130 lines
10 KiB
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130 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.302.5357" ID-GBIF-Dataset="7719763b-73d3-41c0-a2a1-7b556211c510" ID-PMC="PMC3689142" ID-Pensoft-Pub="1313-2970-302-13" ID-PubMed="23794898" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-302-13" ModsDocOrigin="ZooKeys 302" ModsDocTitle="Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species" checkinTime="1451247279908" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Stoev, Pavel & Spiegel, Didier Vanden" docDate="2013" docId="A9239820B4BD8CF2486F1913007163E6" docLanguage="en" docName="ZooKeys 302: 13-77" docOrigin="ZooKeys 302" docSource="http://dx.doi.org/10.3897/zookeys.302.5357" docTitle="Opisotretus hagen Golovatch, Geoffroy, Stoev & Spiegel, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="41" masterDocId="E803FFEEFFE9B5412A1DFF9BFFA2CB26" masterDocTitle="Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species" masterLastPageNumber="77" masterPageNumber="13" pageNumber="40" updateTime="1668155849204" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stoev, Pavel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Spiegel, Didier Vanden</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>302</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>13</mods:start>
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<mods:end>77</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.302.5357</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.302.5357</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-302-13</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044251" LSID="urn:lsid:zoobank.org:act:9CD9B717-7849-4122-B437-ABCF2AC79C48" httpUri="http://treatment.plazi.org/id/A9239820B4BD8CF2486F1913007163E6" lastPageId="28" lastPageNumber="41" pageId="27" pageNumber="40">
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<subSubSection pageId="27" pageNumber="40" type="nomenclature">
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<paragraph pageId="27" pageNumber="40">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:9CD9B717-7849-4122-B437-ABCF2AC79C48" class="Diplopoda" family="Opisotretidae" genus="Opisotretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisotretus hagen" order="Polydesmida" pageId="27" pageNumber="40" phylum="Arthropoda" rank="species" species="hagen">
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<pageBreakToken pageId="27" pageNumber="40" start="start">Opisotretus</pageBreakToken>
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hagen
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</taxonomicName>
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<taxonomicNameLabel pageId="27" pageNumber="40">sp. n.</taxonomicNameLabel>
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Figs 22, 23
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="40" type="type material">
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<paragraph pageId="27" pageNumber="40">Type material.</paragraph>
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<paragraph pageId="27" pageNumber="40">Holotype ♂ (NMNHS), Papua New Guinea, Western Highlands Prov., Mount Hagen, ca 1990 m a.s.l., in town, 22.10.1975, leg. P. Beron (British Speleological Expedition).</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="40" type="paratypes">
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<paragraph pageId="27" pageNumber="40">Paratypes.</paragraph>
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<paragraph pageId="27" pageNumber="40">1 ♀ subadult (19 segments), 1 juv. (fragments) (NMNHS), 1 ♀ subadult (19 segments) (SEM; MNHN JC 340), same locality, together with holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="40" type="diagnosis">
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<paragraph pageId="27" pageNumber="40">Diagnosis.</paragraph>
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<paragraph pageId="27" pageNumber="40">Differs readily fromcongeners by a modified ♂ head (two prominent paramedian tubercles above the antennal sockets), coupled with a less strongly curved, nearly suberect gonopod telopodite, with its apical piece crowned with several peculiar, mostly digitiform outgrowths.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="40" type="name">
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<paragraph pageId="27" pageNumber="40">Name.</paragraph>
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<paragraph pageId="27" pageNumber="40">Referring to the type locality; a noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="41" pageId="27" pageNumber="40" type="description">
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<paragraph pageId="27" pageNumber="40">Description.</paragraph>
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<paragraph pageId="27" pageNumber="40">Length of holotype (and of subadult ♀ paratypes) ca 9 mm, width of midbody pro- and metazona 0.8 and 1.15 mm, respectively. Coloration in alcohol from uniformly pallid to light yellowish.</paragraph>
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<paragraph pageId="27" pageNumber="40">
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Body with 19 (♂) or 20 (♀) segments. All characters like in
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<taxonomicName class="Diplopoda" family="Opisotretidae" genus="Retrodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Retrodesmus cavernicola" order="Polydesmida" pageId="27" pageNumber="40" phylum="Arthropoda" rank="species" species="cavernicola">Retrodesmus cavernicola</taxonomicName>
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sp. n., except as follows.
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</paragraph>
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<paragraph pageId="27" pageNumber="40">♂ head with two round, paramedian, rather high tubercles (t) above antennal sockets (Fig. 23A). Antennae medium-sized, strongly clavate, extending behind segment 2 when stretched dorsally (Figs 22D, 23A).</paragraph>
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<paragraph pageId="27" pageNumber="40">
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In width, collum << segment 2 <head = 3 <4 <5 (6) =15 (♂), thereafter body gradually tapering towards telson. Paraterga rather strongly developed, starting from collum, mostly subhorizontal to slightly declivous, set high, but always lying slightly below a moderately (♀, juv.) to weakly (♂) convex dorsum, with rather faint shoulders frontolaterally (Figs 22
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Caudal corner of postcollum paraterga dentiform, always pointed and extending increasingly well behind rear tergal margin. Lateral edge of paraterga with 2 or 3 small setigerous indentations in poreless and poriferous segments, respectively. Ozopores evident, round, flush open on dorsal surface, located very close to caudal margin at bottom of caudalmost lateral incision (Figs 22C, E, M), lateral tooth being very considerably shorter than medial one. Collum and each following metatergum with 3+3 long bacilliform setae arranged in three regular transverse rows; polygonal bosses flat, but visible (Figs 22
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="27" pageNumber="40">Sterna without modifications, rather broad, strongly setose (Fig. 22H). Legs long, incrassate in ♂ due to prefemora alone (Figs 22N, 23B), ca 1.5 times (♂) as long as midbody height; femora and tarsi longest, subequal in length; sphaerotrichomes or other modified setae missing.</paragraph>
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<paragraph lastPageId="28" lastPageNumber="41" pageId="27" pageNumber="40">
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Gonopod telopodite (Figs 23C, D) only very slightly curved, nearly suberect, unipartite, rather long and slender; apical piece (a) distal to a very short solenomere (sl) rather short, on caudal face with a few finger-shaped ornamentations and a strong,
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<pageBreakToken pageId="28" pageNumber="41" start="start">parabasal</pageBreakToken>
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, subspiniform process (pr). An accessory seminal chamber at base of sl evident, crowned with a hairy pulvillus.
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</paragraph>
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<caption pageId="28" pageNumber="41">
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<paragraph pageId="28" pageNumber="41">
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Figure 22.
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<taxonomicName class="Diplopoda" family="Opisotretidae" genus="Opisotretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisotretus hagen" order="Polydesmida" pageId="28" pageNumber="41" phylum="Arthropoda" rank="species" species="hagen">Opisotretus hagen</taxonomicName>
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sp. n., ♀ subadult, paratype; A, D, G, anterior body part, dorsal, lateral and ventral views, respectively B, E, H midbody segments, dorsal, lateral and ventral views, respectively C, F, I posterior body part, dorsal, lateral and caudal views, respectively J tergal seta, lateral view K tegument texture, dorsal view L limbus, dorsal view M midbody paratergite, subcaudal view N midbody leg; O, claw. - Scale bars: A, G 0.2 mm;
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<normalizedToken originalValue="B–F">B-F</normalizedToken>
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, H, I, K, N 0.1 mm; M 0.05 mm; J, L, O 0.01 mm.
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</paragraph>
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</caption>
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<caption pageId="28" pageNumber="41">
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<paragraph pageId="28" pageNumber="41">
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Figure 23.
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<taxonomicName class="Diplopoda" family="Opisotretidae" genus="Opisotretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisotretus hagen" order="Polydesmida" pageId="28" pageNumber="41" phylum="Arthropoda" rank="species" species="hagen">Opisotretus hagen</taxonomicName>
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sp. n., ♂ holotype; A head, frontal view B midbody leg C, D right gonopod, mesal and lateral views, respectively. - Scale bars: A 1.0; B 0.5 mm; C, D 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="41" type="remarks">
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<paragraph pageId="28" pageNumber="41">Remarks.</paragraph>
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<paragraph pageId="28" pageNumber="41">
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This is the first
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<taxonomicName class="Diplopoda" family="Opisotretidae" genus="Opisotretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisotretus" order="Polydesmida" pageId="28" pageNumber="41" phylum="Arthropoda" rank="genus">Opisotretus</taxonomicName>
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showing ♂ head modifications, thus confirming the character as being only species-specific (
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<bibRefCitation author="Golovatch, SI" journalOrPublisher="Entomologica Basiliensia" pageId="56" pageNumber="69" pagination="15 - 48" title="On the first Polydesmidae, Opisotretidae and Fuhrmannodesmidae from Bhutan (Diplopoda, Polydesmida)." volume="12" year="1988">Golovatch 1988</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |