,280,1116,1142]" pageId="24" pageNumber="25" value="1991-03-04">03.04.1991</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FBD2FE6CFB22" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<materialsCitation id="12E7B9264B5CFF90E7E8FBD2FCE2FB06" ID-GBIF-Occurrence="4104078492" box="[199,809,1151,1178]" collectionCode="RMNH" collectorName="Crus. D." country="India" location="India" pageId="24" pageNumber="25" specimenCount="1">
<emphasis id="90FB6F694B5CFF90E7E8FBD2FEC9FB06" box="[199,258,1151,1177]" italics="true" pageId="24" pageNumber="25">
<collectingCountry id="DA98F3EB4B5CFF90E7E8FBD2FEC9FB06" box="[199,258,1151,1177]" name="India" pageId="24" pageNumber="25">India</collectingCountry>
</emphasis>
: 1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B5CFF90E503FB2DFD4BFB05" box="[556,640,1152,1178]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25" type="Museum">RMNH</collectionCode>
<collectorName id="0F7AD6AD4B5CFF90E5A6FBD2FD16FB06" box="[649,733,1151,1177]" pageId="24" pageNumber="25">Crus.D.</collectorName>
49749
</materialsCitation>
, 
<materialsCitation id="12E7B9264B5CFF90E418FBD2FE69FB22" ID-GBIF-Occurrence="4104078378" collectingDate="1979-03-28" collectorName="J. C. Miquel" country="Kochi (Cochin)" determinerName="Y. Miya" location="Kochi (Cochin)" pageId="24" pageNumber="25" specimenCount="1">
<collectingCountry id="DA98F3EB4B5CFF90E418FBD2FC20FB06" box="[823,1003,1151,1177]" isEnumeration="true" pageId="24" pageNumber="25">Kochi (Cochin)</collectingCountry>
, leg. 
<collectorName id="0F7AD6AD4B5CFF90E304FBD2FB7EFB06" box="[1067,1205,1151,1177]" pageId="24" pageNumber="25">J.C. Miquel</collectorName>
, 
<date id="D63195BB4B5CFF90E3EDFBD2FA8BFB06" box="[1218,1344,1151,1177]" pageId="24" pageNumber="25" value="1979-03-28">
<collectingDate id="C6756C534B5CFF90E3EDFBD2FA8BFB06" box="[1218,1344,1151,1177]" pageId="24" pageNumber="25" value="1979-03-28">28.03.1979</collectingDate>
</date>
(det. 
<determinerName id="57EC5FBF4B5CFF90E2AAFB2DFF19FB22" pageId="24" pageNumber="25">Y. Miya</determinerName>
as 
<taxonomicName id="658FC8F84B5CFF90E7D7FB0EFE5CFB22" authorityName="sensu De Man" authorityYear="1897" box="[248,407,1187,1213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5CFF90E7D7FB0EFE5CFB22" box="[248,407,1187,1213]" italics="true" pageId="24" pageNumber="25">A. euphrosyne</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5CFF96E7E8FB6BFB70FC2E" lastPageId="30" lastPageNumber="31" pageId="24" pageNumber="25" type="description">
<paragraph id="A230B37B4B5CFF90E7E8FB6BFAF3FAF0" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">
<emphasis id="90FB6F694B5CFF90E7E8FB6BFEAEFB7F" bold="true" box="[199,357,1222,1248]" pageId="24" pageNumber="25">Redescription</emphasis>
. See 
<figureCitation id="3AB4AFFE4B5CFF90E688FB6AFDE1FB7E" box="[423,554,1222,1249]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStartId-0="25.[151,250,1892,1916]" captionStartId-1="26.[151,250,1819,1843]" captionStartId-2="27.[151,250,885,909]" captionStartId-3="28.[151,250,1839,1863]" captionStartId-4="29.[151,250,1821,1845]" captionTargetBox-0="[158,1429,181,1868]" captionTargetBox-1="[151,1435,181,1795]" captionTargetBox-2="[151,1435,181,861]" captionTargetBox-3="[174,1408,280,1804]" captionTargetBox-4="[167,1423,194,1791]" captionTargetId-0="figure-15@25.[158,1429,181,1868]" captionTargetId-1="figure-21@26.[151,1435,181,1795]" captionTargetId-2="figure-15@27.[151,1435,181,861]" captionTargetId-3="figure-62@28.[164,1423,273,1815]" captionTargetId-4="figure-15@29.[151,1436,181,1798]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="29" captionText-0="FIGURE 11. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—cephalothorax and chelipeds, dorsal view; B—frontal region, dorsal view; C, D— major (left) cheliped, mesial view, with slightly different angles and illuminations (slightly more ventral and enhancing mesial subdistal ridge on pollex in D). Refer to A for scale." captionText-1="FIGURE 12. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) [A–E] and paralectotype, ovigerous female (cl 15.0 mm) [F, G], both from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—major (left) cheliped, lateral view; B—same, chela, latreral view; C—same, fingers open, mesial view (see also Fig. 10C); D—minor chela in situ, dorsal view (cf. Fig. 3H); E—same, lateral view; F—frontal region, lateral view; G—frontal region and chelipeds in situ, dorsal view. Refer to Fig. 11A for scale in A–E." captionText-2="FIGURE 13. Alpheus eurydactylus De Man, 1920: female (cl 21.3 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49664): A—habitus, lateral view; B—minor (left) chela in situ, mesial view; C—major (right) cheliped in situ, mesial view; D—same, lateral view. Refer to A for scale." captionText-3="FIGURE 14. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–H, J–N]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [I]; A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—same, ischium, lateral view; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—same, basis and ischium, lateral view; M—second pleopod, appendices masculina and interna, anterior view; N—right uropod, dorsal view." captionText-4="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7912324" figureDoi-1="http://doi.org/10.5281/zenodo.7912328" figureDoi-2="http://doi.org/10.5281/zenodo.7912332" figureDoi-3="http://doi.org/10.5281/zenodo.7912334" figureDoi-4="http://doi.org/10.5281/zenodo.7912338" httpUri-0="https://zenodo.org/record/7912324/files/figure.png" httpUri-1="https://zenodo.org/record/7912328/files/figure.png" httpUri-2="https://zenodo.org/record/7912332/files/figure.png" httpUri-3="https://zenodo.org/record/7912334/files/figure.png" httpUri-4="https://zenodo.org/record/7912338/files/figure.png" pageId="24" pageNumber="25">Figs. 11–15</figureCitation>
. Large-sized species of 
<taxonomicName id="658FC8F84B5CFF90E419FB6AFC47FB7E" authorityName="Fabricius" authorityYear="1798" box="[822,908,1223,1249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B5CFF90E419FB6AFC47FB7E" box="[822,908,1223,1249]" italics="true" pageId="24" pageNumber="25">Alpheus</emphasis>
</taxonomicName>
(maximal cl 
<quantity id="65771E9E4B5CFF90E30CFB6AFB43FB7E" box="[1059,1160,1223,1249]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="24" pageNumber="25" unit="mm" value="20.3">20.3 mm</quantity>
, tl ~65.0 mm). Carapace glabrous, finely pitted, especially around cardiac region, without pubescence, with slight grooves. Rostrum short, subtriangular, narrowing towards apex, distally acute, typically falling short of mid-length of first article of antennular peduncle; rostral carina feebly developed, low, flattening posterior to orbital hoods; rostro-orbital furrows distinct, shallow; anterior margin of orbital hoods shallowly concave. Pterygostomial angle broadly rounded; cardiac notch deep.
</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8FAD4FE31F9F5" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">Pleon smooth, finely pitted, without pubescence. Telson broad, ovate-rectangular, slightly tapering towards posterior margin, about 1.3–1.4 times as long as maximal width near proximal margin; lateral margins very shallowly concave in proximal half almost straight in distal half; dorsal surface finely pitted, with shallow, longitudinal, median depression (without setae on edges) and two pairs of stout spiniform setae inserted in deep pits at some distance from lateral margins, first pair near telson mid-length, second pair at about 0.7 of telson length; posterior margin broadly rounded, with two pairs of small spiniform setae at posterolateral angles, lateral shorter than mesial (sometimes broken or missing).</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8F9DEFDEFF8DF" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">Antennular peduncle with stylocerite broad, moderately convex laterally, with acute tip, latter not overreaching distal margin of first article; ventromesial carina with anteriorly rounded tooth, without acute point; second article about 2.2–2.3 times as long as wide. Antennal peduncle with distoventral margin of basicerite unarmed; carpocerite not reaching beyond scaphocerite; scaphocerite with lateral margin straight to very shallowly concave; blade very broad, separated from broad distolateral tooth by deep cleft; anterior margin of blade broadly rounded and reaching distinctly beyond distolateral tooth.</paragraph>
<paragraph id="A230B37B4B5CFF90E7E8F8E7FE54F86C" blockId="24.[151,1437,151,2035]" pageId="24" pageNumber="25">Third maxilliped with antepenultimate article not particularly broadened; penultimate article about 2.5 times as long as wide, with long setae on ventrolateral margin, latter surpassing ultimate article; ultimate article with dense transverse rows of short serrulate setae on mesial surface and some longer setae on dorsal and lateral surfaces, especially near or on apex; coxal lateral plate bluntly produced dorsally; exopod fleshy, broad, reaching end of antepenultimate article.</paragraph>
<caption id="F6F0E3F34B5DFF91E7B8F8C9FD6EF876" ID-DOI="http://doi.org/10.5281/zenodo.7912324" ID-Zenodo-Dep="7912324" httpUri="https://zenodo.org/record/7912324/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1892,1916]" targetBox="[158,1429,181,1868]" targetPageId="25" targetType="figure">
<paragraph id="A230B37B4B5DFF91E7B8F8C9FD6EF876" blockId="25.[151,1436,1892,2025]" pageId="25" pageNumber="26">
<emphasis id="90FB6F694B5DFF91E7B8F8C9FED0F8E2" bold="true" box="[151,283,1892,1917]" pageId="25" pageNumber="26">FIGURE 11</emphasis>
. 
<taxonomicName id="658FC8F84B5DFF91E604F8C8FD66F8E2" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[299,685,1893,1917]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5DFF91E604F8C8FDCCF8E2" box="[299,519,1893,1917]" italics="true" pageId="25" pageNumber="26">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B5DFF91E53EF8C8FD66F8E2" author="De Man, J. G." box="[529,685,1893,1917]" pageId="25" pageNumber="26" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: lectotype, male (cl 13.8 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—cephalothorax and chelipeds, dorsal view; B—frontal region, dorsal view; C, D— major (left) cheliped, mesial view, with slightly different angles and illuminations (slightly more ventral and enhancing mesial subdistal ridge on pollex in D). Refer to A for scale.
</paragraph>
</caption>
<caption id="F6F0E3F34B5EFF92E7B8F8B6FDB1F85C" ID-DOI="http://doi.org/10.5281/zenodo.7912328" ID-Zenodo-Dep="7912328" httpUri="https://zenodo.org/record/7912328/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1819,1843]" targetBox="[151,1435,181,1795]" targetPageId="26" targetType="figure">
<paragraph id="A230B37B4B5EFF92E7B8F8B6FDB1F85C" blockId="26.[151,1436,1819,1987]" pageId="26" pageNumber="27">
<emphasis id="90FB6F694B5EFF92E7B8F8B6FED0F8AC" bold="true" box="[151,283,1819,1843]" pageId="26" pageNumber="27">FIGURE 12</emphasis>
. 
<taxonomicName id="658FC8F84B5EFF92E606F8B6FD6FF8AC" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[297,676,1819,1843]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5EFF92E606F8B6FDC8F8AC" box="[297,515,1819,1843]" italics="true" pageId="26" pageNumber="27">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B5EFF92E523F8B1FD6FF8AC" author="De Man, J. G." box="[524,676,1819,1843]" pageId="26" pageNumber="27" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: lectotype, male (cl 13.8 mm) [A–E] and paralectotype, ovigerous female (cl 15.0 mm) [F, G], both from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—major (left) cheliped, lateral view; B—same, chela, latreral view; C—same, fingers open, mesial view (see also Fig. 10C); D—minor chela 
<emphasis id="90FB6F694B5EFF92E7B8F825FF1EF800" box="[151,213,1928,1951]" italics="true" pageId="26" pageNumber="27">in situ</emphasis>
, dorsal view (cf. Fig. 3H); E—same, lateral view; F—frontal region, lateral view; G—frontal region and chelipeds 
<emphasis id="90FB6F694B5EFF92E277F825FA5DF800" box="[1368,1430,1928,1951]" italics="true" pageId="26" pageNumber="27">in situ</emphasis>
, dorsal view. Refer to Fig. 11A for scale in A–E.
</paragraph>
</caption>
<caption id="F6F0E3F34B5FFF93E7B8FCD8FE2EFC4A" ID-DOI="http://doi.org/10.5281/zenodo.7912332" ID-Zenodo-Dep="7912332" httpUri="https://zenodo.org/record/7912332/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,885,909]" targetBox="[151,1435,181,861]" targetPageId="27" targetType="figure">
<paragraph id="A230B37B4B5FFF93E7B8FCD8FE2EFC4A" blockId="27.[151,1437,885,982]" pageId="27" pageNumber="28">
<emphasis id="90FB6F694B5FFF93E7B8FCD8FEDCFC12" bold="true" box="[151,279,885,909]" pageId="27" pageNumber="28">FIGURE 13</emphasis>
. 
<taxonomicName id="658FC8F84B5FFF93E60CFCD8FD5AFC12" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[291,657,885,909]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5FFF93E60CFCD8FE31FC12" box="[291,506,885,909]" italics="true" pageId="27" pageNumber="28">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B5FFF93E6D0FCDBFD5AFC12" author="De Man, J. G." box="[511,657,885,909]" pageId="27" pageNumber="28" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: female (cl 21.3 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49664): A—habitus, lateral view; B—minor (left) chela 
<emphasis id="90FB6F694B5FFF93E5AFFC37FD74FC2E" box="[640,703,922,945]" italics="true" pageId="27" pageNumber="28">in situ</emphasis>
, mesial view; C—major (right) cheliped 
<emphasis id="90FB6F694B5FFF93E34CFC37FB69FC2E" box="[1123,1186,922,945]" italics="true" pageId="27" pageNumber="28">in situ</emphasis>
, mesial view; D—same, lateral view. Refer to A for scale.
</paragraph>
</caption>
<paragraph id="A230B37B4B5FFF93E7E8FBAEFE24F98A" blockId="27.[151,1437,1027,2026]" pageId="27" pageNumber="28">
Major cheliped of 
<taxonomicName id="658FC8F84B5FFF93E6B5FBA9FDECFB82" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[410,551,1027,1054]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="90FB6F694B5FFF93E6B5FBA9FDECFB82" box="[410,551,1027,1054]" italics="true" pageId="27" pageNumber="28">A. edwardsii</emphasis>
</taxonomicName>
- 
<typeStatus id="7D340DD94B5FFF93E51FFBA9FDAEFB81" box="[560,613,1028,1054]" pageId="27" pageNumber="28">type</typeStatus>
. Merus very stout, about 1.8 times as long as wide; ventromesial margin unarmed distally. Carpus very short, cup-shaped. Chela large, massive, with fingers about 0.6–0.7 length of palm. Mesial face of palm with low mesial transverse ridge in proximal half, extending from proximal edge of subtriangular mesial longitudinal groove to ventral surface of palm; surface smooth, largely without granules, except for very few near ventral shoulder. Lateral face of palm smooth, without granulation; lateral longitudinal groove deep, subrectangular; lateral half with low transverse ridge. Dorsal shoulder pronounced, rounded, sloping with angle of 45º–80º into broad transverse groove, not overhanging. Ventral shoulder pronounced, slightly protruding in lateral view, broadly rounded. Pollex with mesial face granulated over most of its surface, with prominent mesial subdistal ridge marked by conspicuous protuberance proximally and extending to subdistal angle of pollex; area ventral to mesial subdistal ridge conspicuously grooved; distomesial angle distinctly superior to 90º, blunt; lateral face smooth, not granulated; distolateral angle about 90º, broadly rounded. Dactylus slightly longer than pollex, with prominent dorsal ridge, latter twisted mesially and protruding proximally; mesial surface granulated in proximal half; lateral surface largely smooth; dactylar plunger large, stout, clearly distinct from ventral dactylar margin; distal surface of plunger with obliquely flattened ventromesial area, its mesial surface with feebly developed transverse channel. Adhesive discs small.
</paragraph>
<paragraph id="A230B37B4B5FFF93E7E8F9B2FABBF83E" blockId="27.[151,1437,1027,2026]" pageId="27" pageNumber="28">
<collectingRegion id="604B7D994B5FFF93E7E8F9B2FECAF9A6" box="[199,257,1567,1593]" country="Maldives" name="Maale" pageId="27" pageNumber="28">Male</collectingRegion>
minor cheliped with chela strongly balaeniceps. Merus somewhat slenderer than that of major cheliped, about 2.5 times as long as wide; ventral margin with distomesial angle unarmed. Chela moderately swollen, with fingers about as long as palm. Palm subcylindrical, somewhat swollen; mesial face with surface largely smooth, except for very small field of granules on subdistal swelling below mesial longitudinal groove; mesial longitudinal groove well marked, deep; lateral longitudinal groove well marked, deep; dorsal shoulder pronounced, rounded, sloping with 45º or smaller angle into deep dorsal transverse groove; ventral transverse groove moderately deep, bordered by distinct ventral shoulder on lateral surface. Pollex with largely smooth surfaces; mesial surface with row of balaeniceps setae on low crest extending from base to slightly beyond mid-length of pollex; lateral surface with similar row of balaeniceps setae as on mesial surface. Dactylus with well-developed proximal balaeniceps expansion, about 2.0–2.2 times as long as maximal width; lateral and mesial dactylar ridges each with dense rows of balaeniceps setae. Cutting edges of both fingers shallowly excavated on each side of sharp, blade-like ridge.
</paragraph>
<paragraph id="A230B37B4B5FFF94E7E8F806FBEEFF66" blockId="27.[151,1437,1027,2026]" lastBlockId="28.[151,1437,151,249]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Female minor cheliped simple, not balaeniceps. Chela slender, not particularly swollen, with fingers about 1.1–1.2 times length of palm. Palm surface smooth, without granulation (at most with some granules on distomesial surface), with shallow (sometimes traces of) mesial and lateral longitudinal grooves, and with shallow dorsal transverse groove and poorly demarcated ventral shoulder (sometimes not distinct). Fingers simple, with sharp cutting edges, not or slightly gaping when closed; fingertips curved and crossing.</paragraph>
<caption id="F6F0E3F34B58FF94E7B8F882FEACF868" ID-DOI="http://doi.org/10.5281/zenodo.7912334" ID-Zenodo-Dep="7912334" httpUri="https://zenodo.org/record/7912334/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1839,1863]" targetBox="[174,1408,280,1804]" targetPageId="28" targetType="figure">
<paragraph id="A230B37B4B58FF94E7B8F882FEACF868" blockId="28.[151,1436,1839,2039]" pageId="28" pageNumber="29">
<emphasis id="90FB6F694B58FF94E7B8F882FED1F8D8" bold="true" box="[151,282,1839,1863]" pageId="28" pageNumber="29">FIGURE 14</emphasis>
. 
<taxonomicName id="658FC8F84B58FF94E607F882FD6AF8D8" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[296,673,1839,1863]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B58FF94E607F882FDC9F8D8" box="[296,514,1839,1863]" italics="true" pageId="28" pageNumber="29">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B58FF94E525F882FD6AF8D8" author="De Man, J. G." box="[522,673,1839,1863]" pageId="28" pageNumber="29" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–H, J–N]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [I]; A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—same, ischium, lateral view; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—same, basis and ischium, lateral view; M—second pleopod, appendices masculina and interna, anterior view; N—right uropod, dorsal view.
</paragraph>
</caption>
<caption id="F6F0E3F34B59FF95E7B8F8B0FD28F859" ID-DOI="http://doi.org/10.5281/zenodo.7912338" ID-Zenodo-Dep="7912338" httpUri="https://zenodo.org/record/7912338/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1821,1845]" targetBox="[167,1423,194,1791]" targetPageId="29" targetType="figure">
<paragraph id="A230B37B4B59FF95E7B8F8B0FD28F859" blockId="29.[151,1437,1821,1990]" pageId="29" pageNumber="30">
<emphasis id="90FB6F694B59FF95E7B8F8B0FEDCF8AA" bold="true" box="[151,279,1821,1845]" pageId="29" pageNumber="30">FIGURE 15</emphasis>
. 
<taxonomicName id="658FC8F84B59FF95E60DF8B3FD44F8A9" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[290,655,1822,1846]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B59FF95E60DF8B3FE32F8A9" box="[290,505,1822,1846]" italics="true" pageId="29" pageNumber="30">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B59FF95E6D1F8B3FD44F8A9" author="De Man, J. G." box="[510,655,1822,1846]" pageId="29" pageNumber="30" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view.
</paragraph>
</caption>
<paragraph id="A230B37B4B5AFF96E7E8FF3AFE29FE74" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">Second pereiopod with ratio of carpal subarticles approximately equal to 4.2: 2.5: 1: 1: 1.5. Third pereiopod fairly robust; ischium armed with small, stout spiniform seta on ventrolateral surface near proximal margin, sometimes unarmed (especially in larger individuals); merus about 4.2 times as long as maximal width, unarmed; propodus with five or six stout spiniform setae on ventral margin and one pair of spiniform setae (sometimes one seta) on distoventral margin adjacent to dactylus; dactylus about 0.4 and sometimes close to half-length of propodus, spatulate, expanded, as wide as propodus in dorsal view, with row of tufts of setae along mesial ridge. Fourth pereiopod generally similar to third pereiopod, slightly slenderer; merus about five times as long as maximal width; dactylus about 0.4 length of propodus, spatulate. Fifth pereiopod much slenderer than third and fourth pereiopods; ischium unarmed; propodus with three small spiniform setae and well-developed setal brush; dactylus subspatulate, about 0.4 length of propodus.</paragraph>
<paragraph id="A230B37B4B5AFF96E7E8FE59FDD1FDE8" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">
<collectingRegion id="604B7D994B5AFF96E7E8FE59FECAFD91" box="[199,257,500,526]" country="Maldives" name="Maale" pageId="30" pageNumber="31">Male</collectingRegion>
second pleopod with appendix masculina ~0.8 length of appendix interna. Uropod with each protopodal lobe ending in sharp tooth distally; exopod and endopod broad, ovate; diaeresis of exopod complete, almost straight, with small subacute or blunt tooth near small distolateral spiniform seta and adjacent distolateral tooth; lateral margin of exopod broadly convex.
</paragraph>
<paragraph id="A230B37B4B5AFF96E7E8FD2DFBF2FD05" blockId="30.[151,1437,150,2030]" box="[199,1081,639,666]" pageId="30" pageNumber="31">
Eggs numerous (&gt; 
<quantity id="65771E9E4B5AFF96E6BCFD2DFE17FD05" box="[403,476,640,666]" metricMagnitude="0" metricUnit="m" metricValue="5.08" pageId="30" pageNumber="31" unit="in" value="200.0">200 in</quantity>
large females), small; egg diameter less than 
<quantity id="65771E9E4B5AFF96E4F4FDD2FBFFFD05" box="[987,1076,639,666]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="A230B37B4B5AFF96E7E8FD0FFB70FC2E" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">
<emphasis id="90FB6F694B5AFF96E7E8FD0FFEB6FD22" bold="true" box="[199,381,674,701]" pageId="30" pageNumber="31">Colour pattern</emphasis>
. Body uniform dark green, green-brown or brown, without marked banding on carapace or pleon; antennular flagella greenish, bluish proximally; antennal flagella olive-greenish with some bluish tinge proximally; mesial face of major chela olive or greenish brown with large pale orange area, especially near ventral transverse groove; fingers darker olive-green, distally purplish or pinkish; dactylar plunger with mesial side white and blue proximally and bright orange distally and anteriorly; minor chela olive-green with darker fingers; second pereiopods reddish proximally, bright blue distally; ambulatory legs reddish; pleopods pinkish; telson and uropods darker than pleon; uropods with dark, dusky blue patch distally (
<figureCitation id="3AB4AFFE4B5AFF96E454FCD9FC1BFC11" box="[891,976,884,910]" captionStart="FIGURE 16" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[152,1435,181,1885]" captionTargetId="figure-15@31.[152,1435,181,1885]" captionTargetPageId="31" captionText="FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D)." figureDoi="http://doi.org/10.5281/zenodo.7912340" httpUri="https://zenodo.org/record/7912340/files/figure.png" pageId="30" pageNumber="31">Fig. 16</figureCitation>
). One of the Singaporean females (ZRC 1996.47) was accompanied by a short colour note: “overall dark green, chela dactylus purple”.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5AFF96E7E8FC17FD53FC4B" box="[199,664,954,981]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="A230B37B4B5AFF96E7E8FC17FD53FC4B" blockId="30.[151,1437,150,2030]" box="[199,664,954,981]" pageId="30" pageNumber="31">
<materialsCitation id="12E7B9264B5AFF96E7E8FC17FD5EFC4B" ID-GBIF-Occurrence="4104078474" box="[199,661,954,981]" country="Indonesia" location="Surabaya" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="90FB6F694B5AFF96E7E8FC17FEABFC4B" bold="true" box="[199,352,954,980]" pageId="30" pageNumber="31">
<typeStatus id="7D340DD94B5AFF96E7E8FC17FEC9FC4B" box="[199,258,954,980]" pageId="30" pageNumber="31">Type</typeStatus>
locality
</emphasis>
. 
<location id="A750E5A04B5AFF96E641FC17FE12FC4B" LSID="urn:lsid:plazi:treatment:2A26026D4B53FFA9E7B8FEE7FAEAFF68:A750E5A04B5AFF96E641FC17FE12FC4B" box="[366,473,954,980]" country="Indonesia" name="Surabaya" pageId="30" pageNumber="31" stateProvince="Jawa Timur">Surabaya</location>
, 
<collectingRegion id="604B7D994B5AFF96E6CBFC16FDD3FC4A" box="[484,536,955,981]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="31">Java</collectingRegion>
, 
<collectingCountry id="DA98F3EB4B5AFF96E50BFC17FD5EFC4B" box="[548,661,954,980]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5AFF96E7E8FC70FD4CFB39" pageId="30" pageNumber="31" type="distribution">
<paragraph id="A230B37B4B5AFF96E7E8FC70FD4CFB39" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">
<emphasis id="90FB6F694B5AFF96E7E8FC70FE92FC68" bold="true" box="[199,345,989,1015]" pageId="30" pageNumber="31">Distribution</emphasis>
. Tropical Indo-West Pacific from the southern Andaman Sea and 
<collectingCountry id="DA98F3EB4B5AFF96E312FC70FB57FC68" box="[1085,1180,989,1015]" name="Malaysia" pageId="30" pageNumber="31">Malacca</collectingCountry>
Straits to South-China Sea and northern 
<collectingCountry id="DA98F3EB4B5AFF96E647FBADFE04FB85" box="[360,463,1024,1050]" name="Australia" pageId="30" pageNumber="31">Australia</collectingCountry>
(
<figureCitation id="3AB4AFFE4B5AFF96E6CCFBADFD86FB84" box="[483,589,1024,1051]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="30" pageNumber="31">Fig. 51B</figureCitation>
), including coasts of 
<collectingCountry id="DA98F3EB4B5AFF96E460FBADFC77FB85" box="[847,956,1024,1050]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E4E0FBADFBFAFB85" box="[975,1073,1024,1050]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">Sumatra</collectingRegion>
, 
<collectingRegion id="604B7D994B5AFF96E36EFBACFBBEFB84" box="[1089,1141,1025,1051]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="31">Java</collectingRegion>
, 
<collectingRegion id="604B7D994B5AFF96E3AAFBADFB05FB85" box="[1157,1230,1024,1050]" country="Indonesia" name="Papua Barat" pageId="30" pageNumber="31">Papua</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B5AFF96E3C9FBADFA93FB85" box="[1254,1368,1024,1050]" name="Singapore" pageId="30" pageNumber="31">Singapore</collectingCountry>
(East Coast, Mandai, Sungei Buloh, Punggol), 
<collectingCountry id="DA98F3EB4B5AFF96E543FB8EFD18FBA2" box="[620,723,1059,1085]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E5CCFB8EFCF1FBA2" box="[739,826,1059,1085]" country="Malaysia" name="Pulau Pinang" pageId="30" pageNumber="31">Penang</collectingRegion>
, 
<collectingRegion id="604B7D994B5AFF96E469FB8EFC60FBA2" box="[838,939,1059,1085]" country="Malaysia" name="Selangor" pageId="30" pageNumber="31">Selangor</collectingRegion>
, 
<collectingRegion id="604B7D994B5AFF96E497FB8EFC30FBA2" box="[952,1019,1059,1085]" country="Malaysia" name="Johor" pageId="30" pageNumber="31">Johor</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B5AFF96E320FB8EFBB8FBA2" box="[1039,1139,1059,1085]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E3ACFB8EFB24FBA2" box="[1155,1263,1059,1085]" country="Thailand" name="Songkhla" pageId="30" pageNumber="31">Songkhla</collectingRegion>
Lake, 
<collectingRegion id="604B7D994B5AFF96E211FB8EFA5BFBA2" box="[1342,1424,1059,1085]" country="Thailand" name="Pattani" pageId="30" pageNumber="31">Pattani</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B5AFF96E7B8FBEBFEC6FBFF" box="[151,269,1094,1120]" name="Cambodia" pageId="30" pageNumber="31">Cambodia</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B5AFF96E637FBEBFEBCFBFF" box="[280,375,1094,1120]" name="Vietnam" pageId="30" pageNumber="31">Vietnam</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E6AAFBEBFE33FBFF" box="[389,504,1094,1120]" country="Vietnam" name="Hai Phong" pageId="30" pageNumber="31">Haiphong</collectingRegion>
, Nha Trang, 
<collectingRegion id="604B7D994B5AFF96E5A6FBEBFC9CFBFF" box="[649,855,1094,1120]" country="Vietnam" name="Ho Chi Minh City" pageId="30" pageNumber="31">Ho Chi Minh City</collectingRegion>
, Can Gio), 
<collectingCountry id="DA98F3EB4B5AFF96E4F7FBEBFBD5FBFF" box="[984,1054,1094,1120]" name="China" pageId="30" pageNumber="31">China</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B5AFF96E306FBEBFB6CFBFF" box="[1065,1191,1094,1120]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E399FBEBFACCFBFF" box="[1206,1287,1094,1120]" country="Philippines" name="Manila" pageId="30" pageNumber="31">Manila</collectingRegion>
Bay, Panay), and 
<collectingCountry id="DA98F3EB4B5AFF96E7EAFBC4FEE7FB1C" box="[197,300,1129,1155]" name="Australia" pageId="30" pageNumber="31">Australia</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E615FBC4FE09FB1C" box="[314,450,1129,1155]" country="Australia" name="Queensland" pageId="30" pageNumber="31">Queensland</collectingRegion>
) (
<bibRefCitation id="C61ECE8A4B5AFF96E6F7FBC4FDBDFB1C" author="De Man, J. G." box="[472,630,1129,1155]" pageId="30" pageNumber="31" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man 1920</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B5AFF96E5AEFBC4FD77FB1C" author="De Man, J. G." box="[641,700,1129,1155]" pageId="30" pageNumber="31" pagination="1 - 60" refId="ref71119" refString="De Man, J. G. (1924) On a collection of macrurous decapod Crustacea chiefly Penaeidae and Alpheidae from the Indian Archipelago. Archiv fu ¨ r Naturgeschichte, Berlin, Series A, 90, 1 - 60." type="journal article" year="1924">1924</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B5AFF96E5E8FBC4FC85FB1C" author="Marin, I. N." box="[711,846,1129,1155]" pageId="30" pageNumber="31" pagination="369 - 385" refId="ref72908" refString="Marin, I. N. (2021) A new infaunal species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) and a new crustacean association from anoxic mangrove habitats in southern Vietnam. Arthropoda Selecta, 30, 369 - 385. https: // doi. org / 10.15298 / arthsel. 30.3.10" type="journal article" year="2021">Marin 2021</bibRefCitation>
; present study); possibly extending to southern 
<collectingCountry id="DA98F3EB4B5AFF96E24CFBC4FA56FB1C" box="[1379,1437,1129,1155]" name="India" pageId="30" pageNumber="31">India</collectingCountry>
(
<collectingCountry id="DA98F3EB4B5AFF96E7B0FB21FF23FB39" box="[159,232,1164,1190]" name="India" pageId="30" pageNumber="31">Kochi</collectingCountry>
, see discussion below and 
<figureCitation id="3AB4AFFE4B5AFF96E53BFB21FDB1FB39" box="[532,634,1163,1190]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="30" pageNumber="31">Fig. 51B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5AFF96E7E8FB03FC65FB56" box="[199,942,1198,1225]" pageId="30" pageNumber="31" type="vernacular_names">
<paragraph id="A230B37B4B5AFF96E7E8FB03FC65FB56" blockId="30.[151,1437,150,2030]" box="[199,942,1198,1225]" pageId="30" pageNumber="31">
<emphasis id="90FB6F694B5AFF96E7E8FB03FE38FB56" bold="true" box="[199,499,1198,1225]" pageId="30" pageNumber="31">Common name proposed</emphasis>
. De Man’s mangrove snapping shrimp.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5AFF96E7E8FB7FFB84F9D5" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="A230B37B4B5AFF96E7E8FB7FFEF7FA7E" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">
<emphasis id="90FB6F694B5AFF96E7E8FB7FFE0FFB74" bold="true" box="[199,452,1233,1260]" pageId="30" pageNumber="31">Ecology and biology</emphasis>
. 
<taxonomicName id="658FC8F84B5AFF96E6F8FB7CFD06FB74" authorityName="De Man" authorityYear="1920" box="[471,717,1233,1259]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E6F8FB7CFD06FB74" box="[471,717,1233,1259]" italics="true" pageId="30" pageNumber="31">Alpheus eurydactylus</emphasis>
</taxonomicName>
is a largely intertidal and shallow subtidal snapping shrimp (maximal depth probably not exceeding 
<quantity id="65771E9E4B5AFF96E575FB58FD4DFA90" box="[602,646,1269,1295]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="m" value="3.0">3 m</quantity>
) inhabiting estuarine, mangrove mudflats and brackish lagoons, where it lives in deep burrows dug in thick, soft mud, sometimes under large rocks or mangrove debris, among mangrove roots, or in the muddy banks of mangrove creeks. The unusual depth range (
<quantity id="65771E9E4B5AFF96E4D8FA91FBA2FAC9" box="[1015,1129,1340,1366]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="30" pageNumber="31" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20—30 m</quantity>
) given for the female from Ha Tien, 
<collectingCountry id="DA98F3EB4B5AFF96E62DFAF3FEAAFAE7" box="[258,353,1374,1400]" name="Vietnam" pageId="30" pageNumber="31">Vietnam</collectingCountry>
(MNHN-IU-2018-5729), is somewhat questionable. Like most euryhaline decapods present in these habitats, 
<taxonomicName id="658FC8F84B5AFF96E612FA2FFE20FA04" authorityName="De Man" authorityYear="1920" box="[317,491,1409,1436]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E612FA2FFE20FA04" box="[317,491,1409,1436]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
is a resilient shrimp capable of tolerating very low salinities. The ovigerous females of 
<taxonomicName id="658FC8F84B5AFF96E799FA08FEAFFA21" authorityName="De Man" authorityYear="1920" box="[182,356,1444,1471]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E799FA08FEAFFA21" box="[182,356,1444,1471]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
carrying a large number of eggs of relatively small size suggest an extended larval development in this species.
</paragraph>
<paragraph id="A230B37B4B5AFF96E7E8FA47FB84F9D5" blockId="30.[151,1437,150,2030]" pageId="30" pageNumber="31">
This large snapping shrimp is occasionally seen on fish markets and crab / shrimp landings, for instance, in the 
<collectingCountry id="DA98F3EB4B5AFF96E7EFF9A0FEF5F9B8" box="[192,318,1549,1575]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
(
<collectingRegion id="604B7D994B5AFF96E662F9A0FE68F9B8" box="[333,419,1549,1575]" country="Philippines" name="Manila" pageId="30" pageNumber="31">Manila</collectingRegion>
). Interestingly, one of the specimens from 
<collectingCountry id="DA98F3EB4B5AFF96E451F9A0FC29F9B8" box="[894,994,1549,1575]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
(USNM 65557) was accompanied by a note: “kung deet = snapping shrimp, not eaten but used as bait for catfish and drum”.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B5AFFA9E7E8F9FEFAEAFF68" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31" type="discussion">
<paragraph id="A230B37B4B5AFFA8E7E8F9FEFB18FF66" blockId="30.[151,1437,150,2030]" lastBlockId="32.[151,1437,151,2033]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<emphasis id="90FB6F694B5AFF96E7E8F9FEFE7EF9F2" bold="true" box="[199,437,1619,1645]" pageId="30" pageNumber="31">Taxonomic remarks</emphasis>
. 
<taxonomicName id="658FC8F84B5AFF96E6EEF9FEFD64F9F2" authorityName="De Man" authorityYear="1920" box="[449,687,1619,1645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E6EEF9FEFD64F9F2" box="[449,687,1619,1645]" italics="true" pageId="30" pageNumber="31">Alpheus eurydactylus</emphasis>
</taxonomicName>
is morphologically very close to 
<taxonomicName id="658FC8F84B5AFF96E332F9FEFB71F9F2" authorityName="sensu De Man" authorityYear="1897" box="[1053,1210,1619,1645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E332F9FEFB71F9F2" box="[1053,1210,1619,1645]" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
, as redefined above. The two species can be most easily separated from each other by the dorsal shoulder of the major chela sloping with an angle of&gt;45º into the adjacent transverse groove in 
<taxonomicName id="658FC8F84B5AFF96E47FF934FBCBF92C" authorityName="De Man" authorityYear="1920" box="[848,1024,1689,1715]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E47FF934FBCBF92C" box="[848,1024,1689,1715]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B5AFF96E33DF934FB63F92C" box="[1042,1192,1689,1715]" captionStart="FIGURE 11" captionStartId="25.[151,250,1892,1916]" captionTargetBox="[158,1429,181,1868]" captionTargetId="figure-15@25.[158,1429,181,1868]" captionTargetPageId="25" captionText="FIGURE 11. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—cephalothorax and chelipeds, dorsal view; B—frontal region, dorsal view; C, D— major (left) cheliped, mesial view, with slightly different angles and illuminations (slightly more ventral and enhancing mesial subdistal ridge on pollex in D). Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912324" httpUri="https://zenodo.org/record/7912324/files/figure.png" pageId="30" pageNumber="31">Figs. 11C, D</figureCitation>
, 
<figureCitation id="3AB4AFFE4B5AFF96E39AF934FAC2F92C" box="[1205,1289,1689,1716]" captionStart="FIGURE 15" captionStartId="29.[151,250,1821,1845]" captionTargetBox="[167,1423,194,1791]" captionTargetId="figure-15@29.[151,1436,181,1798]" captionTargetPageId="29" captionText="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912338" httpUri="https://zenodo.org/record/7912338/files/figure.png" pageId="30" pageNumber="31">15A, C</figureCitation>
) 
<emphasis id="90FB6F694B5AFF96E234F937FAF2F92C" box="[1307,1337,1690,1715]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
forming a rounded tooth overhanging the transverse groove in 
<taxonomicName id="658FC8F84B5AFF96E43FF911FC78F949" authorityName="sensu De Man" authorityYear="1897" box="[784,947,1724,1750]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E43FF911FC78F949" box="[784,947,1724,1750]" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B5AFF96E4E9F911FB99F949" box="[966,1106,1723,1750]" captionStart="FIGURE 5" captionStartId="14.[151,250,1935,1959]" captionTargetBox="[151,1435,379,1912]" captionTargetId="figure-95@14.[151,1435,379,1912]" captionTargetPageId="14" captionText="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912304" httpUri="https://zenodo.org/record/7912304/files/figure.png" pageId="30" pageNumber="31">Figs. 5C, D</figureCitation>
, 
<figureCitation id="3AB4AFFE4B5AFF96E34FF911FB62F949" box="[1120,1193,1724,1750]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="30" pageNumber="31">7A, B</figureCitation>
). In 
<taxonomicName id="658FC8F84B5AFF96E3CBF911FA5EF949" authorityName="De Man" authorityYear="1920" box="[1252,1429,1724,1750]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E3CBF911FA5EF949" box="[1252,1429,1724,1750]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
, the mesial palmar surface of the major chela is granulated only distally and subdistally (
<figureCitation id="3AB4AFFE4B5AFF96E3ABF972FB27F966" box="[1156,1260,1758,1785]" captionStart="FIGURE 15" captionStartId="29.[151,250,1821,1845]" captionTargetBox="[167,1423,194,1791]" captionTargetId="figure-15@29.[151,1436,181,1798]" captionTargetPageId="29" captionText="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912338" httpUri="https://zenodo.org/record/7912338/files/figure.png" pageId="30" pageNumber="31">Fig. 15C</figureCitation>
), whereas in 
<taxonomicName id="658FC8F84B5AFF96E2ABF972FEDCF884" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E2ABF972FEDCF884" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
, several fields of granules are also present on the central portion of the palmar surface (
<figureCitation id="3AB4AFFE4B5AFF96E22EF8ACFA97F883" box="[1281,1372,1793,1820]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="30" pageNumber="31">Fig. 7A</figureCitation>
). The dactylar plunger of 
<taxonomicName id="658FC8F84B5AFF96E652F888FDE4F8A1" authorityName="De Man" authorityYear="1920" box="[381,559,1828,1855]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E652F888FDE4F8A1" box="[381,559,1828,1855]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
is larger, heavier and not continuous with the ventral edge of the dactylus, as in 
<taxonomicName id="658FC8F84B5AFF96E7F9F8E5FEB3F8FE" authorityName="sensu De Man" authorityYear="1897" box="[214,376,1863,1890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E7F9F8E5FEB3F8FE" box="[214,376,1863,1890]" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
(cf. 
<figureCitation id="3AB4AFFE4B5AFF96E69FF8EAFDDEF8FD" box="[432,533,1863,1890]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="30" pageNumber="31">Figs. 7D</figureCitation>
, 
<figureCitation id="3AB4AFFE4B5AFF96E50CF8EAFD9DF8FD" box="[547,598,1863,1890]" captionStart="FIGURE 15" captionStartId="29.[151,250,1821,1845]" captionTargetBox="[167,1423,194,1791]" captionTargetId="figure-15@29.[151,1436,181,1798]" captionTargetPageId="29" captionText="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912338" httpUri="https://zenodo.org/record/7912338/files/figure.png" pageId="30" pageNumber="31">15D</figureCitation>
). The rostral carina and rostro-orbital furrows are more demarcated in 
<taxonomicName id="658FC8F84B5AFF96E2ABF8E5FEEDF81B" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E2ABF8E5FEEDF81B" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="658FC8F84B5AFF96E650F8C6FDD5F81B" authorityName="sensu De Man" authorityYear="1897" box="[383,542,1898,1925]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E650F8C6FDD5F81B" box="[383,542,1898,1925]" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
, where they are both not or barely distinct (cf. 
<figureCitation id="3AB4AFFE4B5AFF96E309F8C7FB42F81B" box="[1062,1161,1898,1924]" captionStart="FIGURE 6" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[203,1391,197,1776]" captionTargetId="figure-15@15.[183,1404,181,1784]" captionTargetPageId="15" captionText="FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle." figureDoi="http://doi.org/10.5281/zenodo.7912306" httpUri="https://zenodo.org/record/7912306/files/figure.png" pageId="30" pageNumber="31">Figs. 6A</figureCitation>
, 
<figureCitation id="3AB4AFFE4B5AFF96E3BCF8C7FB0CF81B" box="[1171,1223,1898,1924]" captionStart="FIGURE 14" captionStartId="28.[151,250,1839,1863]" captionTargetBox="[174,1408,280,1804]" captionTargetId="figure-62@28.[164,1423,273,1815]" captionTargetPageId="28" captionText="FIGURE 14. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–H, J–N]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [I]; A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—same, dactylus, dorsal view; I—same, ischium, lateral view; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—same, basis and ischium, lateral view; M—second pleopod, appendices masculina and interna, anterior view; N—right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912334" httpUri="https://zenodo.org/record/7912334/files/figure.png" pageId="30" pageNumber="31">14A</figureCitation>
). Some differences between the two species also exist in the sculpture and proportions of the female minor chelae (cf. 
<figureCitation id="3AB4AFFE4B5AFF96E3C2F820FA84F837" box="[1261,1359,1933,1960]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="30" pageNumber="31">Figs. 7H</figureCitation>
, 
<figureCitation id="3AB4AFFE4B5AFF96E273F820FA44F837" box="[1372,1423,1933,1960]" captionStart="FIGURE 15" captionStartId="29.[151,250,1821,1845]" captionTargetBox="[167,1423,194,1791]" captionTargetId="figure-15@29.[151,1436,181,1798]" captionTargetPageId="29" captionText="FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani,Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912338" httpUri="https://zenodo.org/record/7912338/files/figure.png" pageId="30" pageNumber="31">15H</figureCitation>
). The colour patterns of 
<taxonomicName id="658FC8F84B5AFF96E6BFF81CFDF6F855" authorityName="De Man" authorityYear="1920" box="[400,573,1968,1995]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E6BFF81CFDF6F855" box="[400,573,1968,1995]" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B5AFF96E564F81DFD50F855" box="[587,667,1968,1994]" captionStart="FIGURE 16" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[152,1435,181,1885]" captionTargetId="figure-15@31.[152,1435,181,1885]" captionTargetPageId="31" captionText="FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D)." figureDoi="http://doi.org/10.5281/zenodo.7912340" httpUri="https://zenodo.org/record/7912340/files/figure.png" pageId="30" pageNumber="31">Fig. 16</figureCitation>
) and 
<taxonomicName id="658FC8F84B5AFF96E5F9F81CFCBFF855" authorityName="sensu De Man" authorityYear="1897" box="[726,884,1968,1995]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B5AFF96E5F9F81CFCBFF855" box="[726,884,1968,1995]" italics="true" pageId="30" pageNumber="31">A. euphrosyne</emphasis>
</taxonomicName>
(
<figureCitation id="3AB4AFFE4B5AFF96E4ADF81DFC25F855" box="[898,1006,1968,1994]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="19.[151,250,1902,1926]" captionStartId-1="20.[151,250,1875,1899]" captionStartId-2="21.[151,250,1866,1890]" captionTargetBox-0="[151,1435,181,1877]" captionTargetBox-1="[151,1435,181,1851]" captionTargetBox-2="[159,1429,181,1842]" captionTargetId-0="figure-15@19.[151,1435,181,1878]" captionTargetId-1="figure-21@20.[151,1435,181,1851]" captionTargetId-2="figure-15@21.[159,1429,181,1842]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." captionText-1="FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae." captionText-2="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7912312" figureDoi-1="http://doi.org/10.5281/zenodo.7912318" figureDoi-2="http://doi.org/10.5281/zenodo.7912320" httpUri-0="https://zenodo.org/record/7912312/files/figure.png" httpUri-1="https://zenodo.org/record/7912318/files/figure.png" httpUri-2="https://zenodo.org/record/7912320/files/figure.png" pageId="30" pageNumber="31">Fig. 8–10</figureCitation>
) are markedly different. In addition, 
<taxonomicName id="658FC8F84B5AFF96E2ABF81CFEEDF872" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5AFF96E2ABF81CFEEDF872" italics="true" pageId="30" pageNumber="31">A. eurydactylus</emphasis>
</taxonomicName>
is a largely intertidal and shallow subtidal species commonly encountered in soggy mud in mangroves and on estuarine mudflats, whereas 
<taxonomicName id="658FC8F84B64FFA8E50CFF3AFD0AFF2E" authorityName="sensu De Man" authorityYear="1897" box="[547,705,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B64FFA8E50CFF3AFD0AFF2E" box="[547,705,151,177]" italics="true" pageId="32" pageNumber="33">A. euphrosyne</emphasis>
</taxonomicName>
appears to be a subtidal snapping shrimp preferring sandy-muddy bottoms in deeper water (
<quantity id="65771E9E4B64FFA8E697FF16FDD8FF4A" box="[440,531,187,213]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="32" pageNumber="33" unit="m" value="17.5" valueMax="27.0" valueMin="8.0">8–27 m</quantity>
), in brackish lagoons and near estuaries. For separation of 
<taxonomicName id="658FC8F84B64FFA8E380FF16FA95FF4A" authorityName="De Man" authorityYear="1920" box="[1199,1374,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E380FF16FA95FF4A" box="[1199,1374,187,213]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
from other morphologically similar species (especially 
<taxonomicName id="658FC8F84B64FFA8E5EAFF72FCD7FF66" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[709,796,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B64FFA8E5EAFF72FCD7FF66" box="[709,796,223,249]" italics="true" pageId="32" pageNumber="33">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B64FFA8E40CFF72FCB7FF66" bold="true" box="[803,892,223,249]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="8BC8D2124B64FFA8E40CFF72FCB7FF66" box="[803,892,223,249]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) see below or refer to 
<tableCitation id="EF0D86C04B64FFA8E356FF72FB06FF66" box="[1145,1229,223,249]" captionStart="TABLE 1" captionStartId="93.[145,233,465,491]" captionTargetPageId="93" captionText="TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897—A. micro-" httpUri="http://table.plazi.org/id/F6F0E3F34B19FFD5E7BEFE7CF8F8FD90" pageId="32" pageNumber="33" tableUuid="F6F0E3F34B19FFD5E7BEFE7CF8F8FD90">Table 1</tableCitation>
.
</paragraph>
<caption id="F6F0E3F34B5BFF97E7B8F8D9FEBFF868" ID-DOI="http://doi.org/10.5281/zenodo.7912340" ID-Zenodo-Dep="7912340" httpUri="https://zenodo.org/record/7912340/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1908,1932]" targetBox="[152,1435,181,1885]" targetPageId="31" targetType="figure">
<paragraph id="A230B37B4B5BFF97E7B8F8D9FEBFF868" blockId="31.[151,1437,1908,2039]" pageId="31" pageNumber="32">
<emphasis id="90FB6F694B5BFF97E7B8F8D9FED0F813" bold="true" box="[151,283,1908,1932]" pageId="31" pageNumber="32">FIGURE 16</emphasis>
. 
<taxonomicName id="658FC8F84B5BFF97E606F8D8FD6EF812" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[297,677,1909,1933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B5BFF97E606F8D8FDCFF812" box="[297,516,1909,1933]" italics="true" pageId="31" pageNumber="32">Alpheus eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B5BFF97E523F8D8FD6EF812" author="De Man, J. G." box="[524,677,1909,1933]" pageId="31" pageNumber="32" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D).
</paragraph>
</caption>
<paragraph id="A230B37B4B64FFA8E7E8FEAEFCEBFE53" blockId="32.[151,1437,151,2033]" pageId="32" pageNumber="33">
Some previous records of 
<taxonomicName id="658FC8F84B64FFA8E6C0FEAEFD44FE82" authorityName="sensu De Man" authorityYear="1897" box="[495,655,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B64FFA8E6C0FEAEFD44FE82" box="[495,655,259,285]" italics="true" pageId="32" pageNumber="33">A. euphrosyne</emphasis>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B64FFA8E59AFEAEFC17FE82" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[693,988,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B64FFA8E59AFEAEFC17FE82" box="[693,988,259,285]" italics="true" pageId="32" pageNumber="33">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
, for instance those in 
<bibRefCitation id="C61ECE8A4B64FFA8E3F9FEAEFF10FEDE" author="Banner, A. H. &amp; Banner, D. M." pageId="32" pageNumber="33" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B64FFA8E7C9FE8AFEEBFEDE" author="Banner, D. M. &amp; Banner, A. H." box="[230,288,295,321]" pageId="32" pageNumber="33" pagination="1 - 99" refId="ref69349" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">1981</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B64FFA8E604FE8AFEADFEDE" author="Banner, D. M. &amp; Banner, A. H." box="[299,358,295,321]" pageId="32" pageNumber="33" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B64FFA8E65FFE8AFE62FEDE" author="Banner, A. H. &amp; Banner, D. M." box="[368,425,295,321]" pageId="32" pageNumber="33" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
) and 
<bibRefCitation id="C61ECE8A4B64FFA8E6CAFE8AFD6BFEDE" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[485,672,295,321]" pageId="32" pageNumber="33" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B64FFA8E539FE8AFD8DFEDE" box="[534,582,295,321]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. (2019)
</bibRefCitation>
, may actually refer to 
<taxonomicName id="658FC8F84B64FFA8E4B9FE8AFB88FEDE" authorityName="De Man" authorityYear="1920" box="[918,1091,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E4B9FE8AFB88FEDE" box="[918,1091,295,321]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. At least some specimens from 
<collectingCountry id="DA98F3EB4B64FFA8E7B8FEE7FF35FEFB" box="[151,254,330,356]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
identified by D.M. Banner &amp; A.H. Banner as 
<taxonomicName id="658FC8F84B64FFA8E422FEE7FBFEFEFB" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[781,1077,330,356]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B64FFA8E422FEE7FBFEFEFB" box="[781,1077,330,356]" italics="true" pageId="32" pageNumber="33">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
were reexamined by the author and reidentified as 
<taxonomicName id="658FC8F84B64FFA8E65BFEC0FDEEFE19" authorityName="De Man" authorityYear="1920" box="[372,549,364,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E65BFEC0FDEEFE19" box="[372,549,364,391]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. At least part of the material reported as 
<taxonomicName id="658FC8F84B64FFA8E32CFEC0FB6EFE19" authorityName="sensu De Man" authorityYear="1897" box="[1027,1189,364,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B64FFA8E32CFEC0FB6EFE19" box="[1027,1189,364,391]" italics="true" pageId="32" pageNumber="33">A. euphrosyne</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B64FFA8E3FCFEC1FA53FE18" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[1235,1432,364,391]" pageId="32" pageNumber="33" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B64FFA8E228FEC0FAF1FE19" box="[1287,1338,364,390]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. (2019)
</bibRefCitation>
, collected at various localities in southern and eastern 
<collectingCountry id="DA98F3EB4B64FFA8E5D9FE22FCF7FE36" box="[758,828,399,425]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
, could also represent 
<taxonomicName id="658FC8F84B64FFA8E31EFE3DFB2BFE36" authorityName="De Man" authorityYear="1920" box="[1073,1248,399,426]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E31EFE3DFB2BFE36" box="[1073,1248,399,426]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
, at least judging from the detailless illustrations provided by these authors.
</paragraph>
<paragraph id="A230B37B4B64FFA8E7E8FE7BFD1DFB81" blockId="32.[151,1437,151,2033]" pageId="32" pageNumber="33">
<bibRefCitation id="C61ECE8A4B64FFA8E7E8FE7BFE24FE70" author="Banner, D. M. &amp; Banner, A. H." box="[199,495,469,496]" pageId="32" pageNumber="33" pagination="1 - 99" refId="ref69349" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner’s (1981</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B64FFA8E6D2FE78FDFDFE70" author="Banner, A. H. &amp; Banner, D. M." box="[509,566,469,495]" pageId="32" pageNumber="33" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
) records of 
<taxonomicName id="658FC8F84B64FFA8E5EEFE7BFC20FE70" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[705,1003,469,496]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B64FFA8E5EEFE7BFC20FE70" box="[705,1003,469,496]" italics="true" pageId="32" pageNumber="33">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
from the northern and western Indian Ocean cannot be confirmed as 
<taxonomicName id="658FC8F84B64FFA8E6DDFE54FD6AFD8D" authorityName="De Man" authorityYear="1920" box="[498,673,504,531]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E6DDFE54FD6AFD8D" box="[498,673,504,531]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
without examination of their material, which unfortunately remains untraceable. The photograph of a snapping shrimp identified as 
<taxonomicName id="658FC8F84B64FFA8E4A5FDB1FBF2FDAA" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[906,1081,539,566]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B64FFA8E4A5FDB1FBF2FDAA" box="[906,1081,539,566]" italics="true" pageId="32" pageNumber="33">A. malabaricus</emphasis>
</taxonomicName>
from Vembanad Lake, 
<collectingRegion id="604B7D994B64FFA8E27DFDB6FA56FDAA" box="[1362,1437,539,565]" country="India" name="Kerala" pageId="32" pageNumber="33">Kerala</collectingRegion>
(
<bibRefCitation id="C61ECE8A4B64FFA8E7B0FD93FE14FDC7" author="Harikrishnan, M. &amp; Kurup, B. M." box="[159,479,574,601]" pageId="32" pageNumber="33" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup 2008</bibRefCitation>
: fig. 1, specimen on the right side), superficially resembles 
<taxonomicName id="658FC8F84B64FFA8E35AFD92FAE9FDC7" authorityName="De Man" authorityYear="1920" box="[1141,1314,574,601]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E35AFD92FAE9FDC7" box="[1141,1314,574,601]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. However, a short correspondence between the present author and Dr. M. Harikrishnan in 2014 (accompanied by photographs of some morphological details of the preserved material) led to the conclusion that it could be 
<taxonomicName id="658FC8F84B64FFA8E399FD29FAF7FD01" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1206,1340,644,670]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B64FFA8E399FD29FAF7FD01" box="[1206,1340,644,670]" italics="true" pageId="32" pageNumber="33">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B64FFA8E26CFD29FA57FD01" bold="true" box="[1347,1436,644,670]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="8BC8D2124B64FFA8E26CFD29FA57FD01" box="[1347,1436,644,670]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
or another closely related taxon (see below). On the other hand, a colour photograph of a large male from 
<collectingCountry id="DA98F3EB4B64FFA8E27EFD0AFF18FD7B" name="India" pageId="32" pageNumber="33">Kochi, India</collectingCountry>
, sent by Dr. Anil Kumar in 2021 (A. Anker, pers. obs.) shows a snapping shrimp with essentially the same colour pattern as 
<taxonomicName id="658FC8F84B64FFA8E674FD40FDC2FC98" authorityName="De Man" authorityYear="1920" box="[347,521,749,775]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E674FD40FDC2FC98" box="[347,521,749,775]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B64FFA8E560FD40FD65FC98" box="[591,686,749,775]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B64FFA8E5CAFD40FC82FC98" box="[741,841,749,775]" name="Thailand" pageId="32" pageNumber="33">Thailand</collectingCountry>
(
<figureCitation id="3AB4AFFE4B64FFA8E477FD40FC67FC98" box="[856,940,749,775]" captionStart="FIGURE 16" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[152,1435,181,1885]" captionTargetId="figure-15@31.[152,1435,181,1885]" captionTargetPageId="31" captionText="FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D)." figureDoi="http://doi.org/10.5281/zenodo.7912340" httpUri="https://zenodo.org/record/7912340/files/figure.png" pageId="32" pageNumber="33">Fig. 16</figureCitation>
; see also 
<bibRefCitation id="C61ECE8A4B64FFA8E337FD40FB6AFC98" author="Marin, I. N." box="[1048,1185,749,775]" pageId="32" pageNumber="33" pagination="369 - 385" refId="ref72908" refString="Marin, I. N. (2021) A new infaunal species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) and a new crustacean association from anoxic mangrove habitats in southern Vietnam. Arthropoda Selecta, 30, 369 - 385. https: // doi. org / 10.15298 / arthsel. 30.3.10" type="journal article" year="2021">Marin 2021</bibRefCitation>
: fig. 1a). In particular, the general shape and the diagnostic colour of the dactylar plunger of the Indian and the Vietnamese specimens are identical. The only noticeable difference is the distally blue second pereiopod in the specimens from 
<collectingCountry id="DA98F3EB4B64FFA8E211FC9EFA56FCD2" box="[1342,1437,819,845]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B64FFA8E7E8FCFBFEE5FCEF" box="[199,302,854,880]" name="Thailand" pageId="32" pageNumber="33">Thailand</collectingCountry>
; this appendage is entirely reddish in the Indian specimen. Although the presence of 
<taxonomicName id="658FC8F84B64FFA8E3C2FCFBFA57FCEF" authorityName="De Man" authorityYear="1920" box="[1261,1436,854,880]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E3C2FCFBFA57FCEF" box="[1261,1436,854,880]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
in 
<collectingCountry id="DA98F3EB4B64FFA8E798FCD5FF38FC0D" box="[183,243,888,914]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
, or more generally in the Indian Ocean west of Sumatra, cannot be confirmed at this stage, the above observations strongly suggest that the geographical range of 
<taxonomicName id="658FC8F84B64FFA8E412FC31FC20FC2A" authorityName="De Man" authorityYear="1920" box="[829,1003,923,950]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E412FC31FC20FC2A" box="[829,1003,923,950]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
may extend to south-western 
<collectingCountry id="DA98F3EB4B64FFA8E214FC36FABEFC2A" box="[1339,1397,923,949]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
(as indicated in 
<figureCitation id="3AB4AFFE4B64FFA8E60EFC13FE4DFC46" box="[289,390,958,985]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="32" pageNumber="33">Fig. 51B</figureCitation>
). In any case, a more thorough examination of the Indian material (unavailable for the present study) and its molecular comparison with the South-East Asian material will be needed for clarification of the actual distribution of 
<taxonomicName id="658FC8F84B64FFA8E611FBA8FE27FB81" authorityName="De Man" authorityYear="1920" box="[318,492,1028,1055]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E611FBA8FE27FB81" box="[318,492,1028,1055]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
in the Indian Ocean.
</paragraph>
<paragraph id="A230B37B4B64FFA9E7E8FB8AFAEAFF68" blockId="32.[151,1437,151,2033]" lastBlockId="33.[151,1437,151,248]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
The material herein tentatively identified as 
<taxonomicName id="658FC8F84B64FFA8E589FB85FCB9FBDE" authorityName="De Man" authorityYear="1920" box="[678,882,1063,1090]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E589FB85FD75FBDD" box="[678,702,1064,1090]" italics="true" pageId="32" pageNumber="33">A.</emphasis>
cf. 
<emphasis id="90FB6F694B64FFA8E5CCFB8AFCB9FBDE" box="[739,882,1063,1089]" italics="true" pageId="32" pageNumber="33">eurydactylus</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B64FFA8E49EFB8AFBE8FBDE" box="[945,1059,1063,1089]" name="Singapore" pageId="32" pageNumber="33">Singapore</collectingCountry>
is morphologically heterogeneous and possibly contain two different species. The specimens from Jurong, Sungei Buloh and Lim Chu Kang are different from the typical 
<taxonomicName id="658FC8F84B64FFA8E6EEFBC0FDBAFB18" authorityName="De Man" authorityYear="1920" box="[449,625,1133,1159]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E6EEFBC0FDBAFB18" box="[449,625,1133,1159]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
in several details. For instance, in the incomplete male from Lim Chu Kang (ZRC 2014.0724), the third pereiopod merus is about 3.5 times as long as wide, i.e., is relatively broader than in 
<taxonomicName id="658FC8F84B64FFA8E79CFB1EFE94FB52" authorityName="De Man" authorityYear="1920" box="[179,351,1203,1229]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E79CFB1EFE94FB52" box="[179,351,1203,1229]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. The ischia of the third and fourth pereiopods have no trace of a spiniform seta on the ventrolateral margin, as in 
<taxonomicName id="658FC8F84B64FFA8E603FB7BFE24FB6F" authorityName="sensu De Man" authorityYear="1897" box="[300,495,1238,1264]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B64FFA8E603FB7BFE24FB6F" box="[300,495,1238,1264]" italics="true" pageId="32" pageNumber="33">A. microrhynchus</emphasis>
</taxonomicName>
(see below), and unlike the ischia of 
<taxonomicName id="658FC8F84B64FFA8E4A5FB7BFBFCFB6F" authorityName="De Man" authorityYear="1920" box="[906,1079,1238,1264]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E4A5FB7BFBFCFB6F" box="[906,1079,1238,1264]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
, which are usually armed with a stout spiniform seta. The third pereiopod dactylus is proportionally longer, at least 0.6 as long as the propodus. In the second pereiopod, the relative length ratio of the carpal subarticles (4.5: 2: 1: 1: 2) is slightly different from that of the typical 
<taxonomicName id="658FC8F84B64FFA8E61BFA92FE28FAC6" authorityName="De Man" authorityYear="1920" box="[308,483,1343,1369]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E61BFA92FE28FAC6" box="[308,483,1343,1369]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
(4.2: 2.5: 1: 1: 1.5). Unfortunately, the major cheliped of the male from Lim Chu Kang appears to be regenerated, judging from the somewhat unusual proportions of the chela. Nevertheless, it can be characterised by the ventromesial margin of the merus distally unarmed; the dorsal shoulder gently sloping into the adjacent transversal groove; the mesial face of the pollex with some granules, but without mesial subdistal ridge; and the moderately developed, obliquely “sliced” plunger. The rostro-orbital region is aberrant in the absence of a rostrum, probably resulting from an injury (also indicated by the much smaller, clearly regenerated right antennular peduncle). The antennal basicerite is unarmed on both sides, as in the typical 
<taxonomicName id="658FC8F84B64FFA8E4D8F9BCFB6EF9B5" authorityName="De Man" authorityYear="1920" box="[1015,1189,1552,1579]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E4D8F9BCFB6EF9B5" box="[1015,1189,1552,1579]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
, but the carpocerite is longer than in 
<taxonomicName id="658FC8F84B64FFA8E613F999FE27F9D2" authorityName="De Man" authorityYear="1920" box="[316,492,1587,1614]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E613F999FE27F9D2" box="[316,492,1587,1614]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
, reaching far beyond the distal margin of the scaphocerite, as seen in 
<taxonomicName id="658FC8F84B64FFA8E22BF999FEAAF9EF" authority="(Yeo &amp; Ng 1996)" baseAuthorityName="Yeo &amp; Ng" baseAuthorityYear="1996" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="90FB6F694B64FFA8E22BF999FA57F9D2" box="[1284,1436,1587,1614]" italics="true" pageId="32" pageNumber="33">A. paludicola</emphasis>
(
<bibRefCitation id="C61ECE8A4B64FFA8E7B0F9FBFE92F9EF" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[159,345,1622,1649]" pageId="32" pageNumber="33" pagination="37 - 63" refId="ref75202" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (1996) A new species of freshwater snapping shrimp, Alpheus cyanoteles (Decapoda: Caridea: Alpheidae) from Peninsular Malaysia and a redescription of Alpheus paludicola Kemp, 1915. Raffles Bulletin of Zoology, 44, 37 - 63." type="journal article" year="1996">Yeo &amp; Ng 1996</bibRefCitation>
)
</taxonomicName>
. The females from Jurong and Sungei Buloh are similar to the male from Lim Chu Kang in the proportions of the third and second pereiopods, antennules, and antennae. The major cheliped of the complete female from Jurong (ZRC 1979.4.4.6) appears to be regenerated and is generally similar to that of the male, except for the absence of granulation. The frontal margin of the carapace of this specimen bears a minute rostrum, similar to that of 
<taxonomicName id="658FC8F84B64FFA8E627F94FFE04F963" authorityName="sensu De Man" authorityYear="1897" box="[264,463,1762,1788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B64FFA8E627F94FFE04F963" box="[264,463,1762,1788]" italics="true" pageId="32" pageNumber="33">A. microrhynchus</emphasis>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B64FFA8E6D7F94FFD5AF963" baseAuthorityName="Yeo &amp; Ng" baseAuthorityYear="1996" box="[504,657,1762,1788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="paludicola">
<emphasis id="90FB6F694B64FFA8E6D7F94FFD5AF963" box="[504,657,1762,1788]" italics="true" pageId="32" pageNumber="33">A. paludicola</emphasis>
</taxonomicName>
. The minor chelipeds of the females from Jurong and Sungei Buloh (ZRC 1992.8064) are characterised by the palm being smooth on both sides and bearing a slight ventral notch or sinus, and with the fingers slightly longer than the palm; in these features, they are generally concordant with the female minor cheliped of 
<taxonomicName id="658FC8F84B64FFA8E695F8E6FDA3F8FA" authorityName="De Man" authorityYear="1920" box="[442,616,1867,1893]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E695F8E6FDA3F8FA" box="[442,616,1867,1893]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. However, the relatively broad merus of the third pereiopod, the absence of a median dorsal groove on the telson and the very short rostrum of these females are features that do not allow the identification of the Jurong and Sungei Buloh specimens as 
<taxonomicName id="658FC8F84B64FFA8E4A7F83CFBF1F835" authorityName="De Man" authorityYear="1920" box="[904,1082,1936,1963]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E4A7F83CFBF1F835" box="[904,1082,1936,1963]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
. In many respects, the female from Sungei Buloh, for instance, appears to be intermediate between 
<taxonomicName id="658FC8F84B64FFA8E49FF819FBABF852" authorityName="De Man" authorityYear="1920" box="[944,1120,1971,1998]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B64FFA8E49FF819FBABF852" box="[944,1120,1971,1998]" italics="true" pageId="32" pageNumber="33">A. eurydactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B64FFA8E3B5F819FAAAF852" authorityName="sensu De Man" authorityYear="1897" box="[1178,1377,1971,1998]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B64FFA8E3B5F819FAAAF852" box="[1178,1377,1971,1998]" italics="true" pageId="32" pageNumber="33">A. microrhynchus</emphasis>
</taxonomicName>
. The ovigerous female from 
<collectingCountry id="DA98F3EB4B64FFA8E6B3F87BFDE2F86F" box="[412,553,2006,2032]" name="India" pageId="32" pageNumber="33">Kochi, India</collectingCountry>
(RMNH Crus.D.49749), is also problematic, largely for the same reason as the female from Jurong, i.e., in the absence of marked granulation on the mesial face of the major cheliped. A molecular comparison (e.g., using the barcoding segment of the mitochondrial COI gene) of the material herein assigned to 
<taxonomicName id="658FC8F84B65FFA9E2ABFF17FE81FF68" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B65FFA9E2ABFF17FA56FF4B" box="[1412,1437,186,212]" italics="true" pageId="33" pageNumber="34">A.</emphasis>
cf. 
<emphasis id="90FB6F694B65FFA9E794FF70FE81FF68" box="[187,330,221,247]" italics="true" pageId="33" pageNumber="34">eurydactylus</emphasis>
</taxonomicName>
with the typical 
<taxonomicName id="658FC8F84B65FFA9E529FF73FD7FFF68" authorityName="De Man" authorityYear="1920" box="[518,692,221,248]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B65FFA9E529FF73FD7FFF68" box="[518,692,221,248]" italics="true" pageId="33" pageNumber="34">A. eurydactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B65FFA9E5C4FF73FC7BFF68" authorityName="sensu De Man" authorityYear="1897" box="[747,944,221,248]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="90FB6F694B65FFA9E5C4FF73FC7BFF68" box="[747,944,221,248]" italics="true" pageId="33" pageNumber="34">A. microrhynchus</emphasis>
</taxonomicName>
is needed to confirm its identity.
</paragraph>
</subSubSection>
</treatment>
</document>