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<document id="17DBD9DAED2518235FE32FBFE538243B" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB07577BFF6EDE61FBB8F949" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) angulata Prigent &amp; Chen 2008" docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="34" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2EBEBD44E78DB36A018DE7F993820DD9">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="921BA6E5AA7B46436164411D1E770400">Toda, Masanori J.</mods:namePart>
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<mods:namePart id="5A9C511C646B27D4BE911837ADFE5AF5">Bänziger, Hans</mods:namePart>
<mods:affiliation id="1F649E8017E7E6C95E01042A909072D2">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="DBDEE32BED44E87E88C829569A184214">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="D68DB7CE2C67182A9AD3B8095530C0D6">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="D4ADF6DB6F222BB9ED4A757DC2437DE3">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="149DA13F75520359124C52B8F63DF5C7">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<treatment id="03C86C16EB07577BFF6EDE61FBB8F949" ID-DOI="http://doi.org/10.5281/zenodo.5221367" ID-GBIF-Taxon="164612809" ID-Zenodo-Dep="5221367" LSID="urn:lsid:plazi:treatment:03C86C16EB07577BFF6EDE61FBB8F949" httpUri="http://treatment.plazi.org/id/03C86C16EB07577BFF6EDE61FBB8F949" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="C37B8E8BEB07577BFF6EDE61FD12FC29" box="[151,761,873,900]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BDEDD00EB07577BFF6EDE61FD12FC29" blockId="33.[151,761,873,900]" box="[151,761,873,900]" pageId="33" pageNumber="34">
<heading id="D0966A6CEB07577BFF6EDE61FD12FC29" bold="true" box="[151,761,873,900]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B9150112EB07577BFF6EDE61FD12FC29" bold="true" box="[151,761,873,900]" pageId="33" pageNumber="34">
4.
<taxonomicName id="4C61A683EB07577BFF4CDE61FDE3FC2E" ID-CoL="8DBV8" authorityName="Prigent &amp; Chen" authorityYear="2008" box="[181,520,873,900]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="angulata" subGenus="Ashima">
<emphasis id="B9150112EB07577BFF4CDE61FEF6FC2E" bold="true" box="[181,285,873,900]" italics="true" pageId="33" pageNumber="34">Phortica</emphasis>
(
<emphasis id="B9150112EB07577BFED7DE61FE60FC2E" bold="true" box="[302,395,873,900]" italics="true" pageId="33" pageNumber="34">Ashima</emphasis>
)
<emphasis id="B9150112EB07577BFE65DE61FDE3FC2E" bold="true" box="[412,520,873,900]" italics="true" pageId="33" pageNumber="34">angulata</emphasis>
</taxonomicName>
species group, new
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37B8E8BEB07577BFF6EDEB8FBB8F949" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BDEDD00EB07577BFF6EDEB8FC50FB92" blockId="33.[151,1437,944,1763]" pageId="33" pageNumber="34">
<emphasis id="B9150112EB07577BFF6EDEB8FEF9FC61" bold="true" box="[151,274,944,971]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
Dorsomedial, tentorial apodeme ≥ 2/3 basal, parallel portion of dorsolateral, tentorial apodeme (ch.1-0); abdominal tergites III and IV unicolor (ch.21-3); epandrium with triangular or hook-like projection on anteroventral margin (ch.31-2;
<figureCitation id="135AC185EB07577BFEA7DEF1FE5CFBB9" box="[350,439,1017,1043]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas &amp; Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent &amp; Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (DG; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen &amp; Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="33" pageNumber="34">Fig. 3B</figureCitation>
); pregonite (
<figureCitation id="135AC185EB07577BFDB0DEF1FD4AFBB9" box="[585,673,1017,1043]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas &amp; Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent &amp; Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (DG; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen &amp; Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="33" pageNumber="34">Fig. 3C</figureCitation>
) not strongly sclerotized (ch.43-2), plate-like (ch.44-0), with basal extension nearly rectangular to main body and apically broad (ch.49-2).
</paragraph>
<paragraph id="8BDEDD00EB07577BFF3ED948FCA5F9D2" blockId="33.[151,1437,944,1763]" pageId="33" pageNumber="34">
<emphasis id="B9150112EB07577BFF3ED948FE43FBF1" bold="true" box="[199,424,1088,1115]" pageId="33" pageNumber="34">Shared characters.</emphasis>
Arista with dorsal but no ventral branches; apical, leaf-like expansion apically with small, awn-like projection. Palpus with hollow-shaped sense organ (ch.9-2). Some anterior, cibarial, medial sensilla short, apically blunt, peg-like (ch.11-1). Prementum without pubescent patches (ch.12-0). Pseudotracheae in labellum 6 per side (ch.14-2). Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Mid and hind femora and all tibiae entirely yellow (ch.1619-0). Apical, downward curved setula on fore tarsomere V stout, as long as tarsomere V (ch.20-1). Abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0); IV not wider than long (ch.24-0); V wider than long (ch.25-1); IIIV without dense setulae on medial surface (ch.2628-0). Setae on and/or near ventral margin of epandrium (
<figureCitation id="135AC185EB07577BFBC5D835FB7EFAFD" box="[1084,1173,1341,1367]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas &amp; Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent &amp; Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (DG; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen &amp; Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="33" pageNumber="34">Fig. 3B</figureCitation>
) not denser than those on other portions (ch.32-0), longer than others (ch.33-1). Surstylus (
<figureCitation id="135AC185EB07577BFC50D869FBEAFAD1" box="[937,1025,1377,1403]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas &amp; Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent &amp; Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (DG; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen &amp; Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="33" pageNumber="34">Fig. 3B</figureCitation>
) without expansion at caudoventral corner (ch.35-0), with peg-like teeth (ch.37-0). Hypandrium symmetric (ch.40-0); phragma as narrow as lateral gonocoxites (ch.41-1). Pregonite (
<figureCitation id="135AC185EB07577BFDEBD8A1FD8CFA69" box="[530,615,1449,1475]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas &amp; Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent &amp; Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (DG; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen &amp; Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="33" pageNumber="34">Fig. 3C</figureCitation>
) not serrated on ventrodistal portion (ch.50-0), with sensilla on submedial and basal portions (ch.52,53-1). Aedeagal, outer membrane bearing neither spiny ornamentation nor apical, long, narrow tube (ch.55-1, ch.57-0); median rod entirely or partly strongly sclerotized (ch.58-0); free sclerite absent within outer membrane (ch.59-0). Posterior, rod-like process of phallapodeme not longer than aedeagal median rod (ch.62-0). Aedeagal sheath with lateral lobes bearing strongly sclerotized postgonites (ch.65-1); sclerotized, posterior postgonites medially separated from each other (ch.66-0).
</paragraph>
<paragraph id="8BDEDD00EB07577BFF3EDB89FBB8F949" blockId="33.[151,1437,944,1763]" pageId="33" pageNumber="34">
<emphasis id="B9150112EB07577BFF3EDB89FE7AF931" bold="true" box="[199,401,1664,1691]" pageId="33" pageNumber="34">Included species.</emphasis>
This species group comprises African species:
<taxonomicName id="4C61A683EB07577BFC5DDB89FA73F931" authority="Prigent &amp; Chen" authorityName="Prigent &amp; Chen" authorityYear="2008" box="[932,1432,1665,1691]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="angulata" subGenus="Ashima">
<emphasis id="B9150112EB07577BFC5DDB89FBEDF931" box="[932,1030,1665,1691]" italics="true" pageId="33" pageNumber="34">Phortica</emphasis>
(
<emphasis id="B9150112EB07577BFBEFDB89FB81F931" box="[1046,1130,1665,1691]" italics="true" pageId="33" pageNumber="34">Ashima</emphasis>
)
<emphasis id="B9150112EB07577BFB83DB89FB35F931" box="[1146,1246,1665,1691]" italics="true" pageId="33" pageNumber="34">angulata</emphasis>
Prigent &amp; Chen
</taxonomicName>
;
<emphasis id="B9150112EB07577BFF6EDBAEFF43F915" box="[151,168,1702,1727]" italics="true" pageId="33" pageNumber="34">P</emphasis>
.
<taxonomicName id="4C61A683EB07577BFF4EDBADFDCBF915" authority="Prigent &amp; Chen" authorityName="Prigent &amp; Chen" authorityYear="2008" box="[183,544,1701,1727]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="curvispina" subGenus="Ashima">
(
<emphasis id="B9150112EB07577BFF38DBADFF39F915" box="[193,210,1701,1727]" italics="true" pageId="33" pageNumber="34">A</emphasis>
.)
<emphasis id="B9150112EB07577BFF13DBAEFE8BF915" box="[234,352,1702,1727]" italics="true" pageId="33" pageNumber="34">curvispina</emphasis>
Prigent &amp; Chen
</taxonomicName>
*;
<emphasis id="B9150112EB07577BFDC1DBAEFDA2F915" box="[568,585,1702,1727]" italics="true" pageId="33" pageNumber="34">P</emphasis>
.
<taxonomicName id="4C61A683EB07577BFDA1DBADFC2CF915" authority="Prigent &amp; Chen" authorityName="Prigent &amp; Chen" authorityYear="2008" box="[600,967,1701,1727]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="machoruka" subGenus="Ashima">
(
<emphasis id="B9150112EB07577BFD9BDBADFD98F915" box="[610,627,1701,1727]" italics="true" pageId="33" pageNumber="34">A</emphasis>
.)
<emphasis id="B9150112EB07577BFD72DBADFCE2F915" box="[651,777,1701,1727]" italics="true" pageId="33" pageNumber="34">machoruka</emphasis>
Prigent &amp; Chen
</taxonomicName>
; and
<emphasis id="B9150112EB07577BFBFDDBAEFBFEF915" box="[1028,1045,1702,1727]" italics="true" pageId="33" pageNumber="34">P</emphasis>
.
<taxonomicName id="4C61A683EB07577BFBDDDBADFA9CF915" authority="Prigent &amp; Chen" authorityName="Prigent &amp; Chen" authorityYear="2008" box="[1060,1399,1701,1727]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="unispina" subGenus="Ashima">
(
<emphasis id="B9150112EB07577BFBD4DBADFBD5F915" box="[1069,1086,1701,1727]" italics="true" pageId="33" pageNumber="34">A</emphasis>
.)
<emphasis id="B9150112EB07577BFBAEDBAEFB5CF915" box="[1111,1207,1702,1727]" italics="true" pageId="33" pageNumber="34">unispina</emphasis>
Prigent &amp; Chen
</taxonomicName>
*. * The assignment was judged from the original descriptions by
<bibRefCitation id="EFF0A0F1EB07577BFCB2DBC1FBA4F949" author="Prigent &amp; Chen" box="[843,1103,1737,1763]" firstAuthor="Prigent" pageId="33" pageNumber="34" pagination="18 - 30" refId="ref36494" refString="Prigent, S. &amp; Chen, H. W. (2008) A survey of the genus Phortica Schiner from Kenya, Africa (Diptera, Drosophilidae). Zootaxa, 1773 (1), 18 - 30. https: // doi. org / 10.11646 / zootaxa. 1773.1.2" type="journal article" year="2008">Prigent &amp; Chen (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>