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<document id="B26FE5D97B36D6D82A1AF73B2EE2D5FF" ID-DOI="10.1093/zoolinnean/zlad113" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240975" ID-ZooBank="3BD9090F-DFD4-47B0-9CA8-E46CE67D4B98" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1716363357279" checkinUser="plazi" docAuthor="Lepeco, Anderson &amp; Melo, Gabriel A. R." docDate="2024" docId="A93501368459BB2FFF6FFA5DFB857F60" docLanguage="en" docName="zlad113.pdf" docOrigin="Zoological Journal of the Linnean Society 201" docSource="https://www.mendeley.com/catalogue/d31a8eb3-d55c-3269-8d9a-4fae8ddd621b/" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Chrysopsenella Lepeco and Melo 2024, gen. nov." docType="treatment" docVersion="2" lastPageNumber="66" masterDocId="550C794E8451BB26FFDBFFFBFFCA7A20" masterDocTitle="Revisiting the phylogeny of the scolebythid wasps (Hymenoptera: Aculeata) through Bayesian model evaluation and parsimony, with description of a new fossil family of Chrysidoidea" masterLastPageNumber="85" masterPageNumber="57" pageNumber="65" updateTime="1716499870449" updateUser="ExternalLinkService">
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<mods:title id="8CBC490C4204486829FA084E7970E133">Revisiting the phylogeny of the scolebythid wasps (Hymenoptera: Aculeata) through Bayesian model evaluation and parsimony, with description of a new fossil family of Chrysidoidea</mods:title>
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<mods:namePart id="E1A25189D1D77AB1CA988ADFCE1100FE">Lepeco, Anderson</mods:namePart>
<mods:affiliation id="19AC3FD0BCEDB1296B4C49F84E52F5AC">Laboratório de Biologia Comparada e Abelhas, Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. 14040 - 901, Ribeirão Preto, SP, Brazil</mods:affiliation>
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<mods:namePart id="DE2022DAF04597474C56645A0D88C678">Melo, Gabriel A. R.</mods:namePart>
<mods:affiliation id="420DB12DE5EF48E4FF5C3A4D40C64265">Laboratório de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Setor de Ciências Biológicas, Universidade Federal do Paraná. 81531 - 980, Curitiba, PR, Brazil</mods:affiliation>
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<mods:date id="A3E85A579CAB56575979D20D8710DB5C">2024</mods:date>
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<subSubSection id="6986E3AB8459BB2EFF6FFA5DFD1F7FE0" box="[180,725,1445,1472]" pageId="8" pageNumber="65" type="nomenclature">
<paragraph id="2123B0208459BB2EFF6FFA5DFD1F7FE0" blockId="8.[180,725,1445,1472]" box="[180,725,1445,1472]" pageId="8" pageNumber="65">
<heading id="7A6B074C8459BB2EFF6FFA5DFD1F7FE0" bold="true" box="[180,725,1445,1472]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="65" reason="3">
<emphasis id="13E86C328459BB2EFF6FFA5DFD1F7FE0" bold="true" box="[180,725,1445,1472]" pageId="8" pageNumber="65">
Genus † 
<taxonomicName id="E69CCBA38459BB2EFED1FA5EFDBA7FE0" authority="Lepeco and Melo" authorityName="Lepeco and Melo" authorityYear="2024" box="[266,624,1445,1472]" class="Insecta" family="Scolebythidae" genus="Chrysopsenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13E86C328459BB2EFED1FA5EFE7E7FE0" bold="true" box="[266,436,1445,1472]" italics="true" pageId="8" pageNumber="65">Chrysopsenella</emphasis>
Lepeco and Melo
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, 
<taxonomicNameLabel id="08DBD1498459BB2EFDA1FA5DFD1F7FE0" box="[634,725,1446,1472]" pageId="8" pageNumber="65" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="6986E3AB8459BB2EFF5AFA26FF7B7C33" pageId="8" pageNumber="65" type="type_taxon">
<paragraph id="2123B0208459BB2EFF5AFA26FF7B7C33" blockId="8.[129,778,1500,1555]" pageId="8" pageNumber="65">
<emphasis id="13E86C328459BB2EFF5AFA26FF347FD4" box="[129,254,1501,1524]" italics="true" pageId="8" pageNumber="65">
<typeStatus id="FE270E828459BB2EFF5AFA26FF7B7FD4" box="[129,177,1501,1524]" pageId="8" pageNumber="65">Type</typeStatus>
species:
</emphasis>
† 
<taxonomicName id="E69CCBA38459BB2EFEC8FA27FD297FD4" authority="Lepeco and Melo" authorityName="Lepeco and Melo" authorityYear="2024" box="[275,739,1500,1524]" class="Insecta" family="Scolebythidae" genus="Chrysopsenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="euryphaessa" status="sp. nov.">
<emphasis id="13E86C328459BB2EFEC8FA27FDE97FD4" box="[275,547,1500,1524]" italics="true" pageId="8" pageNumber="65">Chrysopsenella euryphaessa</emphasis>
Lepeco and Melo
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, 
<taxonomicNameLabel id="08DBD1498459BB2EFD35FA26FF6D7C33" pageId="8" pageNumber="65" rank="species">sp. nov</taxonomicNameLabel>
..
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<subSubSection id="6986E3AB8459BB2FFF5AF9CFFAA37EE8" lastPageId="9" lastPageNumber="66" pageId="8" pageNumber="65" type="description">
<paragraph id="2123B0208459BB2FFF5AF9CFFAA37EE8" blockId="8.[129,778,1587,1987]" lastBlockId="9.[810,1460,1044,1225]" lastPageId="9" lastPageNumber="66" pageId="8" pageNumber="65">
<emphasis id="13E86C328459BB2EFF5AF9CFFF3D7C6B" box="[129,247,1588,1611]" italics="true" pageId="8" pageNumber="65">Description:</emphasis>
Head globose, round in frontal view, obliquely hypognathous. Maxillary palp with six palpomeres. Labial palp with four palpomeres. Palps relatively short, not reaching the occipital region. Mandibles relatively long, with at least two apical teeth, apicalmost tooth shorter than lower one. Clypeus mostly transverse, short, strongly emarginated by antennal sockets; apex rounded medially, not forming lobe below basal surface level; longitudinal carina absent. Malar space short, about as long as one-third of mandible basal width; without sulcus. Antennal socket directed anteriorly, its lower margin level with clypeal apex; upper portion of antennal rim slightly projected over radicle. Compound eye well-developed, oval, not bulging, occupying most of the head in lateral view; orbits slightly converging dorsally. Ocelli present, organized in a nearly equilateral triangle. Frons gently convex, slightly protuberant above antennal sockets medially; without conspicuous medial line, halves not deepening along midline. Hypostomal bridge as long as 1.5× basal width of mandible; anterior edge as a deep concavity ending at mandibular bases. Occipital carina present, reaching ventral surface of head. Vertex short, widely round in frontal view. Antenna: radicle–scape axis straight; scape relatively long and cylindrical; pedicel clavate; flagellum relatively short, with 11 cylindrical flagellomeres. Pronotum with short pronotal collar, its posterior margin widely incurved, reaching tegula laterally, transition with anterior surface continuous; transverse sulcus on anterior surface present; pronotal flange well developed, preceded by transverse carina; postero-ventral pronotal angle rounded, not projecting towards procoxa. Propleura entirely covered by pronotum in dorsal view; dorsolateral carina with edge directed ventrally; posterior margins diverging ventrally; epicoxal lobes present, small. Prosternum with relatively large basisternum, which is diamond-shaped; deepening along midline; posteriorly depressed in relation to propleuron. Mesoscutum longer than pronotum; notauli deeply impressed, strongly diverging anteriorly; parascutal edges not carinate. Scutellum longer than mesoscutum, with a deep transverse sulcus anteriorly; axilla relatively large. Mesepisternum transversed by a sulcus coming from the prepectus to the mesepisternal pit; prepectus well developed and forming two separate plates, apparently not contacting each other medially; mesocoxal foramen large, wider than half of the posterior surface of mesepisternum, articulation with mesocoxa apparent ventrally. Metanotum short, not covered by scutellum, with shallow sublateral pits. Metepisternum with lateral surfaces separated from its ventral portion by transverse carina; ventrally narrowed, not forming plate between mesocoxae. Metapostnotum forming basal transverse depression, narrowly extending onto propodeum medially. Propodeum relatively short; transition between dorsal and posterior surfaces continuous; spiracle separated from metapleural line by about one-half of its width; posterior surface with longitudinal carina above foramen medially; and with a lateral carina above metacoxa, separating the posterior and lateral surfaces. Propodeal foramen separated from metacoxal foramina by sclerotized bridge. Legs: procoxa somewhat reniform-shaped, inner surface rounded; protrochanter inserted on apico-lateral region of coxa; profemur thicker at midlength, somewhat spindle-shaped; protibial spur with vellum entire, not forming pecten; tarsal claws bidentate, with minute preapical tooth, basal expansions small. Mesofemur without apical expansion to accommodate tibia; meso- and metatibiae with two apical spurs. Metacoxa with strong posterior carina. Forewing with four cells enclosed by tubular veins: costal, radial, 1st cubital, and 1st medial; tubular portion of 
<collectionCode id="478D28E58458BB2FFF55F93EFF697CFD" box="[142,163,1733,1757]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="66">R</collectionCode>
1 becoming nebulous beyond midlength of marginal cell; distal portion of Rs ending very near anterior margin of wing, leaving marginal cell open distally; 2rs-m nebulous; Cu2 not reaching distal portion of A, leaving 2nd cubital cell open distally. Hindwing with venation reduced to Rs + M and A; jugal lobe absent. Metasoma about as long as lengths of head and mesosoma combined. Tergum 1 with dorsal surface gradually slanting towards anterior one; anterior surface with a shallow depression medially; lateral line absent; posterior margin almost straight, interrupted by a deep cleft at the angle between lateral and dorsal surfaces of sclerite. Sternum 1 with a median protuberance; posterior margin without medial reentrance. Posterior segments not retractable within anterior portion of metasoma. Tergum 7 sclerotized and exposed. Sting well developed.
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<emphasis id="13E86C328458BB2FFFAAFF74FF757A86" bold="true" box="[113,191,142,167]" pageId="9" pageNumber="66">Table 2.</emphasis>
Results of steppingstone analyses for the 20 different models applied. Highlighted in bold is the model that retrieved the highest MgL. APRV = among-partition rate variation. ACRV = among-character rate variation. KRS refers to the Kass and Raftery statistic computed with respect to the best model overall.
</paragraph>
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<paragraph id="2123B0208458BB2FFFAAFF0EFA7979E0" pageId="9" pageNumber="66">
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<emphasis id="13E86C328458BB2FFFAAFF0EFF0E7B2D" bold="true" box="[113,196,245,269]" pageId="9" pageNumber="66">Analysis</emphasis>
</th>
<th id="DC7DDB1E845844D9FF32FF0EFE217B2D" box="[233,491,245,269]" gridcol="1" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFF32FF0EFE817B2D" bold="true" box="[233,331,245,269]" pageId="9" pageNumber="66">File name</emphasis>
</th>
<th id="DC7DDB1E845844D9FDD5FF0EFD1D7B2D" box="[526,727,245,269]" gridcol="2" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFDD5FF0EFD1D7B2D" bold="true" box="[526,727,245,269]" pageId="9" pageNumber="66">Partitioning scheme</emphasis>
</th>
<th id="DC7DDB1E845844D9FD20FF0EFC2C7B2D" box="[763,998,245,269]" gridcol="3" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFD20FF0EFCF37B2D" bold="true" box="[763,825,245,269]" pageId="9" pageNumber="66">APRV</emphasis>
</th>
<th id="DC7DDB1E845844D9FBD2FF0EFB067B2D" box="[1033,1228,245,269]" gridcol="4" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFBD2FF0EFB837B2D" bold="true" box="[1033,1097,245,269]" pageId="9" pageNumber="66">ACRV</emphasis>
</th>
<th id="DC7DDB1E845844D9FB34FF0EFA847B2D" box="[1263,1358,245,269]" gridcol="5" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFB34FF0EFAD57B2D" bold="true" box="[1263,1311,245,269]" pageId="9" pageNumber="66">MgL</emphasis>
</th>
<th id="DC7DDB1E845844D9FAAAFF0EFA797B2D" box="[1393,1459,245,269]" gridcol="6" gridrow="0" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFAAAFF0EFA557B2D" bold="true" box="[1393,1439,245,269]" pageId="9" pageNumber="66">KRS</emphasis>
</th>
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<tr id="9FACB262845844D9FFAAFEDEFA797B1D" box="[113,1459,293,317]" gridrow="1" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFEDEFF0E7B1D" box="[113,196,293,317]" gridcol="0" gridrow="1" pageId="9" pageNumber="66">1</th>
<td id="DC7DDB1E845844D9FF32FEDEFE217B1D" box="[233,491,293,317]" gridcol="1" gridrow="1" pageId="9" pageNumber="66">1_unp_equal</td>
<td id="DC7DDB1E845844D9FDD5FEDEFD1D7B1D" box="[526,727,293,317]" gridcol="2" gridrow="1" pageId="9" pageNumber="66">Unpartitioned</td>
<td id="DC7DDB1E845844D9FD20FEDEFC2C7B1D" box="[763,998,293,317]" gridcol="3" gridrow="1" pageId="9" pageNumber="66">N/A</td>
<td id="DC7DDB1E845844D9FBD2FEDEFB067B1D" box="[1033,1228,293,317]" gridcol="4" gridrow="1" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FEDEFA847B1D" box="[1263,1358,293,317]" gridcol="5" gridrow="1" pageId="9" pageNumber="66">–1698.4</td>
<td id="DC7DDB1E845844D9FAAAFEDEFA797B1D" box="[1393,1459,293,317]" gridcol="6" gridrow="1" pageId="9" pageNumber="66">275.64</td>
</tr>
<tr id="9FACB262845844D9FFAAFEBCFA797B7F" box="[113,1459,327,351]" gridrow="2" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFEBCFF0E7B7F" box="[113,196,327,351]" gridcol="0" gridrow="2" pageId="9" pageNumber="66">2</th>
<td id="DC7DDB1E845844D9FF32FEBCFE217B7F" box="[233,491,327,351]" gridcol="1" gridrow="2" pageId="9" pageNumber="66">2_unp_shared</td>
<td id="DC7DDB1E845844D9FDD5FEBCFD1D7B7F" box="[526,727,327,351]" gridcol="2" gridrow="2" pageId="9" pageNumber="66">Unpartitioned</td>
<td id="DC7DDB1E845844D9FD20FEBCFC2C7B7F" box="[763,998,327,351]" gridcol="3" gridrow="2" pageId="9" pageNumber="66">N/A</td>
<td id="DC7DDB1E845844D9FBD2FEBCFB067B7F" box="[1033,1228,327,351]" gridcol="4" gridrow="2" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FEBCFA847B7F" box="[1263,1358,327,351]" gridcol="5" gridrow="2" pageId="9" pageNumber="66">–1657.45</td>
<td id="DC7DDB1E845844D9FAAAFEBCFA797B7F" box="[1393,1459,327,351]" gridcol="6" gridrow="2" pageId="9" pageNumber="66">193.74</td>
</tr>
<tr id="9FACB262845844D9FFAAFE92FA797BA1" box="[113,1459,361,385]" gridrow="3" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFE92FF0E7BA1" box="[113,196,361,385]" gridcol="0" gridrow="3" pageId="9" pageNumber="66">3</th>
<td id="DC7DDB1E845844D9FF32FE92FE217BA1" box="[233,491,361,385]" gridcol="1" gridrow="3" pageId="9" pageNumber="66">3_hom_fixed_equal</td>
<td id="DC7DDB1E845844D9FDD5FE92FD1D7BA1" box="[526,727,361,385]" gridcol="2" gridrow="3" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FE92FC2C7BA1" box="[763,998,361,385]" gridcol="3" gridrow="3" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FE92FB067BA1" box="[1033,1228,361,385]" gridcol="4" gridrow="3" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FE92FA847BA1" box="[1263,1358,361,385]" gridcol="5" gridrow="3" pageId="9" pageNumber="66">–1698.39</td>
<td id="DC7DDB1E845844D9FAAAFE92FA797BA1" box="[1393,1459,361,385]" gridcol="6" gridrow="3" pageId="9" pageNumber="66">275.62</td>
</tr>
<tr id="9FACB262845844D9FFAAFE71FA797B83" box="[113,1459,394,419]" gridrow="4" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFE71FF0E7B83" box="[113,196,394,419]" gridcol="0" gridrow="4" pageId="9" pageNumber="66">4</th>
<td id="DC7DDB1E845844D9FF32FE71FE217B83" box="[233,491,394,419]" gridcol="1" gridrow="4" pageId="9" pageNumber="66">4_hom_fixed_shared</td>
<td id="DC7DDB1E845844D9FDD5FE71FD1D7B83" box="[526,727,394,419]" gridcol="2" gridrow="4" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FE71FC2C7B83" box="[763,998,394,419]" gridcol="3" gridrow="4" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FE71FB067B83" box="[1033,1228,394,419]" gridcol="4" gridrow="4" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FE71FA847B83" box="[1263,1358,394,419]" gridcol="5" gridrow="4" pageId="9" pageNumber="66">–1657.35</td>
<td id="DC7DDB1E845844D9FAAAFE71FA797B83" box="[1393,1459,394,419]" gridcol="6" gridrow="4" pageId="9" pageNumber="66">193.54</td>
</tr>
<tr id="9FACB262845844D9FFAAFE57FA797BE5" box="[113,1459,428,453]" gridrow="5" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFE57FF0E7BE5" box="[113,196,428,453]" gridcol="0" gridrow="5" pageId="9" pageNumber="66">5</th>
<td id="DC7DDB1E845844D9FF32FE57FE217BE5" box="[233,491,428,453]" gridcol="1" gridrow="5" pageId="9" pageNumber="66">5_hom_fixed_per</td>
<td id="DC7DDB1E845844D9FDD5FE57FD1D7BE5" box="[526,727,428,453]" gridcol="2" gridrow="5" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FE57FC2C7BE5" box="[763,998,428,453]" gridcol="3" gridrow="5" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FE57FB067BE5" box="[1033,1228,428,453]" gridcol="4" gridrow="5" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FE57FA847BE5" box="[1263,1358,428,453]" gridcol="5" gridrow="5" pageId="9" pageNumber="66">–1635.91</td>
<td id="DC7DDB1E845844D9FAAAFE57FA797BE5" box="[1393,1459,428,453]" gridcol="6" gridrow="5" pageId="9" pageNumber="66">150.66</td>
</tr>
<tr id="9FACB262845844D9FFAAFE35FA797BC6" box="[113,1459,462,486]" gridrow="6" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFE35FF0E7BC6" box="[113,196,462,486]" gridcol="0" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFFA6FE35FF407BC6" bold="true" box="[125,138,462,486]" pageId="9" pageNumber="66">6</emphasis>
</th>
<td id="DC7DDB1E845844D9FF32FE35FE217BC6" box="[233,491,462,486]" gridcol="1" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFF31FE35FE097BC6" bold="true" box="[234,451,462,486]" pageId="9" pageNumber="66">6_hom_linked_equal</emphasis>
</td>
<td id="DC7DDB1E845844D9FDD5FE35FD1D7BC6" box="[526,727,462,486]" gridcol="2" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFDD4FE35FD4E7BC6" bold="true" box="[527,644,462,486]" pageId="9" pageNumber="66">Homoplasy</emphasis>
</td>
<td id="DC7DDB1E845844D9FD20FE35FC2C7BC6" box="[763,998,462,486]" gridcol="3" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFD27FE35FC157BC6" bold="true" box="[764,991,462,486]" pageId="9" pageNumber="66">Linked branch lengths</emphasis>
</td>
<td id="DC7DDB1E845844D9FBD2FE35FB067BC6" box="[1033,1228,462,486]" gridcol="4" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFBD2FE35FBB07BC6" bold="true" box="[1033,1146,462,486]" pageId="9" pageNumber="66">Equal rates</emphasis>
</td>
<td id="DC7DDB1E845844D9FB34FE35FA847BC6" box="[1263,1358,462,486]" gridcol="5" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFB2BFE35FA897BC6" bold="true" box="[1264,1347,462,486]" pageId="9" pageNumber="66">–1560.6</emphasis>
</td>
<td id="DC7DDB1E845844D9FAAAFE35FA797BC6" box="[1393,1459,462,486]" gridcol="6" gridrow="6" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFA52FE35FA5C7BC6" bold="true" box="[1417,1430,462,486]" pageId="9" pageNumber="66">0</emphasis>
</td>
</tr>
<tr id="9FACB262845844D9FFAAFE0BFA797829" box="[113,1459,496,521]" gridrow="7" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFE0BFF0E7829" box="[113,196,496,521]" gridcol="0" gridrow="7" pageId="9" pageNumber="66">7</th>
<td id="DC7DDB1E845844D9FF32FE0BFE217829" box="[233,491,496,521]" gridcol="1" gridrow="7" pageId="9" pageNumber="66">7_hom_linked_shared</td>
<td id="DC7DDB1E845844D9FDD5FE0BFD1D7829" box="[526,727,496,521]" gridcol="2" gridrow="7" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FE0BFC2C7829" box="[763,998,496,521]" gridcol="3" gridrow="7" pageId="9" pageNumber="66">Linked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FE0BFB067829" box="[1033,1228,496,521]" gridcol="4" gridrow="7" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FE0BFA847829" box="[1263,1358,496,521]" gridcol="5" gridrow="7" pageId="9" pageNumber="66">–1568.67</td>
<td id="DC7DDB1E845844D9FAAAFE0BFA797829" box="[1393,1459,496,521]" gridcol="6" gridrow="7" pageId="9" pageNumber="66">16.18</td>
</tr>
<tr id="9FACB262845844D9FFAAFDE9FA79780A" box="[113,1459,530,554]" gridrow="8" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFDE9FF0E780A" box="[113,196,530,554]" gridcol="0" gridrow="8" pageId="9" pageNumber="66">8</th>
<td id="DC7DDB1E845844D9FF32FDE9FE21780A" box="[233,491,530,554]" gridcol="1" gridrow="8" pageId="9" pageNumber="66">8_hom_linked_per</td>
<td id="DC7DDB1E845844D9FDD5FDE9FD1D780A" box="[526,727,530,554]" gridcol="2" gridrow="8" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FDE9FC2C780A" box="[763,998,530,554]" gridcol="3" gridrow="8" pageId="9" pageNumber="66">Linked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FDE9FB06780A" box="[1033,1228,530,554]" gridcol="4" gridrow="8" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FDE9FA84780A" box="[1263,1358,530,554]" gridcol="5" gridrow="8" pageId="9" pageNumber="66">–1569.86</td>
<td id="DC7DDB1E845844D9FAAAFDE9FA79780A" box="[1393,1459,530,554]" gridcol="6" gridrow="8" pageId="9" pageNumber="66">18.56</td>
</tr>
<tr id="9FACB262845844D9FFAAFDCFFA79786C" box="[113,1459,564,588]" gridrow="9" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFDCFFF0E786C" box="[113,196,564,588]" gridcol="0" gridrow="9" pageId="9" pageNumber="66">9</th>
<td id="DC7DDB1E845844D9FF32FDCFFE21786C" box="[233,491,564,588]" gridcol="1" gridrow="9" pageId="9" pageNumber="66">9_hom_unlinked_equal</td>
<td id="DC7DDB1E845844D9FDD5FDCFFD1D786C" box="[526,727,564,588]" gridcol="2" gridrow="9" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FDCFFC2C786C" box="[763,998,564,588]" gridcol="3" gridrow="9" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FDCFFB06786C" box="[1033,1228,564,588]" gridcol="4" gridrow="9" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FDCFFA84786C" box="[1263,1358,564,588]" gridcol="5" gridrow="9" pageId="9" pageNumber="66">–1679.07</td>
<td id="DC7DDB1E845844D9FAAAFDCFFA79786C" box="[1393,1459,564,588]" gridcol="6" gridrow="9" pageId="9" pageNumber="66">236.98</td>
</tr>
<tr id="9FACB262845844D9FFAAFDAEFA79784E" box="[113,1459,597,622]" gridrow="10" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFDAEFF0E784E" box="[113,196,597,622]" gridcol="0" gridrow="10" pageId="9" pageNumber="66">10</th>
<td id="DC7DDB1E845844D9FF32FDAEFE21784E" box="[233,491,597,622]" gridcol="1" gridrow="10" pageId="9" pageNumber="66">10_hom_unlinked_shared</td>
<td id="DC7DDB1E845844D9FDD5FDAEFD1D784E" box="[526,727,597,622]" gridcol="2" gridrow="10" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FDAEFC2C784E" box="[763,998,597,622]" gridcol="3" gridrow="10" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FDAEFB06784E" box="[1033,1228,597,622]" gridcol="4" gridrow="10" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FDAEFA84784E" box="[1263,1358,597,622]" gridcol="5" gridrow="10" pageId="9" pageNumber="66">–1686.81</td>
<td id="DC7DDB1E845844D9FAAAFDAEFA79784E" box="[1393,1459,597,622]" gridcol="6" gridrow="10" pageId="9" pageNumber="66">252.46</td>
</tr>
<tr id="9FACB262845844D9FFAAFD8CFA7978B0" box="[113,1459,631,656]" gridrow="11" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFD8CFF0E78B0" box="[113,196,631,656]" gridcol="0" gridrow="11" pageId="9" pageNumber="66">11</th>
<td id="DC7DDB1E845844D9FF32FD8CFE2178B0" box="[233,491,631,656]" gridcol="1" gridrow="11" pageId="9" pageNumber="66">11_hom_unlinked_per</td>
<td id="DC7DDB1E845844D9FDD5FD8CFD1D78B0" box="[526,727,631,656]" gridcol="2" gridrow="11" pageId="9" pageNumber="66">Homoplasy</td>
<td id="DC7DDB1E845844D9FD20FD8CFC2C78B0" box="[763,998,631,656]" gridcol="3" gridrow="11" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FD8CFB0678B0" box="[1033,1228,631,656]" gridcol="4" gridrow="11" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FD8CFA8478B0" box="[1263,1358,631,656]" gridcol="5" gridrow="11" pageId="9" pageNumber="66">–1689.06</td>
<td id="DC7DDB1E845844D9FAAAFD8CFA7978B0" box="[1393,1459,631,656]" gridcol="6" gridrow="11" pageId="9" pageNumber="66">256.96</td>
</tr>
<tr id="9FACB262845844D9FFAAFD62FA797891" box="[113,1459,665,689]" gridrow="12" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFD62FF0E7891" box="[113,196,665,689]" gridcol="0" gridrow="12" pageId="9" pageNumber="66">12</th>
<td id="DC7DDB1E845844D9FF32FD62FE217891" box="[233,491,665,689]" gridcol="1" gridrow="12" pageId="9" pageNumber="66">12_ana_fixed_equal</td>
<td id="DC7DDB1E845844D9FDD5FD62FD1D7891" box="[526,727,665,689]" gridcol="2" gridrow="12" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FD62FC2C7891" box="[763,998,665,689]" gridcol="3" gridrow="12" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FD62FB067891" box="[1033,1228,665,689]" gridcol="4" gridrow="12" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FD62FA847891" box="[1263,1358,665,689]" gridcol="5" gridrow="12" pageId="9" pageNumber="66">–1698.33</td>
<td id="DC7DDB1E845844D9FAAAFD62FA797891" box="[1393,1459,665,689]" gridcol="6" gridrow="12" pageId="9" pageNumber="66">275.5</td>
</tr>
<tr id="9FACB262845844D9FFAAFD40FA7978F4" box="[113,1459,699,724]" gridrow="13" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFD40FF0E78F4" box="[113,196,699,724]" gridcol="0" gridrow="13" pageId="9" pageNumber="66">13</th>
<td id="DC7DDB1E845844D9FF32FD40FE2178F4" box="[233,491,699,724]" gridcol="1" gridrow="13" pageId="9" pageNumber="66">13_ana_fixed_shared</td>
<td id="DC7DDB1E845844D9FDD5FD40FD1D78F4" box="[526,727,699,724]" gridcol="2" gridrow="13" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FD40FC2C78F4" box="[763,998,699,724]" gridcol="3" gridrow="13" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FD40FB0678F4" box="[1033,1228,699,724]" gridcol="4" gridrow="13" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FD40FA8478F4" box="[1263,1358,699,724]" gridcol="5" gridrow="13" pageId="9" pageNumber="66">–1657.47</td>
<td id="DC7DDB1E845844D9FAAAFD40FA7978F4" box="[1393,1459,699,724]" gridcol="6" gridrow="13" pageId="9" pageNumber="66">193.78</td>
</tr>
<tr id="9FACB262845844D9FFAAFD26FA7978D5" box="[113,1459,733,757]" gridrow="14" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFD26FF0E78D5" box="[113,196,733,757]" gridcol="0" gridrow="14" pageId="9" pageNumber="66">14</th>
<td id="DC7DDB1E845844D9FF32FD26FE2178D5" box="[233,491,733,757]" gridcol="1" gridrow="14" pageId="9" pageNumber="66">14_ana_fixed_per</td>
<td id="DC7DDB1E845844D9FDD5FD26FD1D78D5" box="[526,727,733,757]" gridcol="2" gridrow="14" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FD26FC2C78D5" box="[763,998,733,757]" gridcol="3" gridrow="14" pageId="9" pageNumber="66">Fixed branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FD26FB0678D5" box="[1033,1228,733,757]" gridcol="4" gridrow="14" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FD26FA8478D5" box="[1263,1358,733,757]" gridcol="5" gridrow="14" pageId="9" pageNumber="66">–1656.27</td>
<td id="DC7DDB1E845844D9FAAAFD26FA7978D5" box="[1393,1459,733,757]" gridcol="6" gridrow="14" pageId="9" pageNumber="66">191.38</td>
</tr>
<tr id="9FACB262845844D9FFAAFD04FA797937" box="[113,1459,767,791]" gridrow="15" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFD04FF0E7937" box="[113,196,767,791]" gridcol="0" gridrow="15" pageId="9" pageNumber="66">15</th>
<td id="DC7DDB1E845844D9FF32FD04FE217937" box="[233,491,767,791]" gridcol="1" gridrow="15" pageId="9" pageNumber="66">15_ana_linked_equal</td>
<td id="DC7DDB1E845844D9FDD5FD04FD1D7937" box="[526,727,767,791]" gridcol="2" gridrow="15" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FD04FC2C7937" box="[763,998,767,791]" gridcol="3" gridrow="15" pageId="9" pageNumber="66">Linked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FD04FB067937" box="[1033,1228,767,791]" gridcol="4" gridrow="15" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FD04FA847937" box="[1263,1358,767,791]" gridcol="5" gridrow="15" pageId="9" pageNumber="66">–1689.48</td>
<td id="DC7DDB1E845844D9FAAAFD04FA797937" box="[1393,1459,767,791]" gridcol="6" gridrow="15" pageId="9" pageNumber="66">257.8</td>
</tr>
<tr id="9FACB262845844D9FFAAFCDAFA797919" box="[113,1459,801,825]" gridrow="16" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFCDAFF0E7919" box="[113,196,801,825]" gridcol="0" gridrow="16" pageId="9" pageNumber="66">16</th>
<td id="DC7DDB1E845844D9FF32FCDAFE217919" box="[233,491,801,825]" gridcol="1" gridrow="16" pageId="9" pageNumber="66">16_ana_linked_shared</td>
<td id="DC7DDB1E845844D9FDD5FCDAFD1D7919" box="[526,727,801,825]" gridcol="2" gridrow="16" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FCDAFC2C7919" box="[763,998,801,825]" gridcol="3" gridrow="16" pageId="9" pageNumber="66">Linked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FCDAFB067919" box="[1033,1228,801,825]" gridcol="4" gridrow="16" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FCDAFA847919" box="[1263,1358,801,825]" gridcol="5" gridrow="16" pageId="9" pageNumber="66">–1648.43</td>
<td id="DC7DDB1E845844D9FAAAFCDAFA797919" box="[1393,1459,801,825]" gridcol="6" gridrow="16" pageId="9" pageNumber="66">175.7</td>
</tr>
<tr id="9FACB262845844D9FFAAFCB9FA79797B" box="[113,1459,834,859]" gridrow="17" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFCB9FF0E797B" box="[113,196,834,859]" gridcol="0" gridrow="17" pageId="9" pageNumber="66">17</th>
<td id="DC7DDB1E845844D9FF32FCB9FE21797B" box="[233,491,834,859]" gridcol="1" gridrow="17" pageId="9" pageNumber="66">17_ana_linked_per</td>
<td id="DC7DDB1E845844D9FDD5FCB9FD1D797B" box="[526,727,834,859]" gridcol="2" gridrow="17" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FCB9FC2C797B" box="[763,998,834,859]" gridcol="3" gridrow="17" pageId="9" pageNumber="66">Linked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FCB9FB06797B" box="[1033,1228,834,859]" gridcol="4" gridrow="17" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FCB9FA84797B" box="[1263,1358,834,859]" gridcol="5" gridrow="17" pageId="9" pageNumber="66">–1647.82</td>
<td id="DC7DDB1E845844D9FAAAFCB9FA79797B" box="[1393,1459,834,859]" gridcol="6" gridrow="17" pageId="9" pageNumber="66">174.48</td>
</tr>
<tr id="9FACB262845844D9FFAAFC9FFA79795C" box="[113,1459,868,892]" gridrow="18" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFC9FFF0E795C" box="[113,196,868,892]" gridcol="0" gridrow="18" pageId="9" pageNumber="66">18</th>
<td id="DC7DDB1E845844D9FF32FC9FFE21795C" box="[233,491,868,892]" gridcol="1" gridrow="18" pageId="9" pageNumber="66">18_ana_unlinked_equal</td>
<td id="DC7DDB1E845844D9FDD5FC9FFD1D795C" box="[526,727,868,892]" gridcol="2" gridrow="18" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FC9FFC2C795C" box="[763,998,868,892]" gridcol="3" gridrow="18" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FC9FFB06795C" box="[1033,1228,868,892]" gridcol="4" gridrow="18" pageId="9" pageNumber="66">Equal rates</td>
<td id="DC7DDB1E845844D9FB34FC9FFA84795C" box="[1263,1358,868,892]" gridcol="5" gridrow="18" pageId="9" pageNumber="66">–1750.54</td>
<td id="DC7DDB1E845844D9FAAAFC9FFA79795C" box="[1393,1459,868,892]" gridcol="6" gridrow="18" pageId="9" pageNumber="66">379.92</td>
</tr>
<tr id="9FACB262845844D9FFAAFC7DFA7979BF" box="[113,1459,902,927]" gridrow="19" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFC7DFF0E79BF" box="[113,196,902,927]" gridcol="0" gridrow="19" pageId="9" pageNumber="66">19</th>
<td id="DC7DDB1E845844D9FF32FC7DFE2179BF" box="[233,491,902,927]" gridcol="1" gridrow="19" pageId="9" pageNumber="66">19_ana_unlinked_shared</td>
<td id="DC7DDB1E845844D9FDD5FC7DFD1D79BF" box="[526,727,902,927]" gridcol="2" gridrow="19" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FC7DFC2C79BF" box="[763,998,902,927]" gridcol="3" gridrow="19" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FC7DFB0679BF" box="[1033,1228,902,927]" gridcol="4" gridrow="19" pageId="9" pageNumber="66">Shared gamma</td>
<td id="DC7DDB1E845844D9FB34FC7DFA8479BF" box="[1263,1358,902,927]" gridcol="5" gridrow="19" pageId="9" pageNumber="66">–1694.15</td>
<td id="DC7DDB1E845844D9FAAAFC7DFA7979BF" box="[1393,1459,902,927]" gridcol="6" gridrow="19" pageId="9" pageNumber="66">267.14</td>
</tr>
<tr id="9FACB262845844D9FFAAFC53FA7979E1" box="[113,1459,936,961]" gridrow="20" pageId="9" pageNumber="66">
<th id="DC7DDB1E845844D9FFAAFC53FF0E79E1" box="[113,196,936,961]" gridcol="0" gridrow="20" pageId="9" pageNumber="66">20</th>
<td id="DC7DDB1E845844D9FF32FC53FE2179E1" box="[233,491,936,961]" gridcol="1" gridrow="20" pageId="9" pageNumber="66">20_ana_unlinked_per</td>
<td id="DC7DDB1E845844D9FDD5FC53FD1D79E1" box="[526,727,936,961]" gridcol="2" gridrow="20" pageId="9" pageNumber="66">Anatomical</td>
<td id="DC7DDB1E845844D9FD20FC53FC2C79E1" box="[763,998,936,961]" gridcol="3" gridrow="20" pageId="9" pageNumber="66">Unlinked branch lengths</td>
<td id="DC7DDB1E845844D9FBD2FC53FB0679E1" box="[1033,1228,936,961]" gridcol="4" gridrow="20" pageId="9" pageNumber="66">Per partition gamma</td>
<td id="DC7DDB1E845844D9FB34FC53FA8479E1" box="[1263,1358,936,961]" gridcol="5" gridrow="20" pageId="9" pageNumber="66">–1694.8</td>
<td id="DC7DDB1E845844D9FAAAFC53FA7979E1" box="[1393,1459,936,961]" gridcol="6" gridrow="20" pageId="9" pageNumber="66">268.44</td>
</tr>
</table>
</paragraph>
<subSubSection id="6986E3AB8458BB2FFCF1FB11FB857F60" pageId="9" pageNumber="66" type="etymology">
<paragraph id="2123B0208458BB2FFCF1FB11FB857F60" blockId="9.[810,1459,1258,1344]" pageId="9" pageNumber="66">
<emphasis id="13E86C328458BB2FFCF1FB11FC537F22" box="[810,921,1258,1282]" italics="true" pageId="9" pageNumber="66">Etymology:</emphasis>
The genus name is based on the combination of the Greek words 
<emphasis id="13E86C328458BB2FFC6AFAF2FC3C7F01" box="[945,1014,1289,1313]" italics="true" pageId="9" pageNumber="66">chrysos</emphasis>
, meaning ‘gold’, and 
<emphasis id="13E86C328458BB2FFB1CFAF2FADE7F01" box="[1223,1300,1289,1313]" italics="true" pageId="9" pageNumber="66">psenella</emphasis>
, the diminutive form of 
<emphasis id="13E86C328458BB2FFCA0FAD2FC6C7F60" box="[891,934,1321,1344]" italics="true" pageId="9" pageNumber="66">psen</emphasis>
, meaning ‘wasp’.
</paragraph>
</subSubSection>
</treatment>
</document>