341 lines
40 KiB
XML
341 lines
40 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6060906" ID-GBIF-Dataset="b19b3fd5-7380-49de-a706-5e8376384274" ID-GBIF-Taxon="126098808" ID-Zenodo-Dep="6060906" checkinTime="1483955123882" checkinUser="plazi" docAuthor="Jean- Paul Haenni & Dalton de Souza Amorim" docDate="2016" docId="725BEB09FFEFD27BFE0F10CC4209FEC2" docLanguage="en" docName="127-142_Haenni & Amorim.pdf" docOrigin="Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 89" docStyle="DocumentStyle{}" docTitle="Octaseps" docType="treatment" docVersion="8" lastPageId="7" lastPageNumber="134" masterDocId="8E629371FFEED27CFFD8122F4105FFE4" masterDocTitle="Two new Afrotropical genera of Scatopsidae (Diptera)" masterLastPageNumber="142" masterPageNumber="127" pageId="1" pageNumber="128" updateTime="1644757963397" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new Afrotropical genera of Scatopsidae (Diptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jean- Paul Haenni</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dalton de Souza Amorim</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Mitteilungen Der Schweizerischen Entomologischen Gesellschaft</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>89</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>127</mods:start>
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<mods:end>142</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.19262</mods:identifier>
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<mods:identifier type="GBIF-Dataset">b19b3fd5-7380-49de-a706-5e8376384274</mods:identifier>
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<mods:identifier type="Zenodo-Dep">192621</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6060906" ID-GBIF-Taxon="126098808" ID-Zenodo-Dep="6060906" LSID="urn:lsid:plazi:treatment:725BEB09FFEFD27BFE0F10CC4209FEC2" httpUri="http://treatment.plazi.org/id/725BEB09FFEFD27BFE0F10CC4209FEC2" lastPageId="7" lastPageNumber="134" pageId="1" pageNumber="128">
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<subSubSection box="[471,724,739,763]" pageId="1" pageNumber="128" type="nomenclature">
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<paragraph blockId="1.[471,724,739,799]" box="[471,724,739,763]" pageId="1" pageNumber="128">
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<heading box="[471,724,739,763]" centered="true" fontSize="10" level="1" pageId="1" pageNumber="128" reason="1">
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<emphasis bold="true" box="[471,631,739,763]" italics="true" pageId="1" pageNumber="128">
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<taxonomicName box="[471,569,739,763]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">Octaseps</taxonomicName>
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alata
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</emphasis>
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sp. nov.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[546,648,779,799]" pageId="1" pageNumber="128" type="description">
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<paragraph blockId="1.[471,724,739,799]" box="[546,648,779,799]" pageId="1" pageNumber="128">
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(
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<figureCitation box="[552,641,779,799]" captionStart-0="Figs 1 – 2" captionStart-1="Figs 3 – 4" captionStart-2="Figs 5 – 10" captionStartId-0="2.[140,178,1229,1248]" captionStartId-1="3.[155,193,1207,1226]" captionStartId-2="4.[140,178,1328,1347]" captionTargetBox-0="[140,1024,122,1206]" captionTargetBox-1="[155,1039,166,1171]" captionTargetBox-2="[145,1020,127,1285]" captionTargetId-0="figure@2.[140,1024,122,1206]" captionTargetId-1="figure@3.[155,1039,122,1171]" captionTargetId-2="figure@4.[140,1024,122,1288]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs 1 – 2. Octaseps alata sp. nov. ♂: — 1. Wing; — 2. Habitus (wings removed) (photographs Jessica Litman)." captionText-1="Figs 3 – 4. Octaseps alata sp. nov. — 3. Head and thorax, side view; — 4. Thorax and base of abdomen, subdorsal view showing the thickened rim between thorax and abdomen (marked) (photographs Jessica Litman)." captionText-2="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri-0="https://zenodo.org/record/235401/files/figure.png" httpUri-1="https://zenodo.org/record/235402/files/figure.png" httpUri-2="https://zenodo.org/record/235403/files/figure.png" pageId="1" pageNumber="128">Figs 1–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="128" type="nomenclature">
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<paragraph blockId="1.[155,1033,828,881]" pageId="1" pageNumber="128">
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<emphasis box="[155,298,828,852]" italics="true" pageId="1" pageNumber="128">Type locality</emphasis>
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.South Africa, Kwa-Zulu Natal: Karkloof Range, Geekie’s Farm, 29°16’ S30°21’ E.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="128" type="materials_examined">
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<paragraph blockId="1.[155,1040,907,1491]" pageId="1" pageNumber="128">
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<emphasis box="[215,423,909,933]" italics="true" pageId="1" pageNumber="128">Material examined</emphasis>
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.
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<materialsCitation ID-GBIF-Occurrence="1423853304" collectingDate="2000-10-11" collectingDateMax="2000-10-29" collectingDateMin="2000-10-11" collectingMethod="Malaise Trap" collectionCode="MHNN" collectorName="W. Barkemeyer" country="South Africa" latitude="-29.266666" location="Howick District" longLatPrecision="1228" longitude="30.35" municipality="Kwa Zulu Natal" pageId="1" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus box="[436,539,909,933]" pageId="1" pageNumber="128">Holotype</typeStatus>
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:
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<specimenCount box="[562,584,907,933]" pageId="1" pageNumber="128" type="male">♂</specimenCount>
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,labelled:
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<collectingCountry box="[703,826,909,933]" pageId="1" pageNumber="128">Süd-Afrika</collectingCountry>
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,
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<collectingMunicipality box="[842,1028,909,933]" pageId="1" pageNumber="128">Kwa Zulu Natal</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:725BEB09FFEFD27BFE0F10CC4209FEC2:FF2D0CC4FFEFD27DFF431185404DFC26" box="[155,328,938,962]" country="South Africa" latitude="-29.266666" longLatPrecision="1228" longitude="30.35" municipality="Kwa Zulu Natal" name="Howick District" pageId="1" pageNumber="128">Howick District</location>
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,
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<location LSID="urn:lsid:plazi:treatment:725BEB09FFEFD27BFE0F10CC4209FEC2:FF2D0CC4FFEFD27DFE89118543A3FC26" box="[337,678,938,963]" country="South Africa" latitude="-29.266666" longLatPrecision="1228" longitude="30.35" municipality="Kwa Zulu Natal" name="Karkloof Range, Geekie’s Farm" pageId="1" pageNumber="128">Karkloof Range, Geekie’s Farm</location>
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,
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<geoCoordinate box="[687,781,938,963]" direction="south" orientation="latitude" pageId="1" pageNumber="128" precision="925" value="-29.266666">29°16’ S</geoCoordinate>
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<geoCoordinate box="[780,876,939,963]" direction="east" orientation="longitude" pageId="1" pageNumber="128" precision="925" value="30.35">30°21’ E</geoCoordinate>
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,
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<collectingMethod box="[886,1029,938,962]" pageId="1" pageNumber="128">Malaise Trap</collectingMethod>
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,
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<collectingDate box="[155,309,968,992]" pageId="1" pageNumber="128" value="2000-10-11" valueMax="2000-10-29" valueMin="2000-10-11">
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<date box="[155,309,968,992]" pageId="1" pageNumber="128" value="2000-10-11" valueMax="2000-10-29" valueMin="2000-10-11">11–29.X.2000</date>
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</collectingDate>
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,
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<collectorName box="[327,501,968,992]" pageId="1" pageNumber="128">W. Barkemeyer</collectorName>
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leg. in coll.
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<collectionCode box="[659,744,968,991]" collectionName="Switzerland, Neuchatel, Museum d'Histoire Naturelle" pageId="1" pageNumber="128">MHNN</collectionCode>
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. Holotype dissected, slide mounted, in fairly good condition.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="128" type="diagnosis">
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<paragraph blockId="1.[155,1040,907,1491]" pageId="1" pageNumber="128">
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<emphasis box="[215,323,1027,1050]" italics="true" pageId="1" pageNumber="128">Diagnosis</emphasis>
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.Dorsal suture between thorax and abdomen forming asinuous Vshaped thickened rim laterally (
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<figureCitation box="[506,575,1056,1080]" captionStart="Figs 3 – 4" captionStartId="3.[155,193,1207,1226]" captionTargetBox="[155,1039,166,1171]" captionTargetId="figure@3.[155,1039,122,1171]" captionTargetPageId="3" captionText="Figs 3 – 4. Octaseps alata sp. nov. — 3. Head and thorax, side view; — 4. Thorax and base of abdomen, subdorsal view showing the thickened rim between thorax and abdomen (marked) (photographs Jessica Litman)." httpUri="https://zenodo.org/record/235402/files/figure.png" pageId="1" pageNumber="128">Fig. 4</figureCitation>
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). Male terminalia (
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<figureCitation box="[789,901,1056,1080]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="1" pageNumber="128">Figs 9–10</figureCitation>
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) with gonocoxites well-developed, parameres with teeth, aedeagus elongate, with an apically enlarged, flattened lobe.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="132" pageId="1" pageNumber="128" type="description">
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<paragraph blockId="1.[155,1040,907,1491]" lastBlockId="4.[140,1025,1326,1393]" lastPageId="4" lastPageNumber="131" pageId="1" pageNumber="128">
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<emphasis box="[215,341,1145,1168]" italics="true" pageId="1" pageNumber="128">Description</emphasis>
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.Male. 2.8 mm long (in alcohol). Head, body and legs black, hardly shining, densely pilose, especially on notum and on last abdominal segment. Labellum and palpi brownish, halters yellowish brown. Apical half of posterior tibia and all tarsi slightly lighter than remaining legs. Head (
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<figureCitation box="[776,843,1232,1256]" captionStart="Figs 3 – 4" captionStartId="3.[155,193,1207,1226]" captionTargetBox="[155,1039,166,1171]" captionTargetId="figure@3.[155,1039,122,1171]" captionTargetPageId="3" captionText="Figs 3 – 4. Octaseps alata sp. nov. — 3. Head and thorax, side view; — 4. Thorax and base of abdomen, subdorsal view showing the thickened rim between thorax and abdomen (marked) (photographs Jessica Litman)." httpUri="https://zenodo.org/record/235402/files/figure.png" pageId="1" pageNumber="128">Fig. 3</figureCitation>
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) as high as long. Supraantennal eye-bridge well developed; antennae longer than head, flagellum with 8flagellomeres, wider than long, each bearing asingle whorl of setae, last flagellomere longer than two preceding ones; frons setose between antennae and eyebridge; palpus adpressed, elongate, obtusely pointed apically, as long as labella. Thorax with notum longer than wide; spiracular sclerite somewhat elongate, nearly twice as long as high, acute anterodorsally (
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<figureCitation box="[638,706,1408,1432]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="1" pageNumber="128">Fig. 5</figureCitation>
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); an elongate row of about 20 supralar setae; scutellum rounded, with arow of lateral setae but devoid of pilosity on posterior third. Postnotal phragma elongate, as long as notum. Wing 2.4 mm long. Membrane very lightly brownish-tinged; anterior veins yellowish brown, posterior veins hyaline, R4+5 elongate, reaching Cat about 2/3 of wing length; base of A2/CuP somewhat inflated and strongly sclerotized, bearing ashort spinose microchetosity. Stem of halters with arow of setae and knob bearing asubbasal posterior patch of short setae near base (
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<figureCitation box="[524,593,1462,1486]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="2" pageNumber="129">Fig. 6</figureCitation>
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). Abdomen constricted at level of first segment after base, dorsal suture between thorax and abdomen V-shaped, forming laterally a sinuous, thickened elevated rim (
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<figureCitation box="[637,705,1345,1370]" captionStart="Figs 3 – 4" captionStartId="3.[155,193,1207,1226]" captionTargetBox="[155,1039,166,1171]" captionTargetId="figure@3.[155,1039,122,1171]" captionTargetPageId="3" captionText="Figs 3 – 4. Octaseps alata sp. nov. — 3. Head and thorax, side view; — 4. Thorax and base of abdomen, subdorsal view showing the thickened rim between thorax and abdomen (marked) (photographs Jessica Litman)." httpUri="https://zenodo.org/record/235402/files/figure.png" pageId="3" pageNumber="130">Fig. 4</figureCitation>
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); tergites 1–7 normally developed, tergite 2with apair of submedian lunules along anterior margin; sternites 2– 7present, normally sclerotized, 2–4 narrower; pregenital segment 7anteriorly with an entire, narrow, heavily sclerotized line of sclerotization; aweakly sclerotized area also present around spiracle 7, on pleural membranes joining tergite and sternite; tergite 7(
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<figureCitation box="[253,320,1492,1516]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="3" pageNumber="130">Fig. 7</figureCitation>
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) elongate, roof-shaped, with rounded posterior margin; sternite 7 Abbreviations: ae – aedeagus; gc – gonocoxite; ep – epandrium; pa – paramere.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/235401/files/figure.png" pageId="2" pageNumber="129" targetBox="[140,1024,122,1206]" targetPageId="2">
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<paragraph blockId="2.[140,1025,1228,1271]" pageId="2" pageNumber="129">
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Figs 1–2.
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<emphasis box="[229,360,1229,1248]" italics="true" pageId="2" pageNumber="129">
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<taxonomicName box="[229,309,1229,1248]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">Octaseps</taxonomicName>
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alata
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</emphasis>
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sp. nov. ♂: — 1. Wing; — 2. Habitus (wings removed) (photographs Jessica Litman).
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</paragraph>
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</caption>
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<caption httpUri="https://zenodo.org/record/235402/files/figure.png" pageId="3" pageNumber="130" targetBox="[155,1039,166,1171]" targetPageId="3">
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<paragraph blockId="3.[155,1039,1207,1272]" pageId="3" pageNumber="130">
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Figs 3–4.
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<emphasis box="[244,375,1207,1227]" italics="true" pageId="3" pageNumber="130">
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<taxonomicName box="[244,324,1208,1227]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">Octaseps</taxonomicName>
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alata
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</emphasis>
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sp. nov. — 3. Head and thorax, side view; — 4. Thorax and base of abdomen, subdorsal view showing the thickened rim between thorax and abdomen (marked) (photographs Jessica Litman).
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</paragraph>
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</caption>
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<caption httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="4" pageNumber="131" targetBox="[145,1020,127,1285]" targetPageId="4">
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<paragraph blockId="4.[140,1025,1326,1393]" pageId="4" pageNumber="131">
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Figs 5–10.
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<emphasis box="[241,372,1328,1347]" italics="true" pageId="4" pageNumber="131">
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<taxonomicName box="[241,321,1328,1347]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">Octaseps</taxonomicName>
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alata
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</emphasis>
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sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view.
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</paragraph>
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</caption>
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<paragraph blockId="5.[155,1040,123,529]" pageId="5" pageNumber="132">
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(
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<figureCitation box="[163,228,123,148]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="5" pageNumber="132">Fig. 8</figureCitation>
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) shorter, posterior margin with median emargination surrounded by apair of submedian rounded lobes and sublateral shallow emarginations. Genital capsule (
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<figureCitation box="[162,273,182,206]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="5" pageNumber="132">Figs 9–10</figureCitation>
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) rotated 180°; epandrium strongly sclerotized apically, with amedian shining projection; gonocoxites oblique, diverging apically, somewhat wingshaped,heavily sclerotized, strongly pilose apically; parameres complex, heavily sclerotized, teethed both basally and distally; aedeagus elongate, geniculate (
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<figureCitation captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="5" pageNumber="132">Fig. 9</figureCitation>
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), widened medially and spatulate apically, with anearly translucent dorsal process (
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<figureCitation box="[215,295,329,353]" captionStart="Figs 5 – 10" captionStartId="4.[140,178,1328,1347]" captionTargetBox="[145,1020,127,1285]" captionTargetId="figure@4.[140,1024,122,1288]" captionTargetPageId="4" captionText="Figs 5 – 10. Octaseps alata sp. nov. ♂: — 5. Spiracular sclerite; — 6. Halter; — 7. Tergite 7; — 8. Sternite 7; — 9. Genital capsule, ventral view; — 10. Genital capsule, dorsal view." httpUri="https://zenodo.org/record/235403/files/figure.png" pageId="5" pageNumber="132">Fig. 10</figureCitation>
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). Sperm pump lying free in abdominal cavity, separated from genital capsule.
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</paragraph>
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<paragraph blockId="5.[155,1040,123,529]" box="[215,421,387,411]" pageId="5" pageNumber="132">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="132" type="distribution">
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<paragraph blockId="5.[155,1040,123,529]" pageId="5" pageNumber="132">
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<emphasis box="[215,345,417,441]" italics="true" pageId="5" pageNumber="132">Distribution</emphasis>
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.Only known from the type-locality in Kwa Zulu Natal, South Africa.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="132" type="etymology">
|
||
<paragraph blockId="5.[155,1040,123,529]" pageId="5" pageNumber="132">
|
||
<emphasis box="[215,332,475,499]" italics="true" pageId="5" pageNumber="132">Etymology</emphasis>
|
||
. The specific epithet of the name is a Latin adjective meaning «winged», as areference to the shape of the male genital capsule.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="134" pageId="5" pageNumber="132" type="discussion">
|
||
<paragraph blockId="5.[155,1040,594,1528]" pageId="5" pageNumber="132">
|
||
This genus lacks typical features of any of the tribes of
|
||
<taxonomicName box="[751,881,594,618]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="subFamily" subFamily="Scatopsinae">Scatopsinae</taxonomicName>
|
||
, hence placing the genus within the system is particularly challenging. There is no question that
|
||
<taxonomicName box="[155,253,653,676]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[155,253,653,676]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName box="[424,555,652,676]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="subFamily" subFamily="Scatopsinae">Scatopsinae</taxonomicName>
|
||
. This is suggested by the sperm pump lying free in the abdomen, the fold in the wing between M2 and M4,and the presence of setation on the frons between the antennae and the eye-bridge, among other features synapomorphic for the subfamily (
|
||
<bibRefCitation author="Amorim, D. de S" bookContentInfo="172 pp." box="[523,679,740,765]" journalOrPublisher="Thesis, Sao Paulo" pageId="5" pageNumber="132" refString="Amorim, D. de S. 1982. Sistematica filogenetica dos Scatopsidae (Diptera: Oligoneura: Bibionomorpha). - Thesis, Sao Paulo, 172 pp., 76 pl." title="Sistematica filogenetica dos Scatopsidae (Diptera: Oligoneura: Bibionomorpha)" type="book" year="1982">Amorim 1982</bibRefCitation>
|
||
). The lack of setation on the wing membrane and veins also indicates that it does not belong in the
|
||
<taxonomicName box="[850,1040,770,794]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Rhegmoclematini">Rhegmoclematini</taxonomicName>
|
||
(although
|
||
<taxonomicName box="[264,414,799,823]" class="Insecta" family="Scatopsidae" genus="Diamphidicus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[264,414,799,823]" italics="true" pageId="5" pageNumber="132">Diamphidicus</emphasis>
|
||
</taxonomicName>
|
||
also lacks this feature). The presence of pedicelar setae in
|
||
<taxonomicName box="[155,253,829,852]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[155,253,829,852]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
also shows that it does not fit into the
|
||
<taxonomicName box="[707,900,828,852]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Rhegmoclematini">Rhegmoclematini</taxonomicName>
|
||
. Finally, the absence of the tegula of the halter is an apomorphic feature shared by all
|
||
<taxonomicName class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="subFamily" subFamily="Scatopsinae">Scatopsinae</taxonomicName>
|
||
except the
|
||
<taxonomicName box="[316,509,887,911]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Rhegmoclematini">Rhegmoclematini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[155,1040,594,1528]" pageId="5" pageNumber="132">
|
||
The elongate R4+5 joining costa beyond the middle of the wing is aplesiomorphic condition in the evolution of the family. In the
|
||
<taxonomicName box="[758,885,946,970]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="subFamily" subFamily="Scatopsinae">Scatopsinae</taxonomicName>
|
||
R4+5 reaching Cbefore the mid of the wing is seen only in
|
||
<taxonomicName authority="Enderlein, 1912" authorityName="Enderlein" authorityYear="1912" box="[627,994,975,999]" class="Insecta" family="Scatopsidae" genus="Swammerdamella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[627,818,975,999]" italics="true" pageId="5" pageNumber="132">Swammerdamella</emphasis>
|
||
Enderlein, 1912
|
||
</taxonomicName>
|
||
and other higher genera of Swammerdamellini – the relatively long R4+5 in
|
||
<taxonomicName box="[942,1040,1005,1028]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[942,1040,1005,1028]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
certainly excludes the genus from the clade that has
|
||
<taxonomicName authority="Cook, 1978" authorityName="Cook" authorityYear="1978" box="[764,1035,1034,1058]" class="Insecta" family="Scatopsidae" genus="Akorhexoza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[764,890,1034,1058]" italics="true" pageId="5" pageNumber="132">Akorhexoza</emphasis>
|
||
<bibRefCitation author="Cook, E. F." box="[902,1035,1034,1058]" journalOrPublisher="Pan-Pacific Entomologist" pageId="5" pageNumber="132" pagination="31 - 37" part="54" refString="Cook, E. F. 1978. A new genus and five new species of Scatopsidae from California, Mexico, El Salvador and Peru (Diptera). - Pan-Pacific Entomologist 54 (1): 31 - 37." title="A new genus and five new species of Scatopsidae from California, Mexico, El Salvador and Peru (Diptera)" type="journal article" year="1978">Cook, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Enderlein, 1936" authorityName="Enderlein" authorityYear="1936" box="[155,429,1063,1087]" class="Insecta" family="Scatopsidae" genus="Rhexoza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[155,245,1063,1087]" italics="true" pageId="5" pageNumber="132">Rhexoza</emphasis>
|
||
Enderlein, 1936
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Cook, 1975" authorityName="Cook" authorityYear="1975" box="[441,688,1063,1087]" class="Insecta" family="Scatopsidae" genus="Quateiella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[441,554,1063,1087]" italics="true" pageId="5" pageNumber="132">Quateiella</emphasis>
|
||
<bibRefCitation author="Cook, E. F." box="[562,688,1063,1087]" journalOrPublisher="Pan-Pacific Entomologist" pageId="5" pageNumber="132" pagination="62 - 75" part="51" refString="Cook, E. F. 1975. A reconsideration of the Nearctic Rhexoza (Diptera: Scatopsidae). - Pan-Pacific Entomologist 51: 62 - 75." title="A reconsideration of the Nearctic Rhexoza (Diptera: Scatopsidae)" type="journal article" year="1975">Cook, 1975</bibRefCitation>
|
||
</taxonomicName>
|
||
etc. (
|
||
<bibRefCitation author="Amorim, D. de S." box="[750,909,1063,1087]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="132" pagination="41 - 53" part="1640" refString="Amorim, D. de S. 2007. Two new genera of Swammerdamellini (Diptera, Scatopsidae), with a discussion of the position of the species of Rhexoza. - Zootaxa 1640: 41 - 53." title="Two new genera of Swammerdamellini (Diptera, Scatopsidae), with a discussion of the position of the species of Rhexoza" type="journal article" year="2007">Amorim 2007</bibRefCitation>
|
||
). The shape of the anterior thoracic spiracular sclerite is also helpful in this discussion. In all genera of Swammerdamellini but in
|
||
<taxonomicName authority="Freeman, 1990" authorityName="Freeman" authorityYear="1990" box="[548,853,1122,1146]" class="Insecta" family="Scatopsidae" genus="Pararhexosa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[548,686,1122,1146]" italics="true" pageId="5" pageNumber="132">Pararhexosa</emphasis>
|
||
<bibRefCitation author="Freeman, P." box="[693,853,1122,1146]" journalOrPublisher="Entomologist's monthly Magazine" pageId="5" pageNumber="132" pagination="9 - 19" part="126" refString="Freeman, P. 1990. Redescription of seven Oriental species of Scatopsidae (Diptera) described by F. W. Edwards in the genus Scatopse. - Entomologist's monthly Magazine 126: 9 - 19." title="Redescription of seven Oriental species of Scatopsidae (Diptera) described by F. W. Edwards in the genus Scatopse" type="journal article" year="1990">Freeman, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
it is more or less elongated, the posterior end of the sclerite often being more slender than the anterior end. The anterior spiracle sclerite in
|
||
<taxonomicName box="[569,667,1181,1204]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[569,667,1181,1204]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
has adorso-anterior extension, a condition seen in the genera of
|
||
<taxonomicName box="[494,608,1210,1234]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Scatopsini">Scatopsini</taxonomicName>
|
||
, as well as in
|
||
<taxonomicName box="[759,897,1210,1234]" class="Insecta" family="Scatopsidae" genus="Pararhexosa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[759,897,1210,1234]" italics="true" pageId="5" pageNumber="132">Pararhexosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[155,1040,594,1528]" pageId="5" pageNumber="132">
|
||
The male terminalia bearing two pairs of well-developed processes, and a complex elongate aedeagus are features seen in all four genera of
|
||
<taxonomicName box="[838,953,1268,1292]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Scatopsini">Scatopsini</taxonomicName>
|
||
. Indeed,
|
||
<taxonomicName box="[155,253,1298,1321]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[155,253,1298,1321]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
has important similarities with
|
||
<taxonomicName authority="Haenni &Huerta, 2014" authorityName="Haenni &Huerta" authorityYear="2014" box="[601,985,1298,1322]" class="Insecta" family="Scatopsidae" genus="Aztecatopse" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[601,728,1298,1321]" italics="true" pageId="5" pageNumber="132">Aztecatopse</emphasis>
|
||
<bibRefCitation author="Haenni, J. - P. & Huerta, H." box="[736,985,1298,1322]" journalOrPublisher="Revue suisse de Zoologie" pageId="5" pageNumber="132" pagination="249 - 260" part="121" refString="Haenni, J. - P. & Huerta, H. 2014. The identity of Scatopse diabolica Duda, 1928, with description of a new genus from Mexico (Diptera, Scatopsidae). - Revue suisse de Zoologie 121 (2): 249 - 260." title="The identity of Scatopse diabolica Duda, 1928, with description of a new genus from Mexico (Diptera, Scatopsidae)" type="journal article" year="2014">Haenni &Huerta, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
and the discussion in
|
||
<bibRefCitation author="Haenni, J. - P. & Huerta, H." box="[341,598,1327,1351]" journalOrPublisher="Revue suisse de Zoologie" pageId="5" pageNumber="132" pagination="249 - 260" part="121" refString="Haenni, J. - P. & Huerta, H. 2014. The identity of Scatopse diabolica Duda, 1928, with description of a new genus from Mexico (Diptera, Scatopsidae). - Revue suisse de Zoologie 121 (2): 249 - 260." title="The identity of Scatopse diabolica Duda, 1928, with description of a new genus from Mexico (Diptera, Scatopsidae)" type="journal article" year="2014">Haenni &Huerta (2014)</bibRefCitation>
|
||
applies here to aconsiderable extent. In fact, the similarities of
|
||
<taxonomicName box="[411,549,1356,1380]" class="Insecta" family="Scatopsidae" genus="Pararhexosa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[411,549,1356,1380]" italics="true" pageId="5" pageNumber="132">Pararhexosa</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName box="[614,712,1357,1380]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[614,712,1357,1380]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
may actually suggest that the position of
|
||
<taxonomicName box="[278,416,1386,1410]" class="Insecta" family="Scatopsidae" genus="Pararhexosa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[278,416,1386,1410]" italics="true" pageId="5" pageNumber="132">Pararhexosa</emphasis>
|
||
</taxonomicName>
|
||
in the Swammerdamellini is mistaken, and that the genus could more properly belong in the
|
||
<taxonomicName box="[532,647,1415,1439]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Scatopsini">Scatopsini</taxonomicName>
|
||
. The inclusion of
|
||
<taxonomicName box="[842,940,1415,1438]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[842,940,1415,1438]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[948,1040,1416,1440]" pageId="5" pageNumber="132" rank="genus">gen.nov.</taxonomicNameLabel>
|
||
in the
|
||
<taxonomicName box="[230,347,1444,1468]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Scatopsini">Scatopsini</taxonomicName>
|
||
is clearly the best solution for the time being. The peculiar thickened suture between the thorax and the abdomen of
|
||
<taxonomicName box="[761,859,1474,1497]" class="Insecta" family="Scatopsidae" genus="Octaseps" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen.nov.">
|
||
<emphasis box="[761,859,1474,1497]" italics="true" pageId="5" pageNumber="132">Octaseps</emphasis>
|
||
</taxonomicName>
|
||
is unique among the
|
||
<taxonomicName box="[195,322,1503,1527]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
|
||
and is evidently an autapomorphy of the new genus.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/235404/files/figure.png" pageId="6" pageNumber="133" targetBox="[144,1020,127,1271]" targetPageId="6">
|
||
<paragraph blockId="6.[140,1025,1326,1415]" pageId="6" pageNumber="133">
|
||
Figs 11–14:
|
||
<taxonomicName authority="Cook" authorityName="Cook" box="[249,481,1328,1347]" class="Insecta" family="Scatopsidae" genus="Psacotes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="processa">
|
||
<emphasis box="[249,410,1328,1346]" italics="true" pageId="6" pageNumber="133">Psacotes processa</emphasis>
|
||
(Cook)
|
||
</taxonomicName>
|
||
: — 11. ♂ genital capsule, ventral view; — 12: ♂ genital capsule, dorsal view; — 13: ♀ terminalia, dorsal view; — 14: ♀ terminalia, ventral view. Abbreviations: ae – aedeagus; ae pl – aedeagal plate; ep – epandrium; pa – paramere; st – sternite; t – tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[155,1039,123,294]" pageId="7" pageNumber="134">
|
||
Within the
|
||
<taxonomicName box="[338,454,123,147]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="tribe" tribe="Scatopsini">Scatopsini</taxonomicName>
|
||
, the new genus may be more closely related to
|
||
<taxonomicName authority="Cook, 1974" authorityName="Cook" authorityYear="1974" class="Insecta" family="Scatopsidae" genus="Apiloscatopse" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="7" pageNumber="134">Apiloscatopse</emphasis>
|
||
Cook, 1974
|
||
</taxonomicName>
|
||
– agenus mainly Holarctic in distribution, also known from an Afrotropical species,
|
||
<taxonomicName authority="Cook, 1965" authorityName="Cook" authorityYear="1965" box="[415,690,182,206]" class="Insecta" family="Scatopsidae" genus="Apiloscatopse" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="species" species="labellata">
|
||
<emphasis box="[415,539,182,206]" italics="true" pageId="7" pageNumber="134">A. labellata</emphasis>
|
||
(
|
||
<bibRefCitation author="Cook, E. F." box="[553,682,182,206]" editor="Hanstrom, B." journalOrPublisher="Stockholm: Statens Naturvetenskapliga Foskinsrad" pageId="7" pageNumber="134" pagination="469 - 481" part="11" refString="Cook, E. F. 1965 a. V. Diptera (Nematocera): Scatopsidae. In: Hanstrom, B., Brinck, P. & Rudebeck, G. (eds), South African animal life. Results of the Lund University Expedition in 1950 - 1951, Volume 11. - Stockholm: Statens Naturvetenskapliga Foskinsrad, pp. 469 - 481." title="V. Diptera (Nematocera): Scatopsidae" type="journal article" volumeTitle="South African animal life. Results of the Lund University Expedition in 1950 - 1951" year="1965">Cook, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, from South Africa – or to
|
||
<taxonomicName authority="Cook, 1974" authorityName="Cook" authorityYear="1974" class="Insecta" family="Scatopsidae" genus="Pharsoreichertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="7" pageNumber="134">Pharsoreichertella</emphasis>
|
||
Cook, 1974
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName box="[448,573,211,235]" class="Insecta" family="Scatopsidae" genus="Apiloscatopse" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="species" species="labellata">
|
||
<emphasis box="[448,573,211,235]" italics="true" pageId="7" pageNumber="134">A. labellata</emphasis>
|
||
</taxonomicName>
|
||
was originally described based on afemale and placed in the genus
|
||
<taxonomicName authority="Cook, 1965a" authorityName="Cook" authorityYear="1965" box="[409,667,241,265]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[409,504,241,264]" italics="true" pageId="7" pageNumber="134">Scatopse</emphasis>
|
||
(
|
||
<bibRefCitation author="Cook, E. F." box="[518,660,241,265]" editor="Hanstrom, B." journalOrPublisher="Stockholm: Statens Naturvetenskapliga Foskinsrad" pageId="7" pageNumber="134" pagination="469 - 481" part="11" refString="Cook, E. F. 1965 a. V. Diptera (Nematocera): Scatopsidae. In: Hanstrom, B., Brinck, P. & Rudebeck, G. (eds), South African animal life. Results of the Lund University Expedition in 1950 - 1951, Volume 11. - Stockholm: Statens Naturvetenskapliga Foskinsrad, pp. 469 - 481." title="V. Diptera (Nematocera): Scatopsidae" type="journal article" volumeTitle="South African animal life. Results of the Lund University Expedition in 1950 - 1951" year="1965" yearSuffix="a">Cook, 1965a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Its precise position within the scatopsines, however, should be still considered as uncertain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |