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<mods:title id="51A5ACF3608BAEFEAA3BB8405BE9C210">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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25)
<taxonomicName id="4C61A683EB3D5741FF3FDE9CFCF1FC04" ID-CoL="4GS5H" authority="Chen &amp; Gao, 2005" authorityName="Chen &amp; Gao in Chen et al." authorityYear="2005" box="[198,794,916,943]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="longipenis" subGenus="Ashima">
<emphasis id="B9150112EB3D5741FF3FDE9CFEC5FC05" bold="true" box="[198,302,916,943]" italics="true" pageId="27" pageNumber="28">Phortica</emphasis>
(
<emphasis id="B9150112EB3D5741FEC6DE9CFE77FC05" bold="true" box="[319,412,916,943]" italics="true" pageId="27" pageNumber="28">Ashima</emphasis>
)
<emphasis id="B9150112EB3D5741FE54DE9CFDC0FC05" bold="true" box="[429,555,916,943]" italics="true" pageId="27" pageNumber="28">longipenis</emphasis>
<bibRefCitation id="EFF0A0F1EB3D5741FDCBDE9CFCF1FC04" author="Chen &amp; Gao" box="[562,794,916,942]" firstAuthor="Chen" pageId="27" pageNumber="28" pagination="3951 - 3978" refId="ref33869" refString="Chen, H. W., Gao, J. J. &amp; Wen, S. Y. (2005 a) Species diversity of the genus Phortica Schiner in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae [sic]). Journal of Natural History, 39, 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article" year="2005">Chen &amp; Gao, 2005</bibRefCitation>
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<paragraph id="8BDEDD00EB3D5741FF6EDEB0FF14FC79" blockId="27.[151,794,916,979]" box="[151,255,952,979]" pageId="27" pageNumber="28">
(
<figureCitation id="135AC185EB3D5741FF59DEB0FF13FC79" box="[160,248,952,979]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (AG; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, DH; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="27" pageNumber="28">Fig. 6H</figureCitation>
)
</paragraph>
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<paragraph id="8BDEDD00EB3D5741FF6EDEFEFCECFBA4" blockId="27.[151,775,1013,1069]" box="[151,775,1013,1038]" pageId="27" pageNumber="28">
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<treatmentCitation id="0AC0FB11EB3D5741FF6EDEFEFCE8FBA4" author="Chen, H. W. &amp; Gao, J. J. &amp; Wen, S. Y." box="[151,771,1013,1038]" httpUri="http://treatment.plazi.org/id/C638879BFF85FFB20258EB23FC5ACC3E" page="3957" pageId="27" pageNumber="28" year="2005">
<taxonomicName id="4C61A683EB3D5741FF6EDEFEFCE8FBA4" ID-CoL="4GS5H" authority="Chen &amp; Gao in Chen et al., 2005 a: 3957" authorityName="Chen &amp; Gao in Chen et al." authorityPageNumber="3957" authorityYear="2005" box="[151,771,1013,1038]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="B9150112EB3D5741FF6EDEFEFE8BFBA4" box="[151,352,1014,1038]" italics="true" pageId="27" pageNumber="28">Phortica longipenis</emphasis>
Chen &amp; Gao in
<bibRefCitation id="EFF0A0F1EB3D5741FDFFDEFDFCE8FBA4" author="Chen" box="[518,771,1013,1038]" etAl="et al." firstAuthor="Chen" pageId="27" pageNumber="28" pagination="3951 - 3978" refId="ref33869" refString="Chen, H. W., Gao, J. J. &amp; Wen, S. Y. (2005 a) Species diversity of the genus Phortica Schiner in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae [sic]). Journal of Natural History, 39, 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article" year="2005" yearSuffix="a">
Chen
<emphasis id="B9150112EB3D5741FDBBDEFEFD84FBA4" box="[578,623,1014,1038]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
., 2005a: 3957
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</paragraph>
<paragraph id="8BDEDD00EB3D5741FF6ED91CFD33FB87" blockId="27.[151,775,1013,1069]" box="[151,728,1044,1069]" pageId="27" pageNumber="28">
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<treatmentCitation id="0AC0FB11EB3D5741FF6ED91CFD3FFB87" author="Chen, H. W. &amp; Maca, J." box="[151,724,1044,1069]" page="507" pageId="27" pageNumber="28" year="2012">
<taxonomicName id="4C61A683EB3D5741FF6ED91CFD3FFB87" ID-CoL="4GS5H" authority=": Chen &amp; Maca, 2012: 507" authorityName=": Chen &amp; Maca" authorityPageNumber="507" authorityYear="2012" box="[151,724,1044,1069]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="longipenis" subGenus="Ashima">
<emphasis id="B9150112EB3D5741FF6ED91CFF1BFB86" box="[151,240,1044,1068]" italics="true" pageId="27" pageNumber="28">Phortica</emphasis>
(
<emphasis id="B9150112EB3D5741FF06D91CFEA0FB86" box="[255,331,1044,1068]" italics="true" pageId="27" pageNumber="28">Ashima</emphasis>
)
<emphasis id="B9150112EB3D5741FEA0D91CFE28FB86" box="[345,451,1044,1068]" italics="true" pageId="27" pageNumber="28">longipenis</emphasis>
:
<bibRefCitation id="EFF0A0F1EB3D5741FE29D91CFD3FFB87" author="Chen &amp; Maca" box="[464,724,1044,1069]" firstAuthor="Chen" pageId="27" pageNumber="28" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. &amp; Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen &amp; Máca, 2012: 507</bibRefCitation>
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<subSubSection id="C37B8E8BEB3D5741FF6ED95BFD62F9E8" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BDEDD00EB3D5741FF6ED95BFCB5FB55" blockId="27.[151,1437,1107,1638]" pageId="27" pageNumber="28">
<emphasis id="B9150112EB3D5741FF6ED95BFEF9FBC4" bold="true" box="[151,274,1107,1134]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Apical, leaf-like expansion of arista much narrower than long, apically with 1 long, medial and 2 minute, lateral, awn-like projections (“
<figureCitation id="135AC185EB3D5741FDE6D970FD90FB38" box="[543,635,1144,1170]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="27" pageNumber="28">Fig. 4A</figureCitation>
” in
<bibRefCitation id="EFF0A0F1EB3D5741FD52D970FC92FB38" author="Chen" box="[683,889,1143,1171]" etAl="et al." firstAuthor="Chen" pageId="27" pageNumber="28" pagination="3951 - 3978" refId="ref33869" refString="Chen, H. W., Gao, J. J. &amp; Wen, S. Y. (2005 a) Species diversity of the genus Phortica Schiner in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae [sic]). Journal of Natural History, 39, 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article" year="2005" yearSuffix="a">
Chen
<emphasis id="B9150112EB3D5741FD17D971FCCAFB38" box="[750,801,1144,1170]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2005a
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); strongly sclerotized arch on anterior margin of aedeagal sheath recurved at anterior ends as postgonites, posteriorly with one large and one small postgonites (projections); one anterior postgonite curved inward, longer, strongly sclerotized, apically with small claw, but the other curved dorsad, shorter, less sclerotized, without claw (
<figureCitation id="135AC185EB3D5741FD0ED9ECFCBAFB55" box="[759,849,1252,1279]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (AG; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, DH; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="27" pageNumber="28">Fig. 6H</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB3D5741FF3ED80FFE33FAC0" blockId="27.[151,1437,1107,1638]" pageId="27" pageNumber="28">
<emphasis id="B9150112EB3D5741FF3ED80FFDE5FA88" bold="true" box="[199,526,1287,1314]" pageId="27" pageNumber="28">Supplementary description</emphasis>
(not repeating characters common to
<taxonomicName id="4C61A683EB3D5741FC3AD801FB90FA88" authorityName="Gong, Zhu &amp; Chen" authorityYear="2018" box="[963,1147,1288,1315]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="semiannulata">
<emphasis id="B9150112EB3D5741FC3AD801FC3FFA88" box="[963,980,1289,1314]" italics="true" pageId="27" pageNumber="28">P</emphasis>
.
<emphasis id="B9150112EB3D5741FC1DD800FB90FA88" box="[996,1147,1288,1314]" italics="true" pageId="27" pageNumber="28">semiannulata</emphasis>
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). Supracervical setae approximately 16. Arista with arb = 13/0. Cibarial, posterior sensilla 67 per side. Pregonite without sensilla on submedial portion (ch.52-0).
</paragraph>
<paragraph id="8BDEDD00EB3D5741FF3ED87CFBBCFA7D" blockId="27.[151,1437,1107,1638]" pageId="27" pageNumber="28">
The antisymmetry is observed in the postgonites: in A-type, the large, posterior postgonite and strongly sclerotized, inward curved, anterior postgonite is present on the right side (
<figureCitation id="135AC185EB3D5741FC50D890FBEFFA19" box="[937,1028,1432,1459]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (AG; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, DH; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="27" pageNumber="28">Fig. 6H</figureCitation>
), while the small, posterior one and less sclerotized, dorsad curved, anterior one on the left side; but in B-type,
<emphasis id="B9150112EB3D5741FC26D8B5FBBBFA7C" box="[991,1104,1469,1494]" italics="true" pageId="27" pageNumber="28">vice versa</emphasis>
.
</paragraph>
<paragraph id="8BDEDD00EB3D5741FF3ED8D7FD62F9E8" blockId="27.[151,1437,1107,1638]" pageId="27" pageNumber="28">
<emphasis id="B9150112EB3D5741FF3ED8D7FE2DFA50" bold="true" box="[199,454,1503,1530]" pageId="27" pageNumber="28">Specimens examined.</emphasis>
<materialsCitation id="3B09D75DEB3D5741FE34D8E8FD42FA50" ID-GBIF-Occurrence="2636268360" box="[461,681,1504,1530]" country="China" location="China" pageId="27" pageNumber="28" specimenCount="4" specimenCount-male="4">
<collectingCountry id="F3769D90EB3D5741FE34D8E8FDFFFA50" box="[461,532,1504,1530]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
:
<specimenCount id="9D671689EB3D5741FDE6D8E8FDA9FA50" box="[543,578,1504,1530]" pageId="27" pageNumber="28" type="male">4♂</specimenCount>
(A-type)
</materialsCitation>
<materialsCitation id="3B09D75DEB3D5741FD49D8E8FDA0F9B4" ID-GBIF-Occurrence="2636268337" collectingDate="2000-08-17" collectionCode="SEHU" collectorName="H. - W. Chen" country="China" elevation="1840" location="Binchuan" municipality="Mt. Jizu" pageId="27" pageNumber="28" specimenCount="6" specimenCount-male="6" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D671689EB3D5741FD49D8E8FD38FA50" box="[688,723,1504,1530]" pageId="27" pageNumber="28" type="male">6♂</specimenCount>
(B-type)
<typeStatus id="54DA63A2EB3D5741FCB9D8E9FC45FA51" box="[832,942,1505,1531]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
,
<collectingMunicipality id="6BBA477AEB3D5741FC40D8E9FBF3FA50" box="[953,1048,1504,1531]" pageId="27" pageNumber="28">Mt. Jizu</collectingMunicipality>
,
<location id="8EBE8BDBEB3D5741FBDAD8E8FB7AFA50" LSID="urn:lsid:plazi:treatment:03C86C16EB3D5741FF6EDE9CFD4BF9CC:8EBE8BDBEB3D5741FBDAD8E8FB7AFA50" box="[1059,1169,1504,1530]" country="China" municipality="Mt. Jizu" name="Binchuan" pageId="27" pageNumber="28" stateProvince="Yunnan">Binchuan</location>
,
<collectingRegion id="49A513E2EB3D5741FB62D8E9FB1FFA51" box="[1179,1268,1505,1531]" country="China" name="Yunnan" pageId="27" pageNumber="28">Yunnan</collectingRegion>
,
<elevation id="004C3A33EB3D5741FB07D8E8FA7CFA50" box="[1278,1431,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="27" pageNumber="28" unit="m" value="1840.0">
<quantity id="4C9970E5EB3D5741FB07D8E8FAB1FA51" box="[1278,1370,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="27" pageNumber="28" unit="m" value="1840.0">1,840 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFBC0EB3D5741FF6EDB0CFECAF9B4" box="[151,289,1540,1566]" pageId="27" pageNumber="28" value="2000-08-17">
<collectingDate id="EF9B0228EB3D5741FF6EDB0CFECAF9B4" box="[151,289,1540,1566]" pageId="27" pageNumber="28" value="2000-08-17">17.viii.2000</collectingDate>
</date>
,
<collectorName id="2694B8D6EB3D5741FED4DB0DFE58F9B4" box="[301,435,1540,1567]" pageId="27" pageNumber="28">H.-W. Chen</collectorName>
leg. (
<collectionCode id="ED7045C5EB3D5741FE0DDB0CFDA8F9B4" box="[500,579,1540,1566]" pageId="27" pageNumber="28">SEHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B09D75DEB3D5741FDAFDB0CFD6EF9E8" ID-GBIF-Occurrence="2636268347" collectingDate="2008-07-25" collectionCode="SEHU" collectorName="H. Banziger" country="Thailand" elevation="1250" location="Doi Suthep" municipality="Sangwal" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingCountry id="F3769D90EB3D5741FDAFDB0CFD55F9B4" box="[598,702,1540,1566]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
:
<specimenCount id="9D671689EB3D5741FD30DB0CFD07F9B4" box="[713,748,1540,1566]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
(A-type), above
<collectingMunicipality id="6BBA477AEB3D5741FC57DB0CFBFAF9B4" box="[942,1041,1540,1566]" pageId="27" pageNumber="28">Sangwal</collectingMunicipality>
,
<location id="8EBE8BDBEB3D5741FBE4DB0CFB4BF9B4" LSID="urn:lsid:plazi:treatment:03C86C16EB3D5741FF6EDE9CFD4BF9CC:8EBE8BDBEB3D5741FBE4DB0CFB4BF9B4" box="[1053,1184,1540,1566]" country="Thailand" municipality="Sangwal" name="Doi Suthep" pageId="27" pageNumber="28" stateProvince="Chiang Mai">Doi Suthep</location>
,
<collectingRegion id="49A513E2EB3D5741FB52DB0CFADFF9B4" box="[1195,1332,1540,1566]" country="Thailand" name="Chiang Mai" pageId="27" pageNumber="28">Chiang Mai</collectingRegion>
,
<elevation id="004C3A33EB3D5741FAB9DB0BFF26F9E8" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="27" pageNumber="28" unit="m" value="1250.0">
<quantity id="4C9970E5EB3D5741FAB9DB0BFA77F9B5" box="[1344,1436,1539,1567]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="27" pageNumber="28" unit="m" value="1250.0">1,250 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFBC0EB3D5741FF20DB2FFEB7F9E8" box="[217,348,1575,1602]" pageId="27" pageNumber="28" value="2008-07-25">
<collectingDate id="EF9B0228EB3D5741FF20DB2FFEB7F9E8" box="[217,348,1575,1602]" pageId="27" pageNumber="28" value="2008-07-25">25.vii.2008</collectingDate>
</date>
,
<collectorName id="2694B8D6EB3D5741FE9EDB21FE05F9E8" box="[359,494,1576,1603]" pageId="27" pageNumber="28">H. Bänziger</collectorName>
leg. (
<collectionCode id="ED7045C5EB3D5741FDD4DB20FD97F9E8" box="[557,636,1576,1602]" pageId="27" pageNumber="28">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB3D5741FF3EDB43FD4BF9CC" box="[199,672,1611,1638]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BDEDD00EB3D5741FF3EDB43FD4BF9CC" blockId="27.[151,1437,1107,1638]" box="[199,672,1611,1638]" pageId="27" pageNumber="28">
<emphasis id="B9150112EB3D5741FF3EDB43FE8AF9CC" bold="true" box="[199,353,1611,1638]" pageId="27" pageNumber="28">Distribution.</emphasis>
<collectingCountry id="F3769D90EB3D5741FE9EDB44FE41F9CC" box="[359,426,1612,1638]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
(
<collectingRegion id="49A513E2EB3D5741FE40DB44FDFEF9CC" box="[441,533,1612,1638]" country="China" name="Yunnan" pageId="27" pageNumber="28">Yunnan</collectingRegion>
),
<collectingCountry id="F3769D90EB3D5741FDDEDB44FD7AF9CC" box="[551,657,1612,1638]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
*.
</paragraph>
</subSubSection>
</treatment>
</document>