277 lines
38 KiB
XML
277 lines
38 KiB
XML
<document id="56B514146D02ACC57F15066301F114AB" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFF94FF9FFF3DF989FCF7E3EE" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana minneriya Huber 2019, sp. n." docType="treatment" docVersion="14" lastPageNumber="5" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="2" updateTime="1698716853299" updateUser="ExternalLinkService">
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<mods:titleInfo id="1B0534B6037A728D0E0B1A4862A89051">
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<mods:title id="8AEFBD5A0A46B12C59E15FFC7770F484">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F2322DAA199409691A2D073566F842A1" type="personal">
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<mods:roleTerm id="13C80F9242F8D5ECFD709E16EEFE8B97">Author</mods:roleTerm>
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<mods:namePart id="C03A75094FE4700FB5EBF695C973028B">Huber, Bernhard A.</mods:namePart>
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<mods:typeOfResource id="F748C8C42EB7F7BA7351C36CCA37D8D0">text</mods:typeOfResource>
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<mods:title id="1E2EBAD5DED6FAC076E6A83788925640">Zootaxa</mods:title>
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<mods:part id="39C8361745DB7CCC92B8269A28E1368B">
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<mods:date id="B17F5E153D25B72F9663165C290D6B97">2019</mods:date>
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<mods:number id="F04EEAE5F07EC0F92B90A5D4C844C4F4">2019-01-23</mods:number>
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<mods:number id="4C04A107EE9AF7078B8B35600D29BD84">4550</mods:number>
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<mods:classification id="2FC202797B4FDA1D8DABA85B592D12E3">journal article</mods:classification>
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<mods:identifier id="6853135FCAF90A26B93D0F9FA59926DC" type="DOI">10.11646/zootaxa.4550.1.1</mods:identifier>
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<treatment id="CA3B104CFF94FF9FFF3DF989FCF7E3EE" ID-DOI="http://doi.org/10.5281/zenodo.4581568" ID-GBIF-Taxon="154534123" ID-Zenodo-Dep="4581568" LSID="urn:lsid:plazi:treatment:CA3B104CFF94FF9FFF3DF989FCF7E3EE" httpUri="http://treatment.plazi.org/id/CA3B104CFF94FF9FFF3DF989FCF7E3EE" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
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<subSubSection id="0A88F2D1FF94FF9AFF3DF989FE7BE728" box="[151,461,1576,1602]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="422DA15AFF94FF9AFF3DF989FE7BE728" blockId="1.[151,461,1576,1635]" box="[151,461,1576,1602]" pageId="1" pageNumber="2">
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<heading id="19651636FF94FF9AFF3DF989FE7BE728" bold="true" box="[151,461,1576,1602]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="70E67D48FF94FF9AFF3DF989FE7BE728" bold="true" box="[151,461,1576,1602]" pageId="1" pageNumber="2">
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<taxonomicName id="8592DAD9FF94FF9AFF3DF989FE32E728" ID-CoL="68H98" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[151,388,1576,1602]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="minneriya" status="sp. nov.">
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<emphasis id="70E67D48FF94FF9AFF3DF989FE32E728" bold="true" box="[151,388,1576,1602]" italics="true" pageId="1" pageNumber="2">Belisana minneriya</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="6BD5C033FF94FF9AFE21F989FE7BE728" box="[395,461,1576,1602]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF94FF9AFF3DF9EBFE72E709" box="[151,452,1610,1635]" pageId="1" pageNumber="2" type="description">
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||
<paragraph id="422DA15AFF94FF9AFF3DF9EBFE72E709" blockId="1.[151,461,1576,1635]" box="[151,452,1610,1635]" pageId="1" pageNumber="2">
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||
<figureCitation id="DAA9BDDFFF94FF9AFF3DF9EBFEABE709" box="[151,285,1610,1635]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[151,1436,187,1603]" captionTargetId="figure@2.[151,1436,187,1603]" captionTargetPageId="2" captionText="FIGURES 1–9. Belisana, live specimens. 1–3. B. minneriya sp. n., males and female with egg-sac from Minneriya. 4–5. B. keyti Huber, 2005, male and female with egg-sac from Hakgala. 6–7. B. gowindahela sp. n., male from Ethagala, female with egg-sac from Dematagala. 8–9. B. ratnapura Huber, 2005, male from Mapalena Ella, female with egg-sac from Viharekele." httpUri="https://zenodo.org/record/2625032/files/figure.png" pageId="1" pageNumber="2">Figures 1–3</figureCitation>
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,
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<figureCitation id="DAA9BDDFFF94FF9AFE80F9EBFEC4E709" box="[298,370,1610,1635]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="1" pageNumber="2">10–16</figureCitation>
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,
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<figureCitation id="DAA9BDDFFF94FF9AFED4F9EBFE72E709" box="[382,452,1610,1635]" captionStart="FIGURES 36–41" captionStartId="10.[151,264,1936,1959]" captionTargetBox="[164,1427,375,1921]" captionTargetId="figure@10.[164,1427,375,1923]" captionTargetPageId="10" captionText="FIGURES 36–41. Cleared female genitalia in ventral and dorsal views. 36–37. Belisana minneriya sp. n. (ZFMK Ar 20000). 38–39. Belisana gowindahela sp. n. (ZFMK Ar 20007). 40–41. B. ratnapura Huber, 2005 (ZFMK Ar 20015). Abbreviations: po, pocket(s); pp, pore plate. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/2625042/files/figure.png" pageId="1" pageNumber="2">36–37</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF94FF9AFF3DF933FC17E7C2" box="[151,929,1682,1704]" pageId="1" pageNumber="2" type="reference_group">
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||
<paragraph id="422DA15AFF94FF9AFF3DF933FC17E7C2" blockId="1.[151,929,1682,1704]" box="[151,929,1682,1704]" pageId="1" pageNumber="2">
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<taxonomicName id="8592DAD9FF94FF9AFF3DF933FF46E7C2" ID-CoL="398Y" box="[151,240,1682,1704]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="70E67D48FF94FF9AFF3DF933FF46E7C2" box="[151,240,1682,1704]" italics="true" pageId="1" pageNumber="2">Belisana</emphasis>
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</taxonomicName>
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SL37:
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<bibRefCitation id="2603DCABFF94FF9AFE90F933FE58E7C2" author="Eberle, J. & Dimitrov, D. & Valdez-Mondragon, A. & Huber, B. A." box="[314,494,1682,1704]" pageId="1" pageNumber="2" pagination="141" refId="ref26630" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. & Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
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Eberle
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<emphasis id="70E67D48FF94FF9AFE28F932FE00E7C2" box="[386,438,1682,1704]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
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2018
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</bibRefCitation>
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(molecular data).
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<bibRefCitation id="2603DCABFF94FF9AFD0DF933FCEAE7C2" author="Huber, B. A. & Eberle, J. & Dimitrov, D." box="[679,860,1682,1704]" pageId="1" pageNumber="2" pagination="51 - 101" refId="ref27545" refString="Huber, B. A., Eberle, J. & Dimitrov, D. (2018) The phylogeny of pholcid spiders (Araneae, Pholcidae): a critical evaluation of relationships suggested by molecular data. ZooKeys, 789, 51 - 101. https: // doi. org / 10.3897 / zookeys. 789.22781" type="journal article" year="2018">
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Huber
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<emphasis id="70E67D48FF94FF9AFD47F932FC96E7C2" box="[749,800,1682,1704]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
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2018
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||
</bibRefCitation>
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||
: fig. 8.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF94FF9AFF3DF974FE14E614" pageId="1" pageNumber="2" type="diagnosis">
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||
<paragraph id="422DA15AFF94FF9AFF3DF974FE14E614" blockId="1.[151,1437,1749,2026]" pageId="1" pageNumber="2">
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||
<emphasis id="70E67D48FF94FF9AFF3DF974FEBCE784" bold="true" box="[151,266,1749,1774]" pageId="1" pageNumber="2">Diagnosis</emphasis>
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. Easily distinguished from Sri Lankan congeners by being the only known litter-dwelling species (i.e., brown color, very short legs: tibia 1 ~1.5); also by male chelicerae with long apophyses (
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<figureCitation id="DAA9BDDFFF94FF9AFB26F958FB56E678" box="[1164,1248,1785,1810]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
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; similar in some species from outside
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<collectingCountry id="3A85E1CAFF94FF9AFE2CF8BCFE4EE65C" box="[390,504,1821,1846]" name="Sri Lanka" pageId="1" pageNumber="2">Sri Lanka</collectingCountry>
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, e.g.
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<taxonomicName id="8592DAD9FF94FF9AFD9EF8BFFCE1E65C" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[564,855,1821,1846]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dodabetta">
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<emphasis id="70E67D48FF94FF9AFD9EF8BFFD72E65C" box="[564,708,1821,1846]" italics="true" pageId="1" pageNumber="2">B. dodabetta</emphasis>
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Huber, 2005
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</taxonomicName>
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from southern
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<collectingCountry id="3A85E1CAFF94FF9AFBACF8BCFBF5E65C" box="[1030,1091,1821,1846]" name="India" pageId="1" pageNumber="2">India</collectingCountry>
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; see Huber 2005: fig. 599), by procursus with distinctive dorsal sclerite (
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<figureCitation id="DAA9BDDFFF94FF9AFDC0F8E0FD5AE630" box="[618,748,1857,1882]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="1" pageNumber="2">Figs 13–14</figureCitation>
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), and by short and wide epigynal plate with distinctive lateral pair of humps (
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<figureCitation id="DAA9BDDFFF94FF9AFEE8F8C4FE23E614" box="[322,405,1893,1918]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="1" pageNumber="2">Fig. 15</figureCitation>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF94FF9AFF6DF828FB37E6C8" box="[199,1153,1929,1954]" pageId="1" pageNumber="2" type="etymology">
|
||
<paragraph id="422DA15AFF94FF9AFF6DF828FB37E6C8" blockId="1.[151,1437,1749,2026]" box="[199,1153,1929,1954]" pageId="1" pageNumber="2">
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||
<emphasis id="70E67D48FF94FF9AFF6DF828FEF0E6C8" bold="true" box="[199,326,1929,1954]" pageId="1" pageNumber="2">Etymology</emphasis>
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||
. The specific name is derived from the
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||
<typeStatus id="9D291FF8FF94FF9AFCAEF82BFC82E6C8" box="[772,820,1930,1954]" pageId="1" pageNumber="2">type</typeStatus>
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||
locality (noun in apposition).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF94FF9FFF6DF80CFBC4E1BF" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="materials_examined">
|
||
<paragraph id="422DA15AFF94FF9AFF6DF80CFC8CE680" blockId="1.[151,1437,1749,2026]" pageId="1" pageNumber="2">
|
||
<emphasis id="70E67D48FF94FF9AFF6DF80CFEC4E6AC" bold="true" box="[199,370,1965,1990]" pageId="1" pageNumber="2">Type material</emphasis>
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||
.
|
||
<materialsCitation id="F2FAAB07FF94FF9AFE2EF80FFC80E680" ID-GBIF-Occurrence="2234226292" collectingDate="2017-03-10" collectionCode="ZFMK" collectorName="B. A. Huber" country="Sri Lanka" elevation="150" latitude="8.047" location="Minneriya Forest" longLatPrecision="77" longitude="80.832" municipality="Ar" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="North Central" typeStatus="holotype">
|
||
<collectingCountry id="3A85E1CAFF94FF9AFE2EF80FFD95E6AC" box="[388,547,1966,1990]" name="Sri Lanka" pageId="1" pageNumber="2">SRI LANKA</collectingCountry>
|
||
:
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||
<specimenCount id="54946AD3FF94FF9AFD9BF80FFDF0E6AF" box="[561,582,1966,1989]" pageId="1" pageNumber="2" type="male">♂</specimenCount>
|
||
<typeStatus id="9D291FF8FF94FF9AFDF8F80CFD01E6AC" box="[594,695,1965,1990]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
|
||
,
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||
<collectionCode id="2483399FFF94FF9AFD6CF80FFCA2E6AF" box="[710,788,1966,1989]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="2" type="Museum">ZFMK</collectionCode>
|
||
(
|
||
<collectingMunicipality id="A2493B20FF94FF9AFC8DF80FFCF0E6AC" box="[807,838,1966,1990]" pageId="1" pageNumber="2">Ar</collectingMunicipality>
|
||
19999),
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||
<collectingRegion id="80566FB8FF94FF9AFC1EF80CFB73E6AC" box="[948,1221,1965,1990]" country="Sri Lanka" name="North Central" pageId="1" pageNumber="2">North Central Province</collectingRegion>
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||
,
|
||
<location id="474DF781FF94FF9AFB7CF80CFA2AE6AF" LSID="urn:lsid:plazi:treatment:CA3B104CFF94FF9FFF3DF989FCF7E3EE:474DF781FF94FF9AFB7CF80CFA2AE6AF" box="[1238,1436,1965,1990]" country="Sri Lanka" latitude="8.047" longLatPrecision="77" longitude="80.832" municipality="Ar" name="Minneriya Forest" pageId="1" pageNumber="2" stateProvince="North Central">Minneriya Forest</location>
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||
(
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||
<geoCoordinate id="27A6C79DFF94FF9AFF35F873FEB7E680" box="[159,257,2002,2026]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="55" value="8.047">8.047°N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="27A6C79DFF94FF9AFEA6F873FECCE680" box="[268,378,2002,2026]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="55" value="80.832">80.832°E</geoCoordinate>
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||
),
|
||
<elevation id="C9BF4669FF94FF9AFE27F870FDA7E680" box="[397,529,2001,2026]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="2" unit="m" value="150.0">
|
||
<quantity id="856A0CBFFF94FF9AFE27F870FE62E683" box="[397,468,2001,2026]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="2" unit="m" value="150.0">150 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="362C879AFF94FF9AFDB7F870FD20E680" box="[541,662,2001,2026]" pageId="1" pageNumber="2" value="2017-03-10">
|
||
<collectingDate id="26687E72FF94FF9AFDB7F870FD20E680" box="[541,662,2001,2026]" pageId="1" pageNumber="2" value="2017-03-10">10.iii.2017</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="EF67C48CFF94FF9AFD0DF873FC98E680" box="[679,814,2001,2026]" pageId="1" pageNumber="2">B.A. Huber</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="16EDF1D2FF97FF99FF3DF9F5FACCE7C3" httpUri="https://zenodo.org/record/2625032/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,264,1620,1643]" targetBox="[151,1436,187,1603]" targetPageId="2">
|
||
<paragraph id="422DA15AFF97FF99FF3DF9F5FACCE7C3" blockId="2.[151,1436,1620,1705]" pageId="2" pageNumber="3">
|
||
<emphasis id="70E67D48FF97FF99FF3DF9F5FE83E700" bold="true" box="[151,309,1620,1643]" pageId="2" pageNumber="3">FIGURES 1–9</emphasis>
|
||
.
|
||
<taxonomicName id="8592DAD9FF97FF99FEEEF9F4FE2BE701" box="[324,413,1621,1643]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E67D48FF97FF99FEEEF9F4FE2BE701" box="[324,413,1621,1643]" italics="true" pageId="2" pageNumber="3">Belisana</emphasis>
|
||
</taxonomicName>
|
||
, live specimens. 1–3.
|
||
<taxonomicName id="8592DAD9FF97FF99FD2EF9F7FCBFE701" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[644,777,1622,1643]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="minneriya" status="sp. nov.">
|
||
<emphasis id="70E67D48FF97FF99FD2EF9F7FCBFE701" box="[644,777,1622,1643]" italics="true" pageId="2" pageNumber="3">B. minneriya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="70E67D48FF97FF99FCBBF9F7FCFAE701" bold="true" box="[785,844,1622,1643]" pageId="2" pageNumber="3">
|
||
<taxonomicNameLabel id="6BD5C033FF97FF99FCBBF9F7FCFAE701" box="[785,844,1622,1643]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, males and female with egg-sac from Minneriya. 4–5.
|
||
<taxonomicName id="8592DAD9FF97FF99FA2CF9F7FEF8E7E0" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="keyti">
|
||
<emphasis id="70E67D48FF97FF99FA2CF9F7FF71E7E0" italics="true" pageId="2" pageNumber="3">B. keyti</emphasis>
|
||
Huber, 2005
|
||
</taxonomicName>
|
||
, male and female with egg-sac from Hakgala. 6–7.
|
||
<taxonomicName id="8592DAD9FF97FF99FCF5F9D4FC48E7E0" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[863,1022,1652,1674]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gowindahela" status="sp. nov.">
|
||
<emphasis id="70E67D48FF97FF99FCF5F9D4FC48E7E0" box="[863,1022,1652,1674]" italics="true" pageId="2" pageNumber="3">B. gowindahela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="70E67D48FF97FF99FBAFF9D5FB89E7E3" bold="true" box="[1029,1087,1652,1673]" pageId="2" pageNumber="3">
|
||
<taxonomicNameLabel id="6BD5C033FF97FF99FBAFF9D5FB89E7E3" box="[1029,1087,1652,1673]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male from Ethagala, female with egg-sac from Dematagala. 8–9.
|
||
<taxonomicName id="8592DAD9FF97FF99FE70F935FD55E7C3" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[474,739,1683,1705]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ratnapura">
|
||
<emphasis id="70E67D48FF97FF99FE70F935FDEBE7C3" box="[474,605,1684,1705]" italics="true" pageId="2" pageNumber="3">B. ratnapura</emphasis>
|
||
Huber, 2005
|
||
</taxonomicName>
|
||
, male from Mapalena Ella, female with egg-sac from Viharekele.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="422DA15AFF97FF99FF6DF96FFE96E689" blockId="2.[151,1437,1742,2019]" pageId="2" pageNumber="3">
|
||
<emphasis id="70E67D48FF97FF99FF6DF96FFDB7E78D" bold="true" box="[199,513,1742,1767]" pageId="2" pageNumber="3">Other material examined</emphasis>
|
||
.
|
||
<materialsCitation id="F2FAAB07FF97FF99FDBFF96EFBF1E78D" ID-GBIF-Occurrence="2234226208" box="[533,1095,1742,1767]" collectionCode="ZFMK" country="Sri Lanka" location="SRI LANKA" pageId="2" pageNumber="3" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<collectingCountry id="3A85E1CAFF97FF99FDBFF96EFD03E78D" box="[533,693,1743,1767]" name="Sri Lanka" pageId="2" pageNumber="3">SRI LANKA</collectingCountry>
|
||
:
|
||
<specimenCount id="54946AD3FF97FF99FD6FF96EFD5EE78D" box="[709,744,1743,1767]" pageId="2" pageNumber="3" type="male">1♂</specimenCount>
|
||
<specimenCount id="54946AD3FF97FF99FD5CF96EFCAEE78D" box="[758,792,1743,1767]" pageId="2" pageNumber="3" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="2483399FFF97FF99FC87F971FCCDE78D" box="[813,891,1744,1767]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="3" type="Museum">ZFMK</collectionCode>
|
||
(Ar 20000), and
|
||
</materialsCitation>
|
||
<materialsCitation id="F2FAAB07FF97FF99FBFFF96EFEAAE689" ID-GBIF-Occurrence="2234226153" collectingDate="2017-03-10" collectionCode="ZFMK, COLOR, BODY" collectorName="B. A. Huber" country="Sri Lanka" elevation="150" latitude="8.047" location="Minneriya Forest" longLatPrecision="77" longitude="80.832" municipality="Ar" pageId="2" pageNumber="3" specimenCode="SL123" specimenCount="1" specimenCount-female="1" stateProvince="North Central" typeStatus="holotype">
|
||
<specimenCount id="54946AD3FF97FF99FBFFF96EFBCEE78D" box="[1109,1144,1743,1767]" pageId="2" pageNumber="3" type="female">1♀</specimenCount>
|
||
in pure ethanol,
|
||
<collectionCode id="2483399FFF97FF99FAE4F971FA2AE78D" box="[1358,1436,1744,1767]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="3" type="Museum">ZFMK</collectionCode>
|
||
(
|
||
<specimenCode id="12340921FF97FF99FF35F952FF58E661" box="[159,238,1779,1803]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="3" type="Museum">SL123</specimenCode>
|
||
), same data as holotype.
|
||
<emphasis id="70E67D48FF97FF99FF6DF8B7FE28E645" bold="true" box="[199,414,1814,1839]" pageId="2" pageNumber="3">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="9D291FF8FF97FF99FE04F8B7FDA0E645" box="[430,534,1814,1839]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
). MEASUREMENTS. Total body length 1.0, carapace width 0.47. Distance PME- PME 50 µm, diameter PME 60 µm, distance PME-ALE 10 µm; AME absent. Sternum width/length: 0.34/0.28. Leg 1: 5.75 (1.50 + 0.20 + 1.55 + 1.75 + 0.75), tibia 2: 0.95, tibia 3: 0.65, tibia 4: 1.00; tibia 1 L/d: 36.
|
||
<collectionCode id="2483399FFF97FF99FF6DF822FEA0E6F3" box="[199,278,1923,1947]" pageId="2" pageNumber="3">COLOR</collectionCode>
|
||
(in ethanol). Prosoma and legs ochre-yellow, abdomen ochre-gray; body and legs without dark marks.
|
||
<collectionCode id="2483399FFF97FF99FF6DF809FEBEE6D4" box="[199,264,1960,1983]" pageId="2" pageNumber="3">BODY</collectionCode>
|
||
. Habitus as in
|
||
<figureCitation id="DAA9BDDFFF97FF99FE04F807FDA4E6D5" box="[430,530,1958,1983]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[151,1436,187,1603]" captionTargetId="figure@2.[151,1436,187,1603]" captionTargetPageId="2" captionText="FIGURES 1–9. Belisana, live specimens. 1–3. B. minneriya sp. n., males and female with egg-sac from Minneriya. 4–5. B. keyti Huber, 2005, male and female with egg-sac from Hakgala. 6–7. B. gowindahela sp. n., male from Ethagala, female with egg-sac from Dematagala. 8–9. B. ratnapura Huber, 2005, male from Mapalena Ella, female with egg-sac from Viharekele." httpUri="https://zenodo.org/record/2625032/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
|
||
; ocular area barely raised; carapace without median furrow; clypeus and sternum unmodified
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="16EDF1D2FF96FF98FF3DF8FEFE0DE69A" httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,264,1887,1910]" targetBox="[151,1436,205,1836]" targetPageId="3">
|
||
<paragraph id="422DA15AFF96FF98FF3DF8FEFE0DE69A" blockId="3.[151,1436,1887,2032]" pageId="3" pageNumber="4">
|
||
<emphasis id="70E67D48FF96FF98FF3DF8FEFEF8E61E" bold="true" box="[151,334,1887,1910]" pageId="3" pageNumber="4">FIGURES 10–16</emphasis>
|
||
.
|
||
<taxonomicName id="8592DAD9FF96FF98FEF6F8FEFD95E61C" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[348,547,1887,1910]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="minneriya" status="sp. nov.">
|
||
<emphasis id="70E67D48FF96FF98FEF6F8FEFD95E61C" box="[348,547,1887,1910]" italics="true" pageId="3" pageNumber="4">Belisana minneriya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="70E67D48FF96FF98FD80F8C1FDD5E61F" bold="true" box="[554,611,1888,1909]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="6BD5C033FF96FF98FD80F8C1FDD5E61F" box="[554,611,1888,1909]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="422DA15AFF91FF9FFF6DFF39FBC4E1BF" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="5">
|
||
CHELICERAE. As in
|
||
<figureCitation id="DAA9BDDFFF91FF9FFE18FF36FDB1E1DA" box="[434,519,151,176]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
|
||
, with pair of small apophyses proximally laterally and pair of long apophyses directed toward lateral and slightly downward; distance between tips of apophyses 0.35.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A88F2D1FF91FF9FFF6DFF41FAA3E30A" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="422DA15AFF91FF9FFF6DFF41FDCCE00F" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="5">
|
||
PALPS. As in
|
||
<figureCitation id="DAA9BDDFFF91FF9FFEC9FF7EFE55E192" box="[355,483,223,248]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="4" pageNumber="5">Figs 10–11</figureCitation>
|
||
; coxa unmodified; trochanter with short and wide rounded process ventrally; femur without process; procursus with transparent retrolateral flap, with distinctive dorsal sclerite and complex membranous elements distally (
|
||
<figureCitation id="DAA9BDDFFF91FF9FFDADFE86FD38E02A" box="[519,654,295,320]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="4" pageNumber="5">Figs 13–14</figureCitation>
|
||
); bulb with long and slender apophysis curved distally and simple embolus lying parallel to bulbal apophysis.
|
||
</paragraph>
|
||
<paragraph id="422DA15AFF91FF9FFF6DFED1FB32E0C7" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="5">LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 22%; prolateral trichobothrium apparently absent on tibia 1; tarsus 1 with ~15 pseudosegments.</paragraph>
|
||
<paragraph id="422DA15AFF91FF9FFF6DFE16FD76E0BA" blockId="4.[151,1437,151,645]" box="[199,704,439,464]" pageId="4" pageNumber="5">
|
||
<emphasis id="70E67D48FF91FF9FFF6DFE16FEB2E0BA" bold="true" box="[199,260,439,464]" pageId="4" pageNumber="5">Male</emphasis>
|
||
(variation). Tibia
|
||
<quantity id="856A0CBFFF91FF9FFE7AFE19FE4DE0BA" box="[464,507,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="5" unit="in" value="1.0">1 in</quantity>
|
||
other male: 1.40.
|
||
</paragraph>
|
||
<paragraph id="422DA15AFF91FF9FFF6DFE7DFA3EE357" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="5">
|
||
<emphasis id="70E67D48FF91FF9FFF6DFE7DFEA8E09F" bold="true" box="[199,286,476,501]" pageId="4" pageNumber="5">Female</emphasis>
|
||
. In general similar to male (
|
||
<figureCitation id="DAA9BDDFFF91FF9FFDCEFE7DFD1CE09E" box="[612,682,476,501]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1620,1643]" captionTargetBox="[151,1436,187,1603]" captionTargetId="figure@2.[151,1436,187,1603]" captionTargetPageId="2" captionText="FIGURES 1–9. Belisana, live specimens. 1–3. B. minneriya sp. n., males and female with egg-sac from Minneriya. 4–5. B. keyti Huber, 2005, male and female with egg-sac from Hakgala. 6–7. B. gowindahela sp. n., male from Ethagala, female with egg-sac from Dematagala. 8–9. B. ratnapura Huber, 2005, male from Mapalena Ella, female with egg-sac from Viharekele." httpUri="https://zenodo.org/record/2625032/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
|
||
); tibia 1: 1.5 (missing in second female). Epigynum as in
|
||
<figureCitation id="DAA9BDDFFF91FF9FFAEEFE7DFA21E09F" box="[1348,1431,476,501]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
|
||
, short but wide plate barely darker than rest of abdomen, with distinctive pair of lateral humps carrying pair of pockets, distance between pockets 315 µm. Internal genitalia as in
|
||
<figureCitation id="DAA9BDDFFF91FF9FFC29FD85FC6FE357" box="[899,985,548,573]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1887,1910]" captionTargetBox="[151,1436,205,1836]" captionTargetId="figure@3.[151,1436,205,1836]" captionTargetPageId="3" captionText="FIGURES 10–16. Belisana minneriya sp. n. (ZFMK Ar 20000). 10–11. Left male palp, prolateral and retrolateral views. 12. Male chelicerae, frontal view. 13–14. Left procursus, prolateral and retrolateral views. 15. Epigynum, ventral view. 16. Cleared female genitalia, dorsal view. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; ds, dorsal sclerite; e, embolus; f, femur; p, procursus; pbs, proximal bulbal sclerite; rf, retrolateral flap; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.2 mm (10– 12, 15–16), 0.1 mm (13–14)." httpUri="https://zenodo.org/record/2625034/files/figure.png" pageId="4" pageNumber="5">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="DAA9BDDFFF91FF9FFC4FFD85FB9BE357" box="[997,1069,548,573]" captionStart="FIGURES 36–41" captionStartId="10.[151,264,1936,1959]" captionTargetBox="[164,1427,375,1921]" captionTargetId="figure@10.[164,1427,375,1923]" captionTargetPageId="10" captionText="FIGURES 36–41. Cleared female genitalia in ventral and dorsal views. 36–37. Belisana minneriya sp. n. (ZFMK Ar 20000). 38–39. Belisana gowindahela sp. n. (ZFMK Ar 20007). 40–41. B. ratnapura Huber, 2005 (ZFMK Ar 20015). Abbreviations: po, pocket(s); pp, pore plate. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/2625042/files/figure.png" pageId="4" pageNumber="5">36–37</figureCitation>
|
||
, with pair of small pore plates.
|
||
</paragraph>
|
||
<paragraph id="422DA15AFF91FF9FFF6DFDE6FAA3E30A" blockId="4.[151,1437,151,645]" box="[199,1301,583,608]" pageId="4" pageNumber="5">
|
||
<emphasis id="70E67D48FF91FF9FFF6DFDE6FECBE30A" bold="true" box="[199,381,583,608]" pageId="4" pageNumber="5">Natural history</emphasis>
|
||
. This species was found in dry leaf litter among low vegetation outside the forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A88F2D1FF91FF9FFF6DFDCDFCF7E3EE" box="[199,833,620,645]" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="422DA15AFF91FF9FFF6DFDCDFCF7E3EE" blockId="4.[151,1437,151,645]" box="[199,833,620,645]" pageId="4" pageNumber="5">
|
||
<emphasis id="70E67D48FF91FF9FFF6DFDCDFEEFE3EF" bold="true" box="[199,345,620,645]" pageId="4" pageNumber="5">Distribution</emphasis>
|
||
. Known from
|
||
<typeStatus id="9D291FF8FF91FF9FFE54FDCCFD98E3EF" box="[510,558,621,645]" pageId="4" pageNumber="5">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="DAA9BDDFFF91FF9FFD7BFDCDFC83E3EE" box="[721,821,620,645]" captionStart="FIGURES 220–221" captionStartId="51.[151,264,919,942]" captionTargetBox="[151,1436,195,898]" captionTargetId="figure@51.[151,1436,195,898]" captionTargetPageId="51" captionText="FIGURES 220–221. Known distribution of Belisana in Sri Lanka." httpUri="https://zenodo.org/record/2625098/files/figure.png" pageId="4" pageNumber="5">Fig. 220</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |