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<document id="F73B2B1FDE8807968E23356CB44F2758" ID-CLB-Dataset="32583" ID-DOI="10.11646/zootaxa.4300.1.2" ID-GBIF-Dataset="1eaf4800-94f1-4618-9d58-a6b71724d11b" ID-ISSN="1175-5326" ID-Zenodo-Dep="837121" ID-ZooBank="1B15875B-A2CF-4E45-9E21-2B803D659176" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501581815757" checkinUser="plazi" docAuthor="Kottelat, Maurice" docDate="2017" docId="D7609840FFA6FF9CFF36F91DFAB599C3" docLanguage="en" docName="zootaxa.4300.1.2.pdf" docOrigin="Zootaxa 4300 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schistura epixenos Kottelat 2017, new species" docType="treatment" docVersion="7" masterDocId="2B59E038FFA7FF9BFFA1FFD8FFE79836" masterDocTitle="Schistura epixenos, a new species of loach from the Nakai Plateau, Laos (Teleostei: Nemacheilidae)" updateTime="1698466097109" updateUser="ExternalLinkService">
<mods:mods id="6891C19304EB5C145295DAA582FC1F61" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14AA8C6B7D3ADFD1AD7BE311052E94E9">
<mods:title id="9A7B0B5EED199736B3D0183DB42B3251">Schistura epixenos, a new species of loach from the Nakai Plateau, Laos (Teleostei: Nemacheilidae)</mods:title>
</mods:titleInfo>
<mods:name id="D3FA6879D801B26530F8F315E16474C0" type="personal">
<mods:role id="87CB809C2E8294AC9C6F5F6BE908D04D">
<mods:roleTerm id="F3BDB77C9845ED9F72ACC321519D4BC1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB00CC4109F80909195584ABFBF9B0A6">Kottelat, Maurice</mods:namePart>
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<mods:typeOfResource id="6F44BB045EA64403A70A57CFD15889A9">text</mods:typeOfResource>
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<mods:title id="B8579FA9719FC0B623454E952DDA9A9E">Zootaxa</mods:title>
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<mods:part id="19266F0465FCACEE9F76A7B7C0917125">
<mods:date id="5C2420BAF0042F283D8082E2FB418983">2017</mods:date>
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<mods:number id="F8832FEF4537B50AB3C9E3531563CA51">4300</mods:number>
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<mods:classification id="E8A39A3010DFC85C2E8331EF11DD669C">journal volume</mods:classification>
<mods:identifier id="F308C811BC27ACCCB2205236F05CB576" type="CLB-Dataset">32583</mods:identifier>
<mods:identifier id="ECD8B886F4EB760FE3220D86A5D34438" type="DOI">10.11646/zootaxa.4300.1.2</mods:identifier>
<mods:identifier id="3F1D8D07418FC87BB9375FD7BC2255FB" type="GBIF-Dataset">1eaf4800-94f1-4618-9d58-a6b71724d11b</mods:identifier>
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<treatment id="D7609840FFA6FF9CFF36F91DFAB599C3" ID-DOI="http://doi.org/10.5281/zenodo.6008587" ID-GBIF-Taxon="132618047" ID-Zenodo-Dep="6008587" LSID="urn:lsid:plazi:treatment:D7609840FFA6FF9CFF36F91DFAB599C3" httpUri="http://treatment.plazi.org/id/D7609840FFA6FF9CFF36F91DFAB599C3" lastPageId="7" pageId="1">
<subSubSection id="17D37ADDFFA6FF9AFF36F91DFEF59F37" pageId="1" type="nomenclature">
<paragraph id="5F762956FFA6FF9AFF36F91DFDFD9ED6" blockId="1.[151,538,1733,1793]" box="[151,538,1733,1760]" pageId="1">
<heading id="043E9E3AFFA6FF9AFF36F91DFDFD9ED6" bold="true" box="[151,538,1733,1760]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="6DBDF544FFA6FF9AFF36F91DFDFD9ED6" bold="true" box="[151,538,1733,1760]" pageId="1">
<taxonomicName id="98C952D5FFA6FF9AFF36F91DFE9B9EE9" authority="Kottelat, 2017" authorityName="Kottelat" authorityYear="2017" box="[151,380,1733,1759]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" phylum="Chordata" rank="species" species="epixenos" status="sp. nov.">
<emphasis id="6DBDF544FFA6FF9AFF36F91DFE9B9EE9" bold="true" box="[151,380,1733,1759]" italics="true" pageId="1">Schistura epixenos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="768E483FFFA6FF9AFE2BF91EFDFD9ED6" box="[394,538,1734,1760]" pageId="1" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5F762956FFA6FF9AFF36F930FEF59F37" blockId="1.[151,538,1733,1793]" box="[151,274,1768,1793]" pageId="1">
(
<figureCitation id="C7F235D3FFA6FF9AFF3EF930FEED9F37" box="[159,266,1768,1793]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1151,1173]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="5.[151,250,968,990]" captionTargetBox-0="[193,1374,220,1100]" captionTargetBox-1="[198,1374,214,1940]" captionTargetBox-2="[161,1400,346,930]" captionTargetId-0="figure@2.[159,1407,193,1129]" captionTargetId-1="figure@3.[189,1398,193,1951]" captionTargetId-2="figure@5.[150,1436,301,947]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 2. Schistura epixenos, MHNG 2767.086, holotype, 59.5 mm SL; Laos: Nakai Plateau: Nam Kathang Gnai." captionText-1="FIGURE 3. Schistura epixenos, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai; a, 56.3 mm SL; b, 57.2 mm SL (right side, reversed); c, 57.0 mm SL (right side, reversed)." captionText-2="FIGURE 4. Schistura epixenos, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai; a, 22.4 mm SL; b, 29.2 mm SL (right side, reversed)." httpUri-0="https://zenodo.org/record/837125/files/figure.png" httpUri-1="https://zenodo.org/record/837127/files/figure.png" httpUri-2="https://zenodo.org/record/837129/files/figure.png" pageId="1">Figs. 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="17D37ADDFFA6FF9AFF36F8E8FEB99FEF" pageId="1" type="materials_examined">
<paragraph id="5F762956FFA6FF9AFF36F8E8FADA9F5B" blockId="1.[151,1436,1840,2009]" pageId="1">
<materialsCitation id="EFA1230BFFA6FF9AFF36F8E8FADA9F5B" ID-GBIF-Occurrence="1585189602" collectingDate="2012-05-07" collectionCode="MHNG" collectorName="M. Kottelat" country="Laos" county="Xe Bangfai" latitude="17.618334" location="Nakai Plateau" longLatPrecision="21" longitude="105.2975" municipality="Nam Kathang Gnai" pageId="1" specimenCode="MHNG 2767.086, 59.5" specimenCount="1" stateProvince="Khammouan ProvinCe" typeStatus="holotype">
<emphasis id="6DBDF544FFA6FF9AFF36F8E8FEEC9F7F" bold="true" box="[151,267,1840,1865]" pageId="1">
<typeStatus id="807297F4FFA6FF9AFF36F8E8FEE19F7F" box="[151,262,1840,1865]" pageId="1" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="0F6F812DFFA6FF9AFEB3F8E9FDF19F7F" box="[274,534,1840,1865]" pageId="1">
<collectionCode id="39D8B193FFA6FF9AFEB3F8E9FE809F7F" box="[274,359,1841,1865]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1">MHNG</collectionCode>
2767.086,
<quantity id="983184B3FFA6FF9AFE44F8E8FDF19F7F" box="[485,534,1840,1865]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="1" unit="mm" value="59.5">59.5</quantity>
</specimenCode>
mm SL;
<collectingCountry id="27DE69C6FFA6FF9AFDDFF8E9FD5E9F7E" box="[638,697,1841,1864]" name="Laos" pageId="1">Laos</collectingCountry>
:
<collectingRegion id="9D0DE7B4FFA6FF9AFD62F8E8FC249F7F" box="[707,963,1840,1865]" country="Laos" name="Khammouan" pageId="1">Khammouan ProvinCe</collectingRegion>
:
<collectingCounty id="B61751DAFFA6FF9AFC6CF8E9FBAA9F7F" box="[973,1101,1840,1865]" pageId="1">Xe Bangfai</collectingCounty>
drainage:
<collectingMunicipality id="BF12B32CFFA6FF9AFB62F8E9FA7B9F7F" box="[1219,1436,1840,1865]" pageId="1">Nam Kathang Gnai</collectingMunicipality>
on edge of
<location id="5A167F8DFFA6FF9AFEB5F88CFE539F5B" LSID="urn:lsid:plazi:treatment:D7609840FFA6FF9CFF36F91DFAB599C3:5A167F8DFFA6FF9AFEB5F88CFE539F5B" box="[276,436,1876,1901]" country="Laos" county="Xe Bangfai" latitude="17.618334" longLatPrecision="21" longitude="105.2975" municipality="Nam Kathang Gnai" name="Nakai Plateau" pageId="1" stateProvince="Khammouan ProvinCe">Nakai Plateau</location>
, near
<location id="5A167F8DFFA6FF9AFE57F88EFD489F58" LSID="urn:lsid:plazi:treatment:D7609840FFA6FF9CFF36F91DFAB599C3:5A167F8DFFA6FF9AFE57F88EFD489F58" box="[502,687,1876,1902]" country="Laos" county="Xe Bangfai" latitude="17.618334" longLatPrecision="21" longitude="105.2975" municipality="Nam Kathang Gnai" name="Ban Khon Keng" pageId="1" stateProvince="Khammouan ProvinCe">Ban Khon Keng</location>
;
<geoCoordinate id="3AFD4F91FFA6FF9AFD1BF88CFCDA9F5B" box="[698,829,1876,1901]" direction="north" orientation="latitude" pageId="1" precision="15" value="17.618334">17°37'06''N</geoCoordinate>
<geoCoordinate id="3AFD4F91FFA6FF9AFCE5F88CFC329F5B" box="[836,981,1876,1901]" direction="east" orientation="longitude" pageId="1" precision="15" value="105.2975">105°17'51''E</geoCoordinate>
;
<collectorName id="F23C4C80FFA6FF9AFC41F88EFB869F5B" box="[992,1121,1876,1901]" pageId="1">M. Kottelat</collectorName>
<emphasis id="6DBDF544FFA6FF9AFBC9F88EFB7E9F5B" box="[1128,1177,1877,1901]" italics="true" pageId="1">et al</emphasis>
.,
<date id="2B770F96FFA6FF9AFB0FF88EFADE9F5B" box="[1198,1337,1877,1902]" pageId="1" value="2012-05-07">
<collectingDate id="3B33F67EFFA6FF9AFB0FF88EFADE9F5B" box="[1198,1337,1877,1902]" pageId="1" value="2012-05-07">7 May 2012</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="5F762956FFA6FF9AFF66F8A0FEB99FEF" blockId="1.[151,1436,1840,2009]" pageId="1">
<materialsCitation id="EFA1230BFFA6FF9AFF66F8A0FEB99FEF" ID-GBIF-Occurrence="1585189601" collectingDate="2012-05-14" collectorName="M. Kottelat" country="Laos" county="Xe Bangfai" latitude="17.651388" location="Nakai Plateau" longLatPrecision="21" longitude="105.237495" municipality="Nam Kathang Gnai" pageId="1" specimenCount="1" stateProvince="Khammouan ProvinCe" typeStatus="paratype">
<emphasis id="6DBDF544FFA6FF9AFF66F8A0FEA19FA7" bold="true" box="[199,326,1912,1937]" pageId="1">
<typeStatus id="807297F4FFA6FF9AFF66F8A0FEA69FA7" box="[199,321,1912,1937]" pageId="1" type="paratype">Paratypes</typeStatus>
.
</emphasis>
CMK 22962, 7,
<quantity id="983184B3FFA6FF9AFDA7F8A0FD4D9FA6" box="[518,682,1912,1937]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" metricValueMax="5.720000000000001" metricValueMin="2.2399999999999998" pageId="1" unit="mm" value="39.8" valueMax="57.2" valueMin="22.4">22.457.2 mm</quantity>
SL; 2 (fixed in ethanol);
<collectingCountry id="27DE69C6FFA6FF9AFC6BF8A1FBE29FA6" box="[970,1029,1913,1936]" name="Laos" pageId="1">Laos</collectingCountry>
:
<collectingRegion id="9D0DE7B4FFA6FF9AFBB1F8A0FAF79FA7" box="[1040,1296,1912,1937]" country="Laos" name="Khammouan" pageId="1">Khammouan ProvinCe</collectingRegion>
:
<collectingCounty id="B61751DAFFA6FF9AFABAF8A1FA7B9FA7" box="[1307,1436,1912,1937]" pageId="1">Xe Bangfai</collectingCounty>
drainage:
<collectingMunicipality id="BF12B32CFFA6FF9AFEA6F846FE049F80" box="[263,483,1949,1974]" pageId="1">Nam Kathang Gnai</collectingMunicipality>
on edge of
<location id="5A167F8DFFA6FF9AFDCCF845FCE99F80" LSID="urn:lsid:plazi:treatment:D7609840FFA6FF9CFF36F91DFAB599C3:5A167F8DFFA6FF9AFDCCF845FCE99F80" box="[621,782,1949,1974]" country="Laos" county="Xe Bangfai" latitude="17.651388" longLatPrecision="21" longitude="105.237495" municipality="Nam Kathang Gnai" name="Nakai Plateau" pageId="1" stateProvince="Khammouan ProvinCe">Nakai Plateau</location>
, near
<location id="5A167F8DFFA6FF9AFCF2F846FC2F9F83" LSID="urn:lsid:plazi:treatment:D7609840FFA6FF9CFF36F91DFAB599C3:5A167F8DFFA6FF9AFCF2F846FC2F9F83" box="[851,968,1950,1973]" country="Laos" county="Xe Bangfai" latitude="17.651388" longLatPrecision="21" longitude="105.237495" municipality="Nam Kathang Gnai" name="Ban Done" pageId="1" stateProvince="Khammouan ProvinCe">Ban Done</location>
;
<geoCoordinate id="3AFD4F91FFA6FF9AFC74F845FBBF9F80" box="[981,1112,1949,1974]" direction="north" orientation="latitude" pageId="1" precision="15" value="17.651388">17°39'05''N</geoCoordinate>
<geoCoordinate id="3AFD4F91FFA6FF9AFBC1F845FB179F80" box="[1120,1264,1949,1974]" direction="east" orientation="longitude" pageId="1" precision="15" value="105.237495">105°14'15''E</geoCoordinate>
;
<collectorName id="F23C4C80FFA6FF9AFB5CF846FA989F80" box="[1277,1407,1949,1974]" pageId="1">M. Kottelat</collectorName>
<emphasis id="6DBDF544FFA6FF9AFA29F846FF4A9FEF" italics="true" pageId="1">et al</emphasis>
.,
<date id="2B770F96FFA6FF9AFF63F819FEBC9FEF" box="[194,347,1985,2009]" pageId="1" value="2012-05-14">
<collectingDate id="3B33F67EFFA6FF9AFF63F819FEBC9FEF" box="[194,347,1985,2009]" pageId="1" value="2012-05-14">14 May 2012</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="0BB679DEFFA5FF99FF36FBA7FADF9CA3" box="[151,1336,1151,1173]" httpUri="https://zenodo.org/record/837125/files/figure.png" pageId="2" targetBox="[193,1374,220,1100]" targetPageId="2">
<paragraph id="5F762956FFA5FF99FF36FBA7FADF9CA3" blockId="2.[151,1336,1151,1173]" box="[151,1336,1151,1173]" pageId="2">
<emphasis id="6DBDF544FFA5FF99FF36FBA7FEF59CA3" bold="true" box="[151,274,1151,1173]" pageId="2">FIGURE 2.</emphasis>
<taxonomicName id="98C952D5FFA5FF99FEB8FBA7FE3F9CA3" authorityName="Kottelat" authorityYear="2017" box="[281,472,1151,1173]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA5FF99FEB8FBA7FE3F9CA3" box="[281,472,1151,1173]" italics="true" pageId="2">Schistura epixenos</emphasis>
</taxonomicName>
, MHNG 2767.086, holotype, 59.5 mm SL; Laos: Nakai Plateau: Nam Kathang Gnai.
</paragraph>
</caption>
<subSubSection id="17D37ADDFFA5FF99FF66FB64FE469DBF" pageId="2" type="diagnosis">
<paragraph id="5F762956FFA5FF99FF66FB64FE469DBF" blockId="2.[151,1437,1212,2029]" pageId="2">
<emphasis id="6DBDF544FFA5FF99FF66FB64FEA69CE3" bold="true" box="[199,321,1212,1237]" pageId="2">Diagnosis.</emphasis>
<taxonomicName id="98C952D5FFA5FF99FEEDFB64FDC49CE2" authorityName="Kottelat" authorityYear="2017" box="[332,547,1212,1236]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA5FF99FEEDFB64FDC49CE2" box="[332,547,1212,1236]" italics="true" pageId="2">Schistura epixenos</emphasis>
</taxonomicName>
is distinguished from the other speCies of the genus in the Xe Bangfai and adjaCent drainages in Central and southern
<collectingCountry id="27DE69C6FFA5FF99FDD1FB39FD419CCE" box="[624,678,1249,1272]" name="Laos" pageId="2">Laos</collectingCountry>
by the following Combination of CharaCters: 1317 dark brown bars on a pale brown body, about as wide as interspaCes, reaChing downwards to the level of peCtoral-fin base; lateral line Complete; proCessus dentiformis narrow, pointed; depth of Caudal pedunCle 1.31.5 times in its length; body slender (13.715.1 % SL); sCales present in predorsal area; pelviC fin not reaChing anus; and pelviC-fin origin below dorsal-fin origin.
</paragraph>
</subSubSection>
<subSubSection id="17D37ADDFFA5FF9CFF66FA4CFDAE98CE" lastPageId="7" pageId="2" type="description">
<paragraph id="5F762956FFA5FF99FF66FA4CFDF89EFB" blockId="2.[151,1437,1212,2029]" pageId="2">
<emphasis id="6DBDF544FFA5FF99FF66FA4CFEBE9D9B" bold="true" box="[199,345,1428,1453]" pageId="2">Description.</emphasis>
See
<figureCitation id="C7F235D3FFA5FF99FE35FA4CFDFB9D9B" box="[404,540,1428,1453]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1151,1173]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="5.[151,250,968,990]" captionTargetBox-0="[193,1374,220,1100]" captionTargetBox-1="[198,1374,214,1940]" captionTargetBox-2="[161,1400,346,930]" captionTargetId-0="figure@2.[159,1407,193,1129]" captionTargetId-1="figure@3.[189,1398,193,1951]" captionTargetId-2="figure@5.[150,1436,301,947]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 2. Schistura epixenos, MHNG 2767.086, holotype, 59.5 mm SL; Laos: Nakai Plateau: Nam Kathang Gnai." captionText-1="FIGURE 3. Schistura epixenos, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai; a, 56.3 mm SL; b, 57.2 mm SL (right side, reversed); c, 57.0 mm SL (right side, reversed)." captionText-2="FIGURE 4. Schistura epixenos, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai; a, 22.4 mm SL; b, 29.2 mm SL (right side, reversed)." httpUri-0="https://zenodo.org/record/837125/files/figure.png" httpUri-1="https://zenodo.org/record/837127/files/figure.png" httpUri-2="https://zenodo.org/record/837129/files/figure.png" pageId="2">Figures 24</figureCitation>
for general appearanCe and
<tableCitation id="124B1CEDFFA5FF99FCC4FA4CFC5C9D9B" box="[869,955,1428,1453]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Morphometric data of type material of Schistura epixenos (n = 6). Range and mean include holotype data." pageId="2">Table 1</tableCitation>
for morphometriC data of
<typeStatus id="807297F4FFA5FF99FB51FA4CFAB59D9B" box="[1264,1362,1428,1453]" pageId="2" type="holotype">holotype</typeStatus>
and
<specimenCount id="49CFE2DFFFA5FF99FA2FFA4CFEE19DE7" pageId="2" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
. A moderately elongate nemaCheiline with body depth slowly inCreasing up to slightly in front of dorsalfin origin. Behind dorsal fin, body depth almost uniform until Caudal-fin base. Dorsal profile Continuous between head and body or with slight ConCavity. Head slightly depressed; body from slightly Compressed anteriorly to Compressed posteriorly. Interorbital area flat. In lateral view, eye flush with or below dorsal profile of head. Cheeks slightly swollen. Snout rounded in dorsal and lateral view. Depth of Caudal pedunCle uniform, 1.31.5 times in its length. Low dorsal keel on posterior third of post-dorsal area. Low ventral keel on posterior half of Caudal pedunCle. Dorsal keel Continuous with upper margin of Caudal fin, but with a shallow ConCavity at their junCtion. Largest reCorded siZe
<quantity id="983184B3FFA5FF99FE2CF96CFE169EFA" box="[397,497,1716,1741]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="2" unit="mm" value="59.5">59.5 mm</quantity>
SL.
</paragraph>
<paragraph id="5F762956FFA5FF99FF66F900FE749FDA" blockId="2.[151,1437,1212,2029]" pageId="2">Dorsal fin with 4 unbranChed and 8½ (6) branChed rays; 2nd or 3rd branChed ray longest; distal margin Convex. PeCtoral fin with 1 unbranChed and 11 (6) branChed rays (inCluding small last ray, usually unbranChed), rounded, reaChing ½–¾ of distanCe to pelviC-fin base. PelviC fin with 1 unbranChed and 7 (6) branChed rays (inCluding small last ray, usually unbranChed); reaChing to anus; triangular, posterior margin straight; origin under dorsal-fin origin; axillary lobe present, entirely free. Anus situated about 2.12.8 eye diameter in front of anal fin. Anal fin with 3 unbranChed and 5½ (6) branChed rays; distal margin straight to slightly Convex. Caudal fin with 9+8 (6) branChed rays; about 67 dorsal and 45 ventral proCurrent rays; forked, lobes rounded, subequal, upper lobe usually slightly longer than lower one.</paragraph>
<caption id="0BB679DEFFA4FF98FF36F86DFCE89FDC" httpUri="https://zenodo.org/record/837127/files/figure.png" pageId="3" targetBox="[198,1374,214,1940]" targetPageId="3">
<paragraph id="5F762956FFA4FF98FF36F86DFCE89FDC" blockId="3.[151,1435,1973,2026]" pageId="3">
<emphasis id="6DBDF544FFA4FF98FF36F86DFEF59FFD" bold="true" box="[151,274,1973,1995]" pageId="3">FIGURE 3.</emphasis>
<taxonomicName id="98C952D5FFA4FF98FEB9F86DFE319FFD" authorityName="Kottelat" authorityYear="2017" box="[280,470,1973,1995]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA4FF98FEB9F86DFE319FFD" box="[280,470,1973,1995]" italics="true" pageId="3">Schistura epixenos</emphasis>
</taxonomicName>
, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai;
<emphasis id="6DBDF544FFA4FF98FB24F86DFB7F9FFD" bold="true" box="[1157,1176,1973,1995]" pageId="3">a,</emphasis>
56.3 mm SL;
<emphasis id="6DBDF544FFA4FF98FA89F86DFADB9FFD" bold="true" box="[1320,1340,1973,1995]" pageId="3">b,</emphasis>
57.2 mm SL (right side, reversed);
<emphasis id="6DBDF544FFA4FF98FE3BF80CFE4C9FDC" bold="true" box="[410,427,2004,2026]" pageId="3">c,</emphasis>
57.0 mm SL (right side, reversed).
</paragraph>
</caption>
<caption id="0BB679DEFFA3FF9FFF36FF40FA8A9899" box="[151,1389,151,175]" pageId="4">
<paragraph id="5F762956FFA3FF9FFF36FF40FA8A9899" blockId="4.[151,1389,151,175]" box="[151,1389,151,175]" pageId="4">
<emphasis id="6DBDF544FFA3FF9FFF36FF40FEED9899" bold="true" box="[151,266,152,175]" pageId="4">TABLE 1.</emphasis>
Morphometric data of type material of
<taxonomicName id="98C952D5FFA3FF9FFD11FF4FFC9F9899" authorityName="Kottelat" authorityYear="2017" box="[688,888,151,175]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA3FF9FFD11FF4FFC9F9899" box="[688,888,151,175]" italics="true" pageId="4">Schistura epixenos</emphasis>
</taxonomicName>
(n=6). Range and mean include holotype data.
</paragraph>
</caption>
<paragraph id="5F762956FFA3FF9FFC83FF14FAED9E69" pageId="4">
<table id="2DC9DBF6FFA30064FF3EFF14FACE9E69" box="[159,1321,204,1631]" gridcols="4" gridrows="35" pageId="4">
<tr id="E1F92B14FFA30064FF3EFF14FACE98D4" box="[159,1321,204,226]" gridrow="0" pageId="4" rowspan-0="1">
<th id="A2284268FFA30064FC83FF14FC9C98D4" box="[802,891,204,226]" gridcol="1" gridrow="0" pageId="4">holotype</th>
<th id="A2284268FFA30064FC4EFF14FB8998D4" box="[1007,1134,204,226]" gridcol="2" gridrow="0" pageId="4">range</th>
<th id="A2284268FFA30064FB51FF14FACE98D4" box="[1264,1321,204,226]" gridcol="3" gridrow="0" pageId="4">mean</th>
</tr>
<tr id="E1F92B14FFA30064FF3EFF2DFACE993D" box="[159,1321,245,267]" gridrow="1" pageId="4" rowspan-3="1">
<th id="A2284268FFA30064FF3EFF2DFE11993D" box="[159,502,245,267]" gridcol="0" gridrow="1" pageId="4">Standard length (mm)</th>
<td id="A2284268FFA30064FC83FF2DFC9C993D" box="[802,891,245,267]" gridcol="1" gridrow="1" pageId="4">59.5</td>
<td id="A2284268FFA30064FC4EFF2DFB89993D" box="[1007,1134,245,267]" gridcol="2" gridrow="1" pageId="4">48.859.5</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFEC6FACE9902" box="[159,1321,286,308]" gridrow="2" pageId="4" rowspan-3="1">
<th id="A2284268FFA30064FF3EFEC6FE119902" box="[159,502,286,308]" gridcol="0" gridrow="2" pageId="4">Total length (mm)</th>
<td id="A2284268FFA30064FC83FEC6FC9C9902" box="[802,891,286,308]" gridcol="1" gridrow="2" pageId="4">72.8</td>
<td id="A2284268FFA30064FC4EFEC6FB899902" box="[1007,1134,286,308]" gridcol="2" gridrow="2" pageId="4">59.372.8</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFE90FACE9968" box="[159,1321,328,350]" gridrow="3" pageId="4" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="A2284268FFA30064FF3EFE90FE119968" box="[159,502,328,350]" gridcol="0" gridrow="3" pageId="4">In percent of standard length</th>
</tr>
<tr id="E1F92B14FFA30064FF3EFEA9FACE99B1" box="[159,1321,369,391]" gridrow="4" pageId="4">
<th id="A2284268FFA30064FF3EFEA9FE1199B1" box="[159,502,369,391]" gridcol="0" gridrow="4" pageId="4">Total length</th>
<td id="A2284268FFA30064FC83FEA9FC9C99B1" box="[802,891,369,391]" gridcol="1" gridrow="4" pageId="4">122.3</td>
<td id="A2284268FFA30064FC4EFEA9FB8999B1" box="[1007,1134,369,391]" gridcol="2" gridrow="4" pageId="4">121.1124.3</td>
<td id="A2284268FFA30064FB51FEA9FACE99B1" box="[1264,1321,369,391]" gridcol="3" gridrow="4" pageId="4">122.5</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFE42FACE9986" box="[159,1321,410,432]" gridrow="5" pageId="4">
<th id="A2284268FFA30064FF3EFE42FE119986" box="[159,502,410,432]" gridcol="0" gridrow="5" pageId="4">Head length (dorsal)</th>
<td id="A2284268FFA30064FC83FE42FC9C9986" box="[802,891,410,432]" gridcol="1" gridrow="5" pageId="4">19.4</td>
<td id="A2284268FFA30064FC4EFE42FB899986" box="[1007,1134,410,432]" gridcol="2" gridrow="5" pageId="4">18.721.2</td>
<td id="A2284268FFA30064FB51FE42FACE9986" box="[1264,1321,410,432]" gridcol="3" gridrow="5" pageId="4">20.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFE1CFACE99EC" box="[159,1321,452,474]" gridrow="6" pageId="4">
<th id="A2284268FFA30064FF3EFE1CFE1199EC" box="[159,502,452,474]" gridcol="0" gridrow="6" pageId="4">Head length (lateral)</th>
<td id="A2284268FFA30064FC83FE1CFC9C99EC" box="[802,891,452,474]" gridcol="1" gridrow="6" pageId="4">23.7</td>
<td id="A2284268FFA30064FC4EFE1CFB8999EC" box="[1007,1134,452,474]" gridcol="2" gridrow="6" pageId="4">21.824.1</td>
<td id="A2284268FFA30064FB51FE1CFACE99EC" box="[1264,1321,452,474]" gridcol="3" gridrow="6" pageId="4">23.3</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFE35FACE9A35" box="[159,1321,493,515]" gridrow="7" pageId="4">
<th id="A2284268FFA30064FF3EFE35FE119A35" box="[159,502,493,515]" gridcol="0" gridrow="7" pageId="4">Predorsal length</th>
<td id="A2284268FFA30064FC83FE35FC9C9A35" box="[802,891,493,515]" gridcol="1" gridrow="7" pageId="4">51.5</td>
<td id="A2284268FFA30064FC4EFE35FB899A35" box="[1007,1134,493,515]" gridcol="2" gridrow="7" pageId="4">51.554.7</td>
<td id="A2284268FFA30064FB51FE35FACE9A35" box="[1264,1321,493,515]" gridcol="3" gridrow="7" pageId="4">53.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFDCEFACE9A1A" box="[159,1321,534,556]" gridrow="8" pageId="4">
<th id="A2284268FFA30064FF3EFDCEFE119A1A" box="[159,502,534,556]" gridcol="0" gridrow="8" pageId="4">Prepelvic length</th>
<td id="A2284268FFA30064FC83FDCEFC9C9A1A" box="[802,891,534,556]" gridcol="1" gridrow="8" pageId="4">49.8</td>
<td id="A2284268FFA30064FC4EFDCEFB899A1A" box="[1007,1134,534,556]" gridcol="2" gridrow="8" pageId="4">49.852.2</td>
<td id="A2284268FFA30064FB51FDCEFACE9A1A" box="[1264,1321,534,556]" gridcol="3" gridrow="8" pageId="4">50.9</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFD98FACE9A60" box="[159,1321,576,598]" gridrow="9" pageId="4">
<th id="A2284268FFA30064FF3EFD98FE119A60" box="[159,502,576,598]" gridcol="0" gridrow="9" pageId="4">Pre-anus length</th>
<td id="A2284268FFA30064FC83FD98FC9C9A60" box="[802,891,576,598]" gridcol="1" gridrow="9" pageId="4">67.1</td>
<td id="A2284268FFA30064FC4EFD98FB899A60" box="[1007,1134,576,598]" gridcol="2" gridrow="9" pageId="4">67.169.1</td>
<td id="A2284268FFA30064FB51FD98FACE9A60" box="[1264,1321,576,598]" gridcol="3" gridrow="9" pageId="4">68.3</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFDB1FACE9A49" box="[159,1321,617,639]" gridrow="10" pageId="4">
<th id="A2284268FFA30064FF3EFDB1FE119A49" box="[159,502,617,639]" gridcol="0" gridrow="10" pageId="4">Pre-anal length</th>
<td id="A2284268FFA30064FC83FDB1FC9C9A49" box="[802,891,617,639]" gridcol="1" gridrow="10" pageId="4">76.4</td>
<td id="A2284268FFA30064FC4EFDB1FB899A49" box="[1007,1134,617,639]" gridcol="2" gridrow="10" pageId="4">76.478.0</td>
<td id="A2284268FFA30064FB51FDB1FACE9A49" box="[1264,1321,617,639]" gridcol="3" gridrow="10" pageId="4">77.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFD4AFACE9A9E" box="[159,1321,658,680]" gridrow="11" pageId="4">
<th id="A2284268FFA30064FF3EFD4AFE119A9E" box="[159,502,658,680]" gridcol="0" gridrow="11" pageId="4">Head depth</th>
<td id="A2284268FFA30064FC83FD4AFC9C9A9E" box="[802,891,658,680]" gridcol="1" gridrow="11" pageId="4">11.9</td>
<td id="A2284268FFA30064FC4EFD4AFB899A9E" box="[1007,1134,658,680]" gridcol="2" gridrow="11" pageId="4">10.111.9</td>
<td id="A2284268FFA30064FB51FD4AFACE9A9E" box="[1264,1321,658,680]" gridcol="3" gridrow="11" pageId="4">11.2</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFD64FACE9AE4" box="[159,1321,700,722]" gridrow="12" pageId="4">
<th id="A2284268FFA30064FF3EFD64FE119AE4" box="[159,502,700,722]" gridcol="0" gridrow="12" pageId="4">Body depth at dorsal-fin origin</th>
<td id="A2284268FFA30064FC83FD64FC9C9AE4" box="[802,891,700,722]" gridcol="1" gridrow="12" pageId="4">14.9</td>
<td id="A2284268FFA30064FC4EFD64FB899AE4" box="[1007,1134,700,722]" gridcol="2" gridrow="12" pageId="4">13.715.1</td>
<td id="A2284268FFA30064FB51FD64FACE9AE4" box="[1264,1321,700,722]" gridcol="3" gridrow="12" pageId="4">14.7</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFD3DFACE9ACD" box="[159,1321,741,763]" gridrow="13" pageId="4">
<th id="A2284268FFA30064FF3EFD3DFE119ACD" box="[159,502,741,763]" gridcol="0" gridrow="13" pageId="4">Depth of caudal peduncle</th>
<td id="A2284268FFA30064FC83FD3DFC9C9ACD" box="[802,891,741,763]" gridcol="1" gridrow="13" pageId="4">11.8</td>
<td id="A2284268FFA30064FC4EFD3DFB899ACD" box="[1007,1134,741,763]" gridcol="2" gridrow="13" pageId="4">11.211.8</td>
<td id="A2284268FFA30064FB51FD3DFACE9ACD" box="[1264,1321,741,763]" gridcol="3" gridrow="13" pageId="4">11.4</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFCD6FACE9B12" box="[159,1321,782,804]" gridrow="14" pageId="4">
<th id="A2284268FFA30064FF3EFCD6FE119B12" box="[159,502,782,804]" gridcol="0" gridrow="14" pageId="4">Length of caudal peduncle</th>
<td id="A2284268FFA30064FC83FCD6FC9C9B12" box="[802,891,782,804]" gridcol="1" gridrow="14" pageId="4">17.2</td>
<td id="A2284268FFA30064FC4EFCD6FB899B12" box="[1007,1134,782,804]" gridcol="2" gridrow="14" pageId="4">14.517.2</td>
<td id="A2284268FFA30064FB51FCD6FACE9B12" box="[1264,1321,782,804]" gridcol="3" gridrow="14" pageId="4">16.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFCE0FACE9B78" box="[159,1321,824,846]" gridrow="15" pageId="4">
<th id="A2284268FFA30064FF3EFCE0FE119B78" box="[159,502,824,846]" gridcol="0" gridrow="15" pageId="4">Head width</th>
<td id="A2284268FFA30064FC83FCE0FC9C9B78" box="[802,891,824,846]" gridcol="1" gridrow="15" pageId="4">15.6</td>
<td id="A2284268FFA30064FC4EFCE0FB899B78" box="[1007,1134,824,846]" gridcol="2" gridrow="15" pageId="4">14.316.2</td>
<td id="A2284268FFA30064FB51FCE0FACE9B78" box="[1264,1321,824,846]" gridcol="3" gridrow="15" pageId="4">15.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFCB9FACE9B41" box="[159,1321,865,887]" gridrow="16" pageId="4">
<th id="A2284268FFA30064FF3EFCB9FE119B41" box="[159,502,865,887]" gridcol="0" gridrow="16" pageId="4">Body width at dorsal-fin origin</th>
<td id="A2284268FFA30064FC83FCB9FC9C9B41" box="[802,891,865,887]" gridcol="1" gridrow="16" pageId="4">13.2</td>
<td id="A2284268FFA30064FC4EFCB9FB899B41" box="[1007,1134,865,887]" gridcol="2" gridrow="16" pageId="4">11.813.2</td>
<td id="A2284268FFA30064FB51FCB9FACE9B41" box="[1264,1321,865,887]" gridcol="3" gridrow="16" pageId="4">12.4</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFC52FACE9B96" box="[159,1321,906,928]" gridrow="17" pageId="4">
<th id="A2284268FFA30064FF3EFC52FE119B96" box="[159,502,906,928]" gridcol="0" gridrow="17" pageId="4">Snout length</th>
<td id="A2284268FFA30064FC83FC52FC9C9B96" box="[802,891,906,928]" gridcol="1" gridrow="17" pageId="4">9.3</td>
<td id="A2284268FFA30064FC4EFC52FB899B96" box="[1007,1134,906,928]" gridcol="2" gridrow="17" pageId="4">9.19.7</td>
<td id="A2284268FFA30064FB51FC52FACE9B96" box="[1264,1321,906,928]" gridcol="3" gridrow="17" pageId="4">9.4</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFC6CFACE9BFC" box="[159,1321,948,970]" gridrow="18" pageId="4">
<th id="A2284268FFA30064FF3EFC6CFE119BFC" box="[159,502,948,970]" gridcol="0" gridrow="18" pageId="4">Eye diameter</th>
<td id="A2284268FFA30064FC83FC6CFC9C9BFC" box="[802,891,948,970]" gridcol="1" gridrow="18" pageId="4">3.7</td>
<td id="A2284268FFA30064FC4EFC6CFB899BFC" box="[1007,1134,948,970]" gridcol="2" gridrow="18" pageId="4">3.44.0</td>
<td id="A2284268FFA30064FB51FC6CFACE9BFC" box="[1264,1321,948,970]" gridcol="3" gridrow="18" pageId="4">3.7</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFC05FACE9BC5" box="[159,1321,989,1011]" gridrow="19" pageId="4">
<th id="A2284268FFA30064FF3EFC05FE119BC5" box="[159,502,989,1011]" gridcol="0" gridrow="19" pageId="4">Interorbital width</th>
<td id="A2284268FFA30064FC83FC05FC9C9BC5" box="[802,891,989,1011]" gridcol="1" gridrow="19" pageId="4">8.2</td>
<td id="A2284268FFA30064FC4EFC05FB899BC5" box="[1007,1134,989,1011]" gridcol="2" gridrow="19" pageId="4">7.48.2</td>
<td id="A2284268FFA30064FB51FC05FACE9BC5" box="[1264,1321,989,1011]" gridcol="3" gridrow="19" pageId="4">7.6</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFBDEFACE9C2A" box="[159,1321,1030,1052]" gridrow="20" pageId="4">
<th id="A2284268FFA30064FF3EFBDEFE119C2A" box="[159,502,1030,1052]" gridcol="0" gridrow="20" pageId="4">Length of dorsal fin</th>
<td id="A2284268FFA30064FC83FBDEFC9C9C2A" box="[802,891,1030,1052]" gridcol="1" gridrow="20" pageId="4">18.0</td>
<td id="A2284268FFA30064FC4EFBDEFB899C2A" box="[1007,1134,1030,1052]" gridcol="2" gridrow="20" pageId="4">15.520.1</td>
<td id="A2284268FFA30064FB51FBDEFACE9C2A" box="[1264,1321,1030,1052]" gridcol="3" gridrow="20" pageId="4">18.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFBE8FACE9C70" box="[159,1321,1072,1094]" gridrow="21" pageId="4">
<th id="A2284268FFA30064FF3EFBE8FE119C70" box="[159,502,1072,1094]" gridcol="0" gridrow="21" pageId="4">Length of upper caudal-fin lobe</th>
<td id="A2284268FFA30064FC83FBE8FC9C9C70" box="[802,891,1072,1094]" gridcol="1" gridrow="21" pageId="4">21.5</td>
<td id="A2284268FFA30064FC4EFBE8FB899C70" box="[1007,1134,1072,1094]" gridcol="2" gridrow="21" pageId="4">21.223.4</td>
<td id="A2284268FFA30064FB51FBE8FACE9C70" box="[1264,1321,1072,1094]" gridcol="3" gridrow="21" pageId="4">22.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFB81FACE9C59" box="[159,1321,1113,1135]" gridrow="22" pageId="4">
<th id="A2284268FFA30064FF3EFB81FE119C59" box="[159,502,1113,1135]" gridcol="0" gridrow="22" pageId="4">Length of median caudal-fin rays</th>
<td id="A2284268FFA30064FC83FB81FC9C9C59" box="[802,891,1113,1135]" gridcol="1" gridrow="22" pageId="4">16.8</td>
<td id="A2284268FFA30064FC4EFB81FB899C59" box="[1007,1134,1113,1135]" gridcol="2" gridrow="22" pageId="4">16.518.2</td>
<td id="A2284268FFA30064FB51FB81FACE9C59" box="[1264,1321,1113,1135]" gridcol="3" gridrow="22" pageId="4">17.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFB5AFACE9CAE" box="[159,1321,1154,1176]" gridrow="23" pageId="4">
<th id="A2284268FFA30064FF3EFB5AFE119CAE" box="[159,502,1154,1176]" gridcol="0" gridrow="23" pageId="4">Length of lower caudal-fin lobe</th>
<td id="A2284268FFA30064FC83FB5AFC9C9CAE" box="[802,891,1154,1176]" gridcol="1" gridrow="23" pageId="4">20.2</td>
<td id="A2284268FFA30064FC4EFB5AFB899CAE" box="[1007,1134,1154,1176]" gridcol="2" gridrow="23" pageId="4">20.223.2</td>
<td id="A2284268FFA30064FB51FB5AFACE9CAE" box="[1264,1321,1154,1176]" gridcol="3" gridrow="23" pageId="4">21.6</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFB74FACE9CF4" box="[159,1321,1196,1218]" gridrow="24" pageId="4">
<th id="A2284268FFA30064FF3EFB74FE119CF4" box="[159,502,1196,1218]" gridcol="0" gridrow="24" pageId="4">Length of anal fin</th>
<td id="A2284268FFA30064FC83FB74FC9C9CF4" box="[802,891,1196,1218]" gridcol="1" gridrow="24" pageId="4">17.4</td>
<td id="A2284268FFA30064FC4EFB74FB899CF4" box="[1007,1134,1196,1218]" gridcol="2" gridrow="24" pageId="4">15.817.5</td>
<td id="A2284268FFA30064FB51FB74FACE9CF4" box="[1264,1321,1196,1218]" gridcol="3" gridrow="24" pageId="4">16.9</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFB0DFACE9CDD" box="[159,1321,1237,1259]" gridrow="25" pageId="4">
<th id="A2284268FFA30064FF3EFB0DFE119CDD" box="[159,502,1237,1259]" gridcol="0" gridrow="25" pageId="4">Length of pelvic fin</th>
<td id="A2284268FFA30064FC83FB0DFC9C9CDD" box="[802,891,1237,1259]" gridcol="1" gridrow="25" pageId="4">16.7</td>
<td id="A2284268FFA30064FC4EFB0DFB899CDD" box="[1007,1134,1237,1259]" gridcol="2" gridrow="25" pageId="4">16.717.7</td>
<td id="A2284268FFA30064FB51FB0DFACE9CDD" box="[1264,1321,1237,1259]" gridcol="3" gridrow="25" pageId="4">17.1</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFB26FACE9D22" box="[159,1321,1278,1300]" gridrow="26" pageId="4">
<th id="A2284268FFA30064FF3EFB26FE119D22" box="[159,502,1278,1300]" gridcol="0" gridrow="26" pageId="4">Length of pectoral fin</th>
<td id="A2284268FFA30064FC83FB26FC9C9D22" box="[802,891,1278,1300]" gridcol="1" gridrow="26" pageId="4">18.7</td>
<td id="A2284268FFA30064FC4EFB26FB899D22" box="[1007,1134,1278,1300]" gridcol="2" gridrow="26" pageId="4">18.120.9</td>
<td id="A2284268FFA30064FB51FB26FACE9D22" box="[1264,1321,1278,1300]" gridcol="3" gridrow="26" pageId="4">19.2</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFAF0FACE9D08" box="[159,1321,1320,1342]" gridrow="27" pageId="4" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="A2284268FFA30064FF3EFAF0FE119D08" box="[159,502,1320,1342]" gridcol="0" gridrow="27" pageId="4">In percent of dorsal head length</th>
</tr>
<tr id="E1F92B14FFA30064FF3EFA89FACE9D51" box="[159,1321,1361,1383]" gridrow="28" pageId="4">
<th id="A2284268FFA30064FF3EFA89FE119D51" box="[159,502,1361,1383]" gridcol="0" gridrow="28" pageId="4">Snout length</th>
<td id="A2284268FFA30064FC83FA89FC9C9D51" box="[802,891,1361,1383]" gridcol="1" gridrow="28" pageId="4">48</td>
<td id="A2284268FFA30064FC4EFA89FB899D51" box="[1007,1134,1361,1383]" gridcol="2" gridrow="28" pageId="4">4449</td>
<td id="A2284268FFA30064FB51FA89FACE9D51" box="[1264,1321,1361,1383]" gridcol="3" gridrow="28" pageId="4">47</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFAA2FACE9DA6" box="[159,1321,1402,1424]" gridrow="29" pageId="4">
<th id="A2284268FFA30064FF3EFAA2FE119DA6" box="[159,502,1402,1424]" gridcol="0" gridrow="29" pageId="4">Eye diameter</th>
<td id="A2284268FFA30064FC83FAA2FC9C9DA6" box="[802,891,1402,1424]" gridcol="1" gridrow="29" pageId="4">19</td>
<td id="A2284268FFA30064FC4EFAA2FB899DA6" box="[1007,1134,1402,1424]" gridcol="2" gridrow="29" pageId="4">1719</td>
<td id="A2284268FFA30064FB51FAA2FACE9DA6" box="[1264,1321,1402,1424]" gridcol="3" gridrow="29" pageId="4">19</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFA7CFACE9D8C" box="[159,1321,1444,1466]" gridrow="30" pageId="4">
<th id="A2284268FFA30064FF3EFA7CFE119D8C" box="[159,502,1444,1466]" gridcol="0" gridrow="30" pageId="4">Interorbital width</th>
<td id="A2284268FFA30064FC83FA7CFC9C9D8C" box="[802,891,1444,1466]" gridcol="1" gridrow="30" pageId="4">42</td>
<td id="A2284268FFA30064FC4EFA7CFB899D8C" box="[1007,1134,1444,1466]" gridcol="2" gridrow="30" pageId="4">3542</td>
<td id="A2284268FFA30064FB51FA7CFACE9D8C" box="[1264,1321,1444,1466]" gridcol="3" gridrow="30" pageId="4">38</td>
</tr>
<tr id="E1F92B14FFA30064FF3EFA15FACE9DD5" box="[159,1321,1485,1507]" gridrow="31" pageId="4" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="A2284268FFA30064FF3EFA15FE119DD5" box="[159,502,1485,1507]" gridcol="0" gridrow="31" pageId="4">In percent of lateral head length</th>
</tr>
<tr id="E1F92B14FFA30064FF3EFA2EFACE9E3A" box="[159,1321,1526,1548]" gridrow="32" pageId="4">
<th id="A2284268FFA30064FF3EFA2EFE119E3A" box="[159,502,1526,1548]" gridcol="0" gridrow="32" pageId="4">Snout length</th>
<td id="A2284268FFA30064FC83FA2EFC9C9E3A" box="[802,891,1526,1548]" gridcol="1" gridrow="32" pageId="4">39</td>
<td id="A2284268FFA30064FC4EFA2EFB899E3A" box="[1007,1134,1526,1548]" gridcol="2" gridrow="32" pageId="4">3843</td>
<td id="A2284268FFA30064FB51FA2EFACE9E3A" box="[1264,1321,1526,1548]" gridcol="3" gridrow="32" pageId="4">40</td>
</tr>
<tr id="E1F92B14FFA30064FF3EF9F8FACE9E00" box="[159,1321,1568,1590]" gridrow="33" pageId="4">
<th id="A2284268FFA30064FF3EF9F8FE119E00" box="[159,502,1568,1590]" gridcol="0" gridrow="33" pageId="4">Eye diameter</th>
<td id="A2284268FFA30064FC83F9F8FC9C9E00" box="[802,891,1568,1590]" gridcol="1" gridrow="33" pageId="4">16</td>
<td id="A2284268FFA30064FC4EF9F8FB899E00" box="[1007,1134,1568,1590]" gridcol="2" gridrow="33" pageId="4">1517</td>
<td id="A2284268FFA30064FB51F9F8FACE9E00" box="[1264,1321,1568,1590]" gridcol="3" gridrow="33" pageId="4">16</td>
</tr>
<tr id="E1F92B14FFA30064FF3EF991FACE9E69" box="[159,1321,1609,1631]" gridrow="34" pageId="4">
<th id="A2284268FFA30064FF3EF991FE119E69" box="[159,502,1609,1631]" gridcol="0" gridrow="34" pageId="4">Interorbital width</th>
<td id="A2284268FFA30064FC83F991FC9C9E69" box="[802,891,1609,1631]" gridcol="1" gridrow="34" pageId="4">34</td>
<td id="A2284268FFA30064FC4EF991FB899E69" box="[1007,1134,1609,1631]" gridcol="2" gridrow="34" pageId="4">3134</td>
<td id="A2284268FFA30064FB51F991FACE9E69" box="[1264,1321,1609,1631]" gridcol="3" gridrow="34" pageId="4">33</td>
</tr>
</table>
</paragraph>
<paragraph id="5F762956FFA3FF9FFF66F97EFD3B9F1C" blockId="4.[151,1437,1702,2015]" pageId="4">Body entirely Covered by sCales, exCept on belly between bases of peCtoral fins. SCales deeply embedded, sparsely set in anterior extremity of predorsal area. Lateral line almost Complete, ending on last brown bar on Caudal pedunCle, with 8895 pores. CephaliC lateral line system with 6 supraorbital, 4 + 11 infraorbital, 9 preoperCulo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="5F762956FFA3FF9EFF66F8EEFDA998CE" blockId="4.[151,1437,1702,2015]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" pageId="4">
Anterior naris pierCed in front side of a pointed flap-like tube. Posterior naris adjaCent to anterior one. Mouth arChed, gape about 2½ times wider than long (
<figureCitation id="C7F235D3FFA3FF9FFD0DF882FD159F45" box="[684,754,1882,1907]" captionStart="FIGURE 5" captionStartId="5.[151,250,1945,1967]" captionTargetBox="[178,1409,1172,1923]" captionTargetId="figure@5.[178,1409,1101,1923]" captionTargetPageId="5" captionText="FIGURE 5. Schistura epixenos, CMK 22962, paratype, 57.0 mm SL; mouth." httpUri="https://zenodo.org/record/837131/files/figure.png" pageId="4">Fig. 5</figureCitation>
). Lips thiCk. Upper lip with small to barely distinCt median notCh, with shallow and thin furrows, edge finely Crenulated. ProCessus dentiformis present. Lower lip with median interruption; median part with 04 weekly-marked sulCi; lateral part not ConspiCuously narrower than median part, smooth. Tip of lower jaw not exposed. A shallow median ConCavity in lower jaw. Inner rostral barbel reaChing base of maxillary barbel; outer one reaChing about below middle of eye. Maxillary barbel reaChing middle of postorbital area. Intestine forming a loop at some distanCe behind stomaCh; stomaCh very slender (
<figureCitation id="C7F235D3FFA2FF9EFBC4FF64FB4D98E3" box="[1125,1194,188,213]" captionStart="FIGURE 6" captionStartId="6.[151,250,1327,1349]" captionTargetBox="[598,992,190,1300]" captionTargetId="figure@6.[585,1005,169,1311]" captionTargetPageId="6" captionText="FIGURE 6. Schistura epixenos, CMK 22962, paratype, 48.8 mm SL; digestive tract. Scale bar 1 mm." httpUri="https://zenodo.org/record/837133/files/figure.png" pageId="5">Fig. 6</figureCitation>
). Air bladder without posterior Chamber in abdominal Cavity.
</paragraph>
<caption id="0BB679DEFFA2FF9EFF36FC10FE739BCB" httpUri="https://zenodo.org/record/837129/files/figure.png" pageId="5" targetBox="[161,1400,346,930]" targetPageId="5">
<paragraph id="5F762956FFA2FF9EFF36FC10FE739BCB" blockId="5.[151,1435,968,1021]" pageId="5">
<emphasis id="6DBDF544FFA2FF9EFF36FC10FEF59BE8" bold="true" box="[151,274,968,990]" pageId="5">FIGURE 4.</emphasis>
<taxonomicName id="98C952D5FFA2FF9EFEB9FC10FE319BE8" authorityName="Kottelat" authorityYear="2017" box="[280,470,968,990]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA2FF9EFEB9FC10FE319BE8" box="[280,470,968,990]" italics="true" pageId="5">Schistura epixenos</emphasis>
</taxonomicName>
, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai;
<emphasis id="6DBDF544FFA2FF9EFB24FC10FB7F9BE8" bold="true" box="[1157,1176,968,990]" pageId="5">a,</emphasis>
22.4 mm SL;
<emphasis id="6DBDF544FFA2FF9EFA89FC10FADB9BE8" bold="true" box="[1320,1340,968,990]" pageId="5">b,</emphasis>
29.2 mm SL (right side, reversed).
</paragraph>
</caption>
<caption id="0BB679DEFFA2FF9EFF36F841FC409F99" box="[151,935,1945,1967]" httpUri="https://zenodo.org/record/837131/files/figure.png" pageId="5" targetBox="[178,1409,1172,1923]" targetPageId="5">
<paragraph id="5F762956FFA2FF9EFF36F841FC409F99" blockId="5.[151,935,1945,1967]" box="[151,935,1945,1967]" pageId="5">
<emphasis id="6DBDF544FFA2FF9EFF36F841FEF59F99" bold="true" box="[151,274,1945,1967]" pageId="5">FIGURE 5.</emphasis>
<taxonomicName id="98C952D5FFA2FF9EFEB8F841FE3F9F99" authorityName="Kottelat" authorityYear="2017" box="[281,472,1945,1967]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA2FF9EFEB8F841FE3F9F99" box="[281,472,1945,1967]" italics="true" pageId="5">Schistura epixenos</emphasis>
</taxonomicName>
, CMK 22962, paratype, 57.0 mm SL; mouth.
</paragraph>
</caption>
<caption id="0BB679DEFFA1FF9DFF36FAF7FB479D73" box="[151,1184,1327,1349]" httpUri="https://zenodo.org/record/837133/files/figure.png" pageId="6" targetBox="[598,992,190,1300]" targetPageId="6">
<paragraph id="5F762956FFA1FF9DFF36FAF7FB479D73" blockId="6.[151,1184,1327,1349]" box="[151,1184,1327,1349]" pageId="6">
<emphasis id="6DBDF544FFA1FF9DFF36FAF7FEF59D73" bold="true" box="[151,274,1327,1349]" pageId="6">FIGURE 6.</emphasis>
<taxonomicName id="98C952D5FFA1FF9DFEB8FAF7FE3F9D73" authorityName="Kottelat" authorityYear="2017" box="[281,472,1327,1349]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA1FF9DFEB8FAF7FE3F9D73" box="[281,472,1327,1349]" italics="true" pageId="6">Schistura epixenos</emphasis>
</taxonomicName>
, CMK 22962, paratype, 48.8 mm SL; digestive tract. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="5F762956FFA1FF9DFF66FAB4FF149D9F" blockId="6.[151,1437,1388,2025]" pageId="6">
<emphasis id="6DBDF544FFA1FF9DFF66FAB4FE5F9DB3" bold="true" box="[199,440,1388,1413]" pageId="6">Sexual dimorphism.</emphasis>
None observed. No speCimen with suborbital flap, groove or slit. No modified peCtoralfin rays.
</paragraph>
<paragraph id="5F762956FFA1FF9DFF66FA6CFA8A9EB6" blockId="6.[151,1437,1388,2025]" pageId="6">
<emphasis id="6DBDF544FFA1FF9DFF66FA6CFEA89DFB" bold="true" box="[199,335,1460,1485]" pageId="6">Coloration.</emphasis>
After one month in formalin. Head and body baCkground Colour yellowish brown; throat, belly and lower part of Caudal pedunCle paler. Unless otherwise stated, all markings dark brown. Top and sides of head dark brown, without distinCtive pattern. Body with 1317 bars (57 predorsal, 34 subdorsal, 57 postdorsal), from dorsal midline and reaChing downwards about to level of peCtoral fin, not Continuous or irregularly Continuous with Contralaterals over baCk; reaChing ventral midline on Caudal pedunCle in largest speCimens. Bars mainly of regular width and shape, from slightly wider to slightly narrower than interspaCes. A narrow inner axial stripe present.
</paragraph>
<paragraph id="5F762956FFA1FF9DFF66F955FE499F27" blockId="6.[151,1437,1388,2025]" pageId="6">Pattern at Caudal-fin base: blaCk bar at base of fin, denser blaCk along base of lowermost 12 rays of upper lobe and 4 uppermost rays of lower lobe; bar extending almost to dorsal and ventral midlines, widening to form a roughly triangular patCh at both ends, on upper and lower proCurrent rays; in some speCimens, upper triangle partly separate from rest of bar.</paragraph>
<paragraph id="5F762956FFA1FF9DFF66F8C4FD499F97" blockId="6.[151,1437,1388,2025]" pageId="6">All fins hyaline, with blaCk pigments along edges of rays and between segments; a small patCh of blaCk pigments on membranes between branChes at branChing points. Pigments more densely set on dorsal surfaCe of peCtoral and pelviC-fin rays. Dorsal fin with a small blaCk spot at base of simple rays and first branChed ray and a narrow grey band along base of remaining rays.</paragraph>
<paragraph id="5F762956FFA1FF9DFF66F874FB8A9FDF" blockId="6.[151,1437,1388,2025]" pageId="6">
Juveniles (only two speCimens, 22.4 and
<quantity id="983184B3FFA1FF9DFD37F875FD1C9FF2" box="[662,763,1965,1989]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="6" unit="mm" value="29.2">29.2 mm</quantity>
SL;
<figureCitation id="C7F235D3FFA1FF9DFC93F874FC909FF3" box="[818,887,1964,1989]" captionStart="FIGURE 4" captionStartId="5.[151,250,968,990]" captionTargetBox="[161,1400,346,930]" captionTargetId="figure@5.[150,1436,301,947]" captionTargetPageId="5" captionText="FIGURE 4. Schistura epixenos, CMK 22962, paratypes; Laos: Nakai Plateau: Nam Kathang Gnai; a, 22.4 mm SL; b, 29.2 mm SL (right side, reversed)." httpUri="https://zenodo.org/record/837129/files/figure.png" pageId="6">Fig. 4</figureCitation>
): bars not sharply Contrasted, irregularly shaped, several bars appearing as ConneCted in pairs at their upper extremity, or paler in middle.
</paragraph>
<paragraph id="5F762956FFA0FF9CFF66FF4FFDAE98CE" blockId="7.[151,1437,151,501]" pageId="7">
<emphasis id="6DBDF544FFA0FF9CFF66FF4FFE719886" bold="true" box="[199,406,151,176]" pageId="7">Notes on biology.</emphasis>
A disseCted female (CMK 22962,
<quantity id="983184B3FFA0FF9CFC89FF40FC699899" box="[808,910,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.88" pageId="7" unit="mm" value="48.8">48.8 mm</quantity>
SL) had an ovary with yellow (probably near ripe) eggs
<quantity id="983184B3FFA0FF9CFEB3FF64FE8D98E2" box="[274,362,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" unit="mm" value="0.8">0.8 mm</quantity>
diameter, in anterior part mixed with white ooCytes about same siZe, and organised in a long narrow string 23 egg-diameters wide.
</paragraph>
</subSubSection>
<subSubSection id="17D37ADDFFA0FF9CFF66FEDCFE8C999B" pageId="7" type="distribution">
<paragraph id="5F762956FFA0FF9CFF66FEDCFE8C999B" blockId="7.[151,1437,151,501]" pageId="7">
<emphasis id="6DBDF544FFA0FF9CFF66FEDCFE16992B" bold="true" box="[199,497,260,285]" pageId="7">Distribution and habitat.</emphasis>
<taxonomicName id="98C952D5FFA0FF9CFE56FEDCFD2E992A" authorityName="Kottelat" authorityYear="2017" box="[503,713,260,284]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="epixenos">
<emphasis id="6DBDF544FFA0FF9CFE56FEDCFD2E992A" box="[503,713,260,284]" italics="true" pageId="7">Schistura epixenos</emphasis>
</taxonomicName>
is presently known only from the Nam Kathang Gnai [also spelt Katang Yai] on the Nakai Plateau. At the sampling sites, the stream was under forest Cover, very narrow, with a dense Cover of branChes, bamboos and shrubs (
<figureCitation id="C7F235D3FFA0FF9CFD15FE94FD1D9952" box="[692,762,332,357]" captionStart="FIGURE 7" captionStartId="7.[151,250,1449,1471]" captionTargetBox="[148,1444,555,1416]" captionTargetId="figure@7.[148,1444,555,1416]" captionTargetPageId="7" captionText="FIGURE 7. Nam Kathang Gnai, Nakai Plateau, Laos; type locality of Schistura epixenos; 7 May 2012." httpUri="https://zenodo.org/record/837135/files/figure.png" pageId="7">Fig. 7</figureCitation>
). The water was shallow (130 Cm deep). The bottom was Covered by thiCk mud (probably the result of traCk ConstruCtion and reCent agriCultural work), at plaCes with thiCk layers of leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="17D37ADDFFA0FF9CFF66FE6FFAB599C3" pageId="7" type="etymology">
<paragraph id="5F762956FFA0FF9CFF66FE6FFAB599C3" blockId="7.[151,1437,151,501]" pageId="7">
<emphasis id="6DBDF544FFA0FF9CFF66FE6FFEA999E6" bold="true" box="[199,334,439,464]" pageId="7">Etymology.</emphasis>
From the ClassiCal Greek
<emphasis id="6DBDF544FFA0FF9CFD30FE6FFD0899F9" box="[657,751,439,463]" italics="true" pageId="7">ἐπίξενΟς</emphasis>
(
<taxonomicName id="98C952D5FFA0FF9CFCA2FE6FFC8D99E6" authorityName="Kottelat" authorityYear="2017" box="[771,874,439,464]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="epixenos">epixenos</taxonomicName>
) meaning stranger, from another Community, a referenCe to the 'stranger' status of this speCies among the fish fauna of Nakai Plateau. A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>