258 lines
26 KiB
XML
258 lines
26 KiB
XML
<document id="B268470F6C83FA0A7A58606161A19CC1" ID-CLB-Dataset="304167" ID-DOI="10.3897/zse.100.128717" ID-GBIF-Dataset="a7f81ef5-5178-48c8-8d40-80b29116daad" ID-ZooBank="FD4C02D2-01F9-4A4C-817B-F81B27F22A41" ID-publisher-id="128717" URI-arpha="A58A01B3-C620-5671-AB59-ABE7916A4ADE" URI-zoobank="http://zoobank.org/FD4C02D2-01F9-4A4C-817B-F81B27F22A41" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1728062842085" checkinUser="pensoft" docAuthor="Klongklaew, Kanyaporn, Poeaim, Supattra & Dumrongrojwattana, Pongrat" docDate="2024" docId="F2A0D22321315721BBF4612DE4450412" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1347-1360" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/128717/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Georissa quinquelirata Klongkaew, Poeaim & Dumrongrojwattana 2024, sp. nov." docType="treatment" docUuid="70D34F82-2195-4F29-89F2-24C53848CF95" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="1360" masterDocId="A58A01B3C6205671AB59ABE7916A4ADE" masterDocTitle="Four new species of Georissa W. Blanford, 1864 (Gastropoda, Hydrocenidae) from Thailand" masterLastPageNumber="1360" masterPageNumber="1347" pageNumber="1347" updateTime="1728065275675" updateUser="admin">
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<mods:mods id="3776C92677E5A6A69183D27AFA3CB624" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="36B7265B10B8D270BA67D968398EA477">
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<mods:title id="2C107A057BFED4EAB7E5F14BF738F0BE">Four new species of Georissa W. Blanford, 1864 (Gastropoda, Hydrocenidae) from Thailand</mods:title>
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<mods:name id="327D9720CB529AA95F9739CF20A72760" type="personal">
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<mods:roleTerm id="CC4ACE0B559248710D7D9E44E7761BA8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="769E42B1E7D03A7131FBB9BE1A4339C8">Klongklaew, Kanyaporn</mods:namePart>
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<mods:affiliation id="5F54F7714301F90361231E8DAAFFC44F">Department of Biology, School of Science, King Mongkut's Institute of Technology Ladkrabang, Bangkok 10520, Thailand</mods:affiliation>
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<mods:name id="62A18AD32B415E71CD84D469097428A1" type="personal">
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<mods:roleTerm id="0DE44EABFA66183AC1DE60D0EF10E3A6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A4F53E689462FB6B1F879548AB1CCDFB">Poeaim, Supattra</mods:namePart>
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<mods:nameIdentifier id="5399AB43B3DBA15A60F740ABFB89FF17" type="ORCID">0000-0003-1293-3820</mods:nameIdentifier>
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<mods:affiliation id="B98055E60CB916964FAA869B085FCCEC">Department of Biology, School of Science, King Mongkut's Institute of Technology Ladkrabang, Bangkok 10520, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="AB16CC61F17759FD4C5A1BDEA4196868" type="personal">
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<mods:roleTerm id="7A7A2928E0A437E1F31C6DC942C12E26">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9749F3D54167CDEA848510A360F53760">Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation id="9CB17A5496B61B02955FD175EE8354D5">Department of Biology, Faculty of Science, Burapha University, Bangsaen, Chonburi 20131, Thailand</mods:affiliation>
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<mods:typeOfResource id="FD7BE7C25997058CB44C86F2E14F540C">text</mods:typeOfResource>
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<mods:title id="6159A54B9A892FED6704812D1DE864D0">Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part id="4897DD37DBFE6AE168D0740BA9508D15">
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<mods:date id="DB84CAC7CDC9A29B2ED4C292E2AEFA1D">2024</mods:date>
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<mods:number id="5BC2CB902FB75D88618FA08381301CC7">2024-10-04</mods:number>
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<mods:detail id="0034BB6E341DEE8EE9F3FEAF7413C878" type="volume">
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<mods:number id="EC49032638E16F53352ECFABF4A5FB7C">100</mods:number>
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<mods:detail id="3F5630CDFA8DBC90F48C77BA411AA279" type="issue">
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<mods:number id="1F8B652E681FBC8CF46DECB29C5E6019">4</mods:number>
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<mods:start id="99F6F2B97707E84805CCAC3559E7EB30">1347</mods:start>
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<mods:end id="C73DC04B27DE4368EAF2D43B8B03DA60">1360</mods:end>
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</mods:extent>
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<mods:classification id="D7245123D263CA6C003E318EAF658485">journal article</mods:classification>
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<mods:identifier id="CB650F805FB248A57E73E971773C257E" type="DOI">10.3897/zse.100.128717</mods:identifier>
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<treatment id="F2A0D22321315721BBF4612DE4450412" ID-DOI="http://doi.org/10.5281/zenodo.13891559" ID-Zenodo-Dep="13891559" ID-arpha="F2A0D223-2131-5721-BBF4-612DE4450412" ID-zoobank="https://zoobank.org/70D34F82-2195-4F29-89F2-24C53848CF95" LSID="urn:lsid:zoobank.org:act:70D34F82-2195-4F29-89F2-24C53848CF95" LSID-ZBK="urn:lsid:zoobank.org:act:70D34F82-2195-4F29-89F2-24C53848CF95" httpUri="http://treatment.plazi.org/id/F2A0D22321315721BBF4612DE4450412">
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<subSubSection id="4C3A68E2E18FA547C339BD99A31F1605" type="nomenclature">
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<paragraph id="5F04D9049C470E8F381A80C6E627FB90">
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<taxonomicName id="F230D2C4660081179F4F4B597F330F85" authority="Klongkaew, Poeaim & Dumrongrojwattana" authorityName="Klongkaew, Poeaim & Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="quinquelirata" status="sp. nov.">
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<emphasis id="9F5E7700A98FCF8CA0CBABDFDC0B2DCD" italics="true">Georissa quinquelirata</emphasis>
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Klongkaew, Poeaim & Dumrongrojwattana
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</taxonomicName>
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<taxonomicNameLabel id="043B2CA1A68C7C992F8D3B335ED1807A" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="487D38792B04E48FCDF6DA91F146BF6A" type="description">
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||
<paragraph id="05E1E870CB44342FC8278C0A096CC16A">
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||
<figureCitation id="EA6F8A488A0088B1844D082A56749BD3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">Figs 2 A</figureCitation>
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,
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<figureCitation id="195E0634FFFB50457FC694E43155E6EC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 A – D</figureCitation>
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,
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<figureCitation id="7647EEABAF74B85A72B9FDD08755D56F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 A – D</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EREBG" type="type material">
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||
<paragraph id="394E00795A4D80CB1933B5F23B494F64">
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||
<heading id="F612905B288435126C9CA644278D49DC" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="63F7056FB1C168281F50FD9232CB023A">
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<materialsCitation id="A84915CE4883301E2C5BEA8827841399" collectingDate="2023-02-25" collectionCode="ZRCBUU" collectorName="P. Dumrongrojwattana" country="Thailand" county="Thap Than district" latitude="15.474138" location="Pathawi limestone hill" longLatPrecision="1" longitude="99.75694" specimenCode="ZRCBUU 0900" specimenCount="1" stateProvince="Uthai Thani Province" typeStatus="holotype">
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||
<emphasis id="F3B43C37E6B48564B55E2618214C0B30" bold="true" italics="true">
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||
<typeStatus id="BD73B7D965D947CB01C5703E11CF7E12" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
•
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<specimenCode id="CC06E787E206E5559A611FCCB3313046">
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||
<abbrev id="ABBRID0E3EBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
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||
0900
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||
</specimenCode>
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||
(Fig.
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<figureCitation id="F146365B0552B8EDDB18A89E900AC722" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 A</figureCitation>
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||
).
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||
<abbrev id="ABBRID0EEFBG" xlink_title="Shell height">SH</abbrev>
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=
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||
<quantity id="6D8372DD8808E1730B18FD6B5ED43924" metricMagnitude="-3" metricUnit="m" metricValue="2.45" unit="mm" value="2.45">2.45 mm</quantity>
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,
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||
<abbrev id="ABBRID0EIFBG" xlink_title="shell width">SW</abbrev>
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||
=
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<quantity id="A36010CE924CB161829B0CC6F1630961" metricMagnitude="-3" metricUnit="m" metricValue="1.63" unit="mm" value="1.63">1.63 mm</quantity>
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||
,
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||
<abbrev id="ABBRID0EMFBG" xlink_title="aperture height">AH</abbrev>
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||
=
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||
<quantity id="0450EAD3D3183A99DC5BC5D3D7A1C828" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.80 mm</quantity>
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||
,
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||
<abbrev id="ABBRID0EQFBG" xlink_title="aperture width">AW</abbrev>
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||
=
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||
<quantity id="1620A6C1B7FFEE93A0DD105AB600778E" metricMagnitude="-4" metricUnit="m" metricValue="9.1" unit="mm" value="0.91">0.91 mm</quantity>
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||
, W = 4.25–4.5
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||
</materialsCitation>
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||
.
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<materialsCitation id="BEE1F0D668DC5D2E7E73C9FE7F926AE7" collectingDate="2023-02-25" collectionCode="ZRCBUU" collectorName="P. Dumrongrojwattana" country="Thailand" county="Thap Than district" latitude="15.474138" location="Pathawi limestone hill" longLatPrecision="1" longitude="99.75694" specimenCode="ZRCBUU 0901" specimenCount="1" stateProvince="Uthai Thani Province" typeStatus="paratype">
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||
<emphasis id="54566CD9CA263D8A3D8B9B53CB5B1A5F" bold="true" italics="true">
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||
<typeStatus id="09EE47F4034ED6DE58EF0118D2E961B6" type="paratype">Paratype</typeStatus>
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||
</emphasis>
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||
•
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<specimenCode id="4D4C0009B9FB023AA41E2B1D8BF01E76">
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||
<abbrev id="ABBRID0EXFBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
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0901
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</specimenCode>
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(Fig.
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<figureCitation id="27DB3C67F75DBD4543AC8FF5230DFFC0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 A – D</figureCitation>
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) (13 shells); Shell measurements:
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<abbrev id="ABBRID0E6FBG" xlink_title="Shell height">SH</abbrev>
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=
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<quantity id="94DB809C772620E12F59B1D281D5BF11" metricMagnitude="-3" metricUnit="m" metricValue="2.135" metricValueMax="2.36" metricValueMin="1.91" unit="mm" value="2.135" valueMax="2.36" valueMin="1.91">1.91–2.36 mm</quantity>
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(2.10 ±
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||
<quantity id="94770DAF9C2FCA54B58DB680A1902D03" metricMagnitude="-4" metricUnit="m" metricValue="1.5" unit="mm" value="0.15">0.15 mm</quantity>
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||
),
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||
<abbrev id="ABBRID0EDGBG" xlink_title="shell width">SW</abbrev>
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=
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<quantity id="8CAE76193FA2368151CCF04F78DC350B" metricMagnitude="-3" metricUnit="m" metricValue="1.485" metricValueMax="1.6" metricValueMin="1.37" unit="mm" value="1.485" valueMax="1.6" valueMin="1.37">1.37–1.60 mm</quantity>
|
||
(1.46 ±
|
||
<quantity id="91AF8C23A94F6D6FF3D5A5456A944366" metricMagnitude="-5" metricUnit="m" metricValue="7" unit="mm" value="0.07">0.07 mm</quantity>
|
||
),
|
||
<abbrev id="ABBRID0EHGBG" xlink_title="aperture height">AH</abbrev>
|
||
=
|
||
<quantity id="B8622DDA948EF42EC278593CC2D34051" metricMagnitude="-4" metricUnit="m" metricValue="7.85" metricValueMax="8.6" metricValueMin="7.1" unit="mm" value="0.7849999999999999" valueMax="0.86" valueMin="0.71">0.71–0.86 mm</quantity>
|
||
(0.78 ±
|
||
<quantity id="339A7AA7118318B7EE1831D20F4A2067" metricMagnitude="-5" metricUnit="m" metricValue="6.0" unit="mm" value="0.06">0.06 mm</quantity>
|
||
),
|
||
<abbrev id="ABBRID0ELGBG" xlink_title="aperture width">AW</abbrev>
|
||
=
|
||
<quantity id="1D931C6D637FC7DE86403E62DEF7680D" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="9.2" metricValueMin="8.0" unit="mm" value="0.86" valueMax="0.92" valueMin="0.8">0.80–0.92 mm</quantity>
|
||
(0.86 ±
|
||
<quantity id="62FD19292DE3B722DB2D7616D6436652" metricMagnitude="-5" metricUnit="m" metricValue="4.0" unit="mm" value="0.04">0.04 mm</quantity>
|
||
), W = 3.75–4.5
|
||
</materialsCitation>
|
||
(all type material from type locality;
|
||
<date id="91305FE549D8F6308E366B1D0968AA23" value="2023-02-25">
|
||
<collectingDate id="2E34B25FB70798B67115CDC0A7975904" value="2023-02-25">25 February 2023</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="967583DC2DD7B3457DD7B458A9CF7874">P. Dumrongrojwattana</collectorName>
|
||
leg).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EPGBG" type="type locality">
|
||
<paragraph id="0BECF2753F677D7C8297359A0787CA58">
|
||
<heading id="9993F91B55B00D33DA08020A73B05776" reason="title">Type locality.</heading>
|
||
</paragraph>
|
||
<paragraph id="1512150407CFE7E250A026A359ECC0B1">
|
||
<collectingCountry id="F1DDD342FF022C622A48B67CEB0BB6CC" name="Thailand">Thailand</collectingCountry>
|
||
,
|
||
<location id="6BB2C7F2CC77846EE963ED0798307D2E" LSID="urn:lsid:plazi:treatment:F2A0D22321315721BBF4612DE4450412:6BB2C7F2CC77846EE963ED0798307D2E" country="Thailand" county="Thap Than district" latitude="15.474138" longLatPrecision="1" longitude="99.75694" name="Pathawi limestone hill" stateProvince="Uthai Thani Province">Pathawi limestone hill</location>
|
||
,
|
||
<collectingCounty id="7DBC0E7D1CD99F2B0C3329EC012D5071">Thap Than district</collectingCounty>
|
||
,
|
||
<collectingRegion id="A7774AB1B4E820A599A656823E3E811C" country="Thailand" name="Uthai Thani">Uthai Thani Province</collectingRegion>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYGBG" specific-use="{"type":"Point","coordinates":[99.756944,15.474139]}">
|
||
<geoCoordinate id="ABF6324DC614C1978A834E4094BCF9DF" degrees="15" direction="north" minutes="28" orientation="latitude" precision="1" seconds="26.9" value="15.474138">15 ° 28 ' 26.9 " N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="35E1B92DF5D92B2DD9DC869E3E43AF9A" degrees="99" direction="east" minutes="45" orientation="longitude" precision="1" seconds="25.0" value="99.75694">99 ° 45 ' 25.0 " E</geoCoordinate>
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</named-content>
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||
.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="SECID0E4GBG" type="etymology">
|
||
<paragraph id="8FA4500DAA30B26C681B7981AEAE3F8E">
|
||
<heading id="9C6A72AC15B308B493628BE517442402" reason="title">Etymology.</heading>
|
||
</paragraph>
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<paragraph id="B8273F15311D72EABB63BE1E6F7DBF32">
|
||
The specific designation “
|
||
<emphasis id="93E13D1C7519E54CE479137484F6367E" italics="true">quinquelirata</emphasis>
|
||
” indicates the number of lirae on the whorl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENHBG" type="diagnosis">
|
||
<paragraph id="AE987EB84A8B3705356F33843903DBE1">
|
||
<heading id="6D0E753E5ACBCFCF8BAB6D54A3AA60B2" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="F7803712B12AE76C26F95FF75BB3358C">Shell minute, ovately conical, orangish to brownish, protoconch round to slightly ellipsoid and wrinkled sculpture. Body whorl have about 5–6 prominent carinae, and between carinae have weak growth lines. Aperture semi-rounded. Umbilicus closed. Operculum corneous with apophysis, paucispiral, opaque white, and semi-rounded.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESHBG" type="description">
|
||
<paragraph id="B9921E251678D9EC30DFD491C71459E3">
|
||
<heading id="9647EE05224CF33DB265FC30B7F2B11B" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="85F3B43246A561A5F92E8059D68AC52D">
|
||
Shell minute, dextral, orangish to brownish, ovately conical with 4.25–4.5 whorls (Fig.
|
||
<figureCitation id="A6E0BB74287192D75CB125F98A3D207E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 A</figureCitation>
|
||
). Protoconch orange, about one whorl, round to slightly ellipsoid, covered with wrinkled sculpture (Fig.
|
||
<figureCitation id="BE6D64FBAC5AC9B6CB9BE9270C076FC0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 B, C</figureCitation>
|
||
). Teleoconch orange, 3.25–3.5 whorls. Body whorl peripherally rounded, sculptured with 5–6 prominent carinae, weak growth lines between carinae, intersected by weak oblique growth lines. Basal part of body whorl with approximately 9–10 densely spaced spiral cords (Figs
|
||
<figureCitation id="DA93CED9DD64EB139F863B32951F2969" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 A</figureCitation>
|
||
,
|
||
<figureCitation id="958C876414A6833CDAA8BBF64FF5E47A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 A</figureCitation>
|
||
). Suture incised. Aperture semi-rounded. Peristome thin, sharp, unexpanded or thickened, not reflected. Umbilicus closed (Figs
|
||
<figureCitation id="6B23C8E310DF6D9865FD3CE1BD88CE82" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 A</figureCitation>
|
||
,
|
||
<figureCitation id="8AFBE5E5A8D1E79771B2A51DAF94B129" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 A</figureCitation>
|
||
). Operculum paucispiral, nucleus submarginal, corneous, opaque white, semi-rounded, thin, smooth outside surface with apophysis, inside surface with an arched peg arising from base (Fig.
|
||
<figureCitation id="9A166FEBD579B1F82B41F63DA7322EE9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUIBG" type="Radula">
|
||
<paragraph id="FAF1BC16C735EDF4DFF6E208D64EDDE0">
|
||
<heading id="896372C9B9CEE6892907B9E18772A6D0" reason="title">Radula.</heading>
|
||
</paragraph>
|
||
<paragraph id="5B11515B30157222F41DD5BB1C4D17C9">
|
||
Ribbon-like, slender, longer, and delicate towards the outside, rhipidoglossate (Fig.
|
||
<figureCitation id="48BED4DBCA836703A79C35B40075CCB4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 A – D</figureCitation>
|
||
). Central tooth five small teeth, trapezoid shape. Lateral teeth five teeth, claw-like, with 9–13 alternately long and short denticles decreasing in size from the first lateral tooth toward the 5
|
||
<superScript id="D831C30222381E6C0159773EA849A84C">th</superScript>
|
||
lateral tooth. Marginal teeth two teeth, claw-like, with 12–14 equal-sized denticles. Radula formula 2: 5: 1 + 1 + 1 + 1 + 1 + 1: 5: 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAJBG" type="Differential diagnosis">
|
||
<paragraph id="938652E1CD0D892B070D6B4725B51792">
|
||
<heading id="F578D0FCF8893EDBE42B1EC7FA3AD767" reason="title">Differential diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="AAC3B7E74CD5C07283B6EAAD0D7912C8">
|
||
<taxonomicName id="9464BB0C54EA2C010DD55B9FF3B541A8" authorityName="Klongklaew & Poeaim & Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="quinquelirata" status="sp. nov.">
|
||
<emphasis id="58823DFAE24DF8D22AC7FA8629D1BDAA" italics="true">Georissa quinquelirata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="045AB390C5741869FFE2D85F14ADA060" rank="species">sp. nov.</taxonomicNameLabel>
|
||
resembles
|
||
<taxonomicName id="306FA8B256694D06B9B286F5E4FB3C55" authority="Godwin-Austen, 1889" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="hungerfordi">
|
||
<emphasis id="22BBDF51D967D4AD4F446B4F206BFA4C" italics="true">G. hungerfordi</emphasis>
|
||
<bibRefCitation id="735237A5CFC37D7C0A83020F6297A719" author="Godwin-Austen HH" issue="3" journalOrPublisher="Proceedings of the Zoological Society of London" pagination="332-335" refId="B11" refString="Godwin-Austen HH (1889) On a collection of land-shells made in Borneo by Mr. A. Everett with supposed new species. Part I. Cyclostomacae. Proceedings of the Zoological Society of London 57 (3): 332 - 335." title="On a collection of land-shells made in Borneo by Mr. A. Everett with supposed new species. Part I. Cyclostomacae." volume="57" year="1889">Godwin-Austen, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="A8B4C1910C19B54A26B3E4F9BD42933E" country="Malaysia" name="Sabah">Sabah</collectingRegion>
|
||
, Borneo,
|
||
<collectingCountry id="8F0194FB922D9ADD5F5C245616A7BD89" name="Malaysia">Malaysia</collectingCountry>
|
||
,
|
||
<taxonomicName id="543310104B5B22473717B73B5AD84BEE" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="hungerfordi">
|
||
<emphasis id="88E2BB9410886FB5711CA49075F5F5AA" italics="true">Georissa hungerfordi</emphasis>
|
||
</taxonomicName>
|
||
differs in having a less convex body whorl, a smooth sculpture, and more spiral lirae (7–10 lirae instead of 5–6 carinae). This new species is also like
|
||
<taxonomicName id="DC80AC9BCD74CDC828B5B0AFD90086A1" authority="Stolixzka, 1871" authorityName="Stolixzka" authorityYear="1871" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="liratula">
|
||
<emphasis id="4EA3DB38129B49B137DFA827F482EDE0" italics="true">G. liratula</emphasis>
|
||
Stolixzka, 1871
|
||
</taxonomicName>
|
||
from Domotha, Moulmein, but differs in having a more slender shell with fewer spiral lirae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |