237 lines
26 KiB
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237 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang & Liang, Hongbin" docDate="2013" docId="28E9333193E16B8DFCC15DD9CA15777A" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Paraphaea minor Shi & Liang 2013, sp. n." docType="treatment" docUuid="C0987D82-6F52-42F8-93A8-D105D2A8D0B1" docUuidSource="ZooBank" docVersion="6" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="18" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="17" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhou, Hongzhang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-04-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>284</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>129</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
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<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152042282" LSID="urn:lsid:zoobank.org:act:C0987D82-6F52-42F8-93A8-D105D2A8D0B1" httpUri="http://treatment.plazi.org/id/28E9333193E16B8DFCC15DD9CA15777A" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection pageId="16" pageNumber="17" type="nomenclature">
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<paragraph pageId="16" pageNumber="17">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:C0987D82-6F52-42F8-93A8-D105D2A8D0B1" authority="Shi & Liang" authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paraphaea minor" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minor" status="sp. n.">Paraphaea minor Shi & Liang</taxonomicName>
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<taxonomicNameLabel pageId="16" pageNumber="17">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="16" pageNumber="17">Habitus: Figs 3</figureCitation>
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<figureCitation captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Type materials, scale bars = 2.0 mm: 31 Lectotype of Plochionus binotatus Dejean [= Paraphaea binotata (Dejean)] 32 Lectotype of Calleida discophora Chaudoir [= Paraphaea binotata (Dejean)] 33 Lectotype of Paraphaea signifera Bates [= Paraphaea binotata (Dejean)] 34 Holotype of Parena formosana Jedlicka [= Paraphaea formosana (Jedlicka)] 35 Holotype of Paraphaea minor sp. n. 36 Holotype of Coptodera philippinensis Jedlicka [= Paraphaea philippinensis (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29679" pageId="16" pageNumber="17">35</figureCitation>
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<figureCitation captionStart="Figure 69" captionStartId="F14" captionText="Figure 69. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Paraphaea minor sp. n., holotype." httpUri="https://binary.pensoft.net/fig/29687" pageId="16" pageNumber="17">male genitalia: Figs 69</figureCitation>
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<figureCitation captionStart="Figures 95–110" captionStartId="F40" captionText="Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29716" pageId="16" pageNumber="17">96</figureCitation>
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<figureCitation captionStart="Figures 111–125" captionStartId="F41" captionText="Figures 111 - 125. Left ovipositors of females, ventral view, scale bars = 0.2 mm: 111 Paraphaea binotata (Dejean), a specimen from Guangxi 112 Paraphaea minor sp. n., a paratype from Hainan 113 Paraphaea philippinensis (Jedlicka), holotype 114 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 115 Physodera dejeani Eschscholtz, a specimen from Yunnan 116 Physodera eschscholtzii Parry, a specimen from Yunnan 117 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 118 Allocota aurata (Bates), a specimen from Shaanxi 119 Allocota bicolor sp. n., a paratype from Guangdong 120 Lachnoderma asperum Bates, a specimen from Zhejiang 121 Dasiosoma bellum (Habu), a specimen from Guangdong 122 Dasiosoma maindroni (Tian & Deuve), a specimen from Tonkin 123 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 124 Orionella lewisii (Bates), a specimen from Zhejiang 125 Endynomena pradieri (Fairmaire), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29719" pageId="16" pageNumber="17">female genitalia: Figs 112</figureCitation>
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<figureCitation captionStart="Figures 126–131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="16" pageNumber="17">127</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="type material">
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<paragraph pageId="16" pageNumber="17">Type material.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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<emphasis bold="true" pageId="16" pageNumber="17">Holotype</emphasis>
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(MNHN): male, body length = 5.8 mm, board mounted, genitalia dissected and deposited in microvial pinned under specimen, "Hoa-Binh / Tonkin / A. de Cooman"; "MUSEUM PARIS / Coll. Ch. ALLUAUD"; "HOLOTYPE ♂/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label][Figs 69, 96, 35].
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<emphasis bold="true" pageId="16" pageNumber="17">Paratypes (</emphasis>
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Total 1 male, 6 females
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<emphasis bold="true" pageId="16" pageNumber="17">)</emphasis>
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:
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<emphasis bold="true" pageId="16" pageNumber="17">Vietnam</emphasis>
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: 1 female (MNHN), "
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<emphasis italics="true" pageId="16" pageNumber="17">Tonkin</emphasis>
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/
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<emphasis italics="true" pageId="16" pageNumber="17">Dap-Can</emphasis>
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"; "MUSEUM PARIS / Ex. Coll. M. MAINDRON / Coll. G. BABAULT 1930"; "
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<emphasis italics="true" pageId="16" pageNumber="17">Anchista</emphasis>
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/
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<emphasis italics="true" pageId="16" pageNumber="17">circumdata</emphasis>
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/
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<emphasis italics="true" pageId="16" pageNumber="17">sp. n</emphasis>
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."; "PARATYPE ♀/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label].
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<emphasis bold="true" pageId="16" pageNumber="17">Thailand</emphasis>
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: 1 female (MNHN), "MUSEUM PARIS / BANGKOK / LARNAUDIE 349-64"; a round yellow label probably meaning Oriental with backside written "
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<emphasis italics="true" pageId="16" pageNumber="17">3kg</emphasis>
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/
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<emphasis italics="true" pageId="16" pageNumber="17">Gk</emphasis>
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"; "PARATYPE ♀/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label]. 1 male (CRS), "Thailand bor. occ. / Pai / Soppong / 28.5-5.6. 1997 / Lgt. M. Snizek"; "PARATYPE ♂/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label].
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<emphasis bold="true" pageId="16" pageNumber="17">Myanmar</emphasis>
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: 1 male (NHML), "
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<emphasis italics="true" pageId="16" pageNumber="17">Toungoo</emphasis>
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"; "
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<emphasis italics="true" pageId="16" pageNumber="17">Anchista</emphasis>
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/
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<emphasis italics="true" pageId="16" pageNumber="17">sp. n</emphasis>
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."; "H. E. Andrewes Coll. / B. M. 1945-97"; "PARATYPE ♂/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label].
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<emphasis bold="true" pageId="16" pageNumber="17">Cambodia</emphasis>
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: 1 female (MNHN), "Cambodge /
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<normalizedToken originalValue="Rég">Reg</normalizedToken>
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. De Chiehreng / G. Thomas 1912.";
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<normalizedToken originalValue="“Muséum">"Museum</normalizedToken>
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Paris / Coll. R.
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<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
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/ 1952"; "PARATYPE ♀/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label].
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<emphasis bold="true" pageId="16" pageNumber="17">China</emphasis>
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: 1 female (HBUM), "2006-XI-15-16 /
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<normalizedToken originalValue="海南昌江县尖峰岭">海南昌江县尖峰岭</normalizedToken>
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/
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<normalizedToken originalValue="任国栋">任国栋</normalizedToken>
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/
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<normalizedToken originalValue="河北大学博物馆”">河北大学博物馆"</normalizedToken>
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[2006-XI-15-16, Hainan, Changjiang, Jianfengling, Ren Guodong leg., Hebei University Museum]; "PARATYPE ♀/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label][Figs 112, 127]. 1 female (IZAS), "采集地：海南五指山 /
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<normalizedToken originalValue="海拔">海拔</normalizedToken>
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<normalizedToken originalValue="日期">日期</normalizedToken>
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2008-
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<emphasis italics="true" pageId="16" pageNumber="17">IV-03</emphasis>
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/
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<normalizedToken originalValue="方法">方法</normalizedToken>
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<normalizedToken originalValue="灯诱">灯诱</normalizedToken>
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<normalizedToken originalValue="采集人：杨玉霞”">采集人：杨玉霞"</normalizedToken>
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[Hainan, Wuzhishan, 2008.IV.03. by light trap, Yang Yuxia leg.]; "PARATYPE ♀/ Paraphaea minor / new species / Des. SHI H. L. 2011" [red label][Fig. 3].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph pageId="16" pageNumber="17">Diagnosis.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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Pronotum glabrous, widest at middle, lateral margins sinuate before hind angles; elytra reddish yellow with dark lateral and apical margins; two setigerous pores present on 3rd interval and one setigerous pore on base of 5th interval; median lobe of aedeagus with apical third slightly expanded, internal sac with sinuous main flagellum (
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<figureCitation captionStart="Figure 69" captionStartId="F14" captionText="Figure 69. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Paraphaea minor sp. n., holotype." httpUri="https://binary.pensoft.net/fig/29687" pageId="16" pageNumber="17">Fig. 69</figureCitation>
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).
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</paragraph>
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<paragraph lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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The new species is most similar to
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1946" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis italics="true" pageId="16" pageNumber="17">Paraphaea formosana</emphasis>
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</taxonomicName>
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in male genitalia. It differs as follows: (1) 3rd interval with two setigerous pores in
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<taxonomicName authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minor">
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<emphasis italics="true" pageId="16" pageNumber="17">Paraphaea minor</emphasis>
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</taxonomicName>
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, but with three or four pores in
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1946" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis italics="true" pageId="16" pageNumber="17">Paraphaea formosana</emphasis>
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</taxonomicName>
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; (2) elytral patterns of these two species are different; (3)
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<taxonomicName authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minor">
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<emphasis italics="true" pageId="16" pageNumber="17">Paraphaea minor</emphasis>
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</taxonomicName>
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with median lobe of aedeagus less expanded subapically, left-lateral margin only slightly sin
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<pageBreakToken pageId="17" pageNumber="18" start="start">uate</pageBreakToken>
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in dorsal view (
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<figureCitation captionStart="Figure 69" captionStartId="F14" captionText="Figure 69. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Paraphaea minor sp. n., holotype." httpUri="https://binary.pensoft.net/fig/29687" pageId="17" pageNumber="18">Fig. 69</figureCitation>
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);
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1946" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis italics="true" pageId="17" pageNumber="18">Paraphaea formosana</emphasis>
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</taxonomicName>
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with left-lateral margin of aedeagal median lobe more distinctly sinuate in dorsal view (
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<figureCitation captionStart="Figure 68. " captionStartId="F13" captionText="Figure 68. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Paraphaea formosana (Jedlicka), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29686" pageId="17" pageNumber="18">Fig. 68</figureCitation>
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); (4)
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<taxonomicName authorityName="Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="minor">
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<emphasis italics="true" pageId="17" pageNumber="18">Paraphaea minor</emphasis>
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</taxonomicName>
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with apical lamella completely rounded, very short, apex wide;
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1946" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis italics="true" pageId="17" pageNumber="18">Paraphaea formosana</emphasis>
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</taxonomicName>
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with apical lamella somewhat triangular, much longer, apex narrower.
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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This new species maybe confused with
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<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="binotata">
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<emphasis italics="true" pageId="17" pageNumber="18">Paraphaea binotata</emphasis>
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</taxonomicName>
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, but can be distinguished by different elytral pattern, and pronotum widest at middle, not at apical third.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="description">
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<paragraph pageId="17" pageNumber="18">Description.</paragraph>
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<paragraph pageId="17" pageNumber="18">
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Body length usually 5.8-6.9 mm (specimens from Hainan length 7.6-7.9 mm); head and pronotum reddish yellow to brown, lateral explanate areas of pronotum paler; mouthparts, antennae and legs yellowish brown or reddish brown; elytra reddish yellow, with dark marginal fascia occupying 7th-9th intervals, fascia conjoined on elytral apex but not on base, elytral lateral margins yellowish brown; ventral side yellowish brown to dark brown.
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<emphasis bold="true" pageId="17" pageNumber="18">Head</emphasis>
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glabrous, without punctures or microsculpture.
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<emphasis bold="true" pageId="17" pageNumber="18">Pronotum</emphasis>
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cordiform, widest near middle; ratio PW/PL 1.38 to 1.52; pronotal base briefly but distinctly lobed; disc glabrous, microsculpture indistinct, without punctures; front angles with a few short setae; lateral margins completely rounded in middle, sinuate before hind angles; front angles wide and round, not projected; hind angles sharp, rectangular or nearly so, slightly pointed; disc moderately convex; basal foveae wide and even, with a few fine punctures.
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<emphasis bold="true" pageId="17" pageNumber="18">Elytra</emphasis>
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with striae distinct, finely punctate; intervals slightly convex, without punctures or microsculpture; 3rd interval with two setigerous pores: basal one placed approximately basal one-third, adjacent to 3rd stria; apical one placed approximately apical one-eighth, adjacent to 2nd stria or in middle of interval; 5th interval with one setigerous pore near base, adjacent to 5th stria; 15-17 umbilical pores on 9th interval.
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<emphasis bold="true" pageId="17" pageNumber="18">Male genitalia.</emphasis>
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Median lobe of aedeagus with apical third slightly expanded; in dorsal view, left-lateral margin moderately sinuate medially, and then gradually narrowed to apex; apical lamella placed on left-ventral side, broadly rounded, not prolongate; internal sac with main flagellum slightly sinuous; basal part of main flagellum moderately bent, so trumpet-form expansion adjacent to left margin, apical margin of trumpet-form expansion not crenulate (
|
||
<figureCitation captionStart="Figure 69" captionStartId="F14" captionText="Figure 69. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Paraphaea minor sp. n., holotype." httpUri="https://binary.pensoft.net/fig/29687" pageId="17" pageNumber="18">Fig. 69</figureCitation>
|
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).
|
||
<emphasis bold="true" pageId="17" pageNumber="18">Female genitalia</emphasis>
|
||
. Spermatheca slender and strongly bent near middle, but not so elongate as in
|
||
<taxonomicName authorityName="Shi & Zhou & Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="binotata">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Paraphaea binotata</emphasis>
|
||
</taxonomicName>
|
||
; spermathecal gland inserted near apical one-third of spermatheca, distinctly expanded at basal part, not bent; spermatheca with ring-sculpture between apex and the gland insertion; spermatheca strongly bent at apical one-third (
|
||
<figureCitation captionStart="Figures 126–131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="17" pageNumber="18">Fig. 127</figureCitation>
|
||
). Apical segment of ovipositor scimitar-shaped, inner margin slightly angulate at apical one-third; length about three times basal width; inner margin setose in apical half; apex slightly widened, with membranous extension long and slender (
|
||
<figureCitation captionStart="Figures 111–125" captionStartId="F41" captionText="Figures 111 - 125. Left ovipositors of females, ventral view, scale bars = 0.2 mm: 111 Paraphaea binotata (Dejean), a specimen from Guangxi 112 Paraphaea minor sp. n., a paratype from Hainan 113 Paraphaea philippinensis (Jedlicka), holotype 114 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 115 Physodera dejeani Eschscholtz, a specimen from Yunnan 116 Physodera eschscholtzii Parry, a specimen from Yunnan 117 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 118 Allocota aurata (Bates), a specimen from Shaanxi 119 Allocota bicolor sp. n., a paratype from Guangdong 120 Lachnoderma asperum Bates, a specimen from Zhejiang 121 Dasiosoma bellum (Habu), a specimen from Guangdong 122 Dasiosoma maindroni (Tian & Deuve), a specimen from Tonkin 123 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 124 Orionella lewisii (Bates), a specimen from Zhejiang 125 Endynomena pradieri (Fairmaire), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29719" pageId="17" pageNumber="18">Fig. 112</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph pageId="17" pageNumber="18">Distribution</paragraph>
|
||
<paragraph pageId="17" pageNumber="18">
|
||
(
|
||
<figureCitation captionStart="Map 2" captionStartId="F46" captionText="Map 2. Known distribution of genus Paraphaea II: ▲ Paraphaea minor sp. n. ◆ Paraphaea philippinensis (Jedlicka) ★ Paraphaea formosana (Jedlicka)." httpUri="https://binary.pensoft.net/fig/29724" pageId="17" pageNumber="18">Map 2</figureCitation>
|
||
). China (Hainan), Vietnam, Thailand, Myanmar, Cambodia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph pageId="17" pageNumber="18">Etymology.</paragraph>
|
||
<paragraph pageId="17" pageNumber="18">
|
||
This species is the smallest one of the genus, so we name it "
|
||
<emphasis italics="true" pageId="17" pageNumber="18">minor</emphasis>
|
||
", a word from Latin meaning
|
||
<normalizedToken originalValue="“small”">"small"</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="18" type="geographical variation">
|
||
<paragraph pageId="17" pageNumber="18">Geographical variation.</paragraph>
|
||
<paragraph pageId="17" pageNumber="18">
|
||
Two females from Hainan (China) (
|
||
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="17" pageNumber="18">Fig. 3</figureCitation>
|
||
) are distinctly larger than the other specimens from South East Asia (7.6-7.9 mm contrasting with 5.8-6.9 mm). But there is no other difference between them and other paratypes from Tonkin in external and female genitalia characters, so we include these two females in type series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |