208 lines
22 KiB
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208 lines
22 KiB
XML
<document ID-CLB-Dataset="9152" ID-DOI="http://dx.doi.org/10.3897/zookeys.1007.59351" ID-GBIF-Dataset="c7bddb47-5b7f-41c5-b6cf-17fa4f0210ef" ID-PMC="PMC7788071" ID-Pensoft-Pub="1313-2970-1007-129" ID-Pensoft-UUID="ADEE7499EE9853E1BDD6ECF092EE97F9" ID-PubMed="33505183" ID-ZooBank="59B6D78F4C814260B82BCE74CDC6A13D" ModsDocID="1313-2970-1007-129" checkinTime="1609377136731" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob & Balke, Michael" docDate="2020" docId="37E1527A36915673A58CB38E09A82CFC" docLanguage="en" docName="ZooKeys 1007: 129-143" docOrigin="ZooKeys 1007" docPubDate="2020-12-30" docSource="http://dx.doi.org/10.3897/zookeys.1007.59351" docTitle="Exocelina amabilis Shaverdo & Balke 2020, sp. nov." docType="treatment" docUuid="C7F77D9E-27BD-446D-9E9E-415D6B6D143C" docUuidSource="ZooBank" docVersion="6" id="ADEE7499EE9853E1BDD6ECF092EE97F9" lastPageNumber="129" masterDocId="ADEE7499EE9853E1BDD6ECF092EE97F9" masterDocTitle="Three new species of Exocelina Broun, 1886 from the southern slopes of the New Guinea central range, with introduction of the Exocelina skalei group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="143" masterPageNumber="129" pageNumber="129" updateTime="1732824863550" updateUser="ExternalLinkService">
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<mods:mods id="D5773F0A9F4FF65410C44535B447D87F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="47226FC6541D8E4A8A216B580FE26023">
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<mods:title id="BB7FFF4E082E24977764269BD39D7ADC">Three new species of Exocelina Broun, 1886 from the southern slopes of the New Guinea central range, with introduction of the Exocelina skalei group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F5C49A72B2A2960B76315B9A36AF1783" type="personal">
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<mods:role id="4D9B9C0D2022256CE79D5734A33959DE">
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<mods:roleTerm id="D95C329F577110C1E2FA66EBC7900B2A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92485FC44A21F8AF4E241DE47C30AE36">Shaverdo, Helena</mods:namePart>
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<mods:affiliation id="51E5C5D5510D30357BB6F56433FC1910">Naturhistorisches Museum Wien, Burgring 7, 1010, Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier id="7C0AB72AA235BC6AA7A42AF1FEA8FB6F" type="email">shaverdo@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name id="652321D8242265216B2119917F76A667" type="personal">
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<mods:role id="08D1C1F5C0F4B872C8CDCEBC9CA0BF12">
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<mods:roleTerm id="70E97B9EA1B24C9FC28D4700161F432B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4163DD588897DCDF82A09170A4996BDD">Surbakti, Suriani</mods:namePart>
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<mods:affiliation id="39E29289D75B16EA01B0144EBBB68839">Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name id="21C1FD27ACBC1061BC0A50A28D24DE09" type="personal">
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<mods:role id="7FE044C8EACD6475783BCD9A73FBFD09">
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<mods:roleTerm id="B39DE8BEE5C4EB5759B547E7825BB5AB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="80AC49F3F159D6597A38DC7C857111F5">Sumoked, Bob</mods:namePart>
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<mods:affiliation id="E3C9B09400DDBCCC0D2CA95D76324424">Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name id="125E08DC14ADCF0DA0254A65F7877C14" type="personal">
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<mods:role id="DBB2F17F3DA7F6396436AB2934B3F9A8">
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<mods:roleTerm id="3E33BD8B95AF20828BD8E2742D277FA8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E7DBBAB6AA885B28B17E2EE73D207FD6">Balke, Michael</mods:namePart>
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<mods:nameIdentifier id="84EC647C7E7CA481EB25B7C87CA8F627" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
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<mods:affiliation id="345A2B7F709B919EA778681160F3EE6A">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany & GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="35B22004148DFC6BB14A1538E61F1CCD">text</mods:typeOfResource>
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<mods:relatedItem id="E85C1D904043F067BE92265569C9C6FA" type="host">
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<mods:titleInfo id="80770CEF5BFCF548C01C1B8B491B78CB">
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<mods:title id="69B2712E4F6CA82ABFA1A8E84727A4CC">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="2CE5BB573CF84D2579FA931EBAC15E06">
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<mods:date id="D154CFF59A63A4580B981FC2852ECDC5">2020</mods:date>
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<mods:detail id="1EC7C0A71C4CD6D2F584B5FDDD50E56B" type="pubDate">
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<mods:number id="D9F7FB58585CC0EA797F3FD40D2E1162">2020-12-30</mods:number>
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</mods:detail>
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<mods:detail id="10FB17D4F81838E5CC15F7775A005FD7" type="volume">
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<mods:number id="D19FFBD6D49631FBE3AF8F377118A2CA">1007</mods:number>
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</mods:detail>
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<mods:extent id="2FEB44CD59A8C521BC000E1DDE519292" unit="page">
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<mods:start id="5735C5E51D253A2BBD544E3F42E0DAF6">129</mods:start>
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<mods:end id="7C847021AD6F25209799C53D3DB44A61">143</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location id="ED24298B38608CDF4933229838B80FEB">
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<mods:url id="20B2EFCDD22BD96339F15ACA98A11227">http://dx.doi.org/10.3897/zookeys.1007.59351</mods:url>
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</mods:location>
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<mods:classification id="A9F821AED1F5F9524AADE9421F32EE04">journal article</mods:classification>
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<mods:identifier id="1E65F24DF023C7A47A6C6EBCAA051A1E" type="DOI">http://dx.doi.org/10.3897/zookeys.1007.59351</mods:identifier>
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<mods:identifier id="9001D81B909E3B7A4A13548EDA4B0058" type="Pensoft-Pub">1313-2970-1007-129</mods:identifier>
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<mods:identifier id="021640DAE994558C5806C8E59C209CA2" type="ZooBank">59B6D78F4C814260B82BCE74CDC6A13D</mods:identifier>
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<mods:identifier id="54FEF33D85790FBDC5313B0DF653552E" type="Pensoft-UUID">ADEE7499EE9853E1BDD6ECF092EE97F9</mods:identifier>
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</mods:mods>
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<treatment id="37E1527A36915673A58CB38E09A82CFC" ID-GBIF-Taxon="176349000" LSID="urn:lsid:zoobank.org:act:C7F77D9E-27BD-446D-9E9E-415D6B6D143C" httpUri="http://treatment.plazi.org/id/37E1527A36915673A58CB38E09A82CFC" lastPageNumber="129" pageId="0" pageNumber="129" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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<subSubSection id="5A71ACAC17962C4D95145851E3B35318" pageId="0" pageNumber="129" type="nomenclature">
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<paragraph id="E1E6B4DE44933B571BF1E9050A01238D" pageId="0" pageNumber="129">
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<taxonomicName id="19603E5F2FAE8A1802CAA05EED7AB2BF" ID-CoL="8LZS2" LSID="http://zoobank.org/C7F77D9E-27BD-446D-9E9E-415D6B6D143C" authority="Shaverdo & Balke" authorityName="Shaverdo & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="sp. nov.">Exocelina amabilis Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel id="A1A0761A7CF416EEFF214B484E95AFEF" pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
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<figureCitation id="9FFD373C5454EC5567F7E4BC29CA6D61" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1007.59351.figure5" httpUri="https://binary.pensoft.net/fig/492244" pageId="0" pageNumber="129">Figs 5</figureCitation>
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<figureCitation id="553987B1EAF8D3932C474BDB7E423F31" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Exocelina amabilis sp. nov., holotype." figureDoi="10.3897/zookeys.1007.59351.figure6" httpUri="https://binary.pensoft.net/fig/492245" pageId="0" pageNumber="129">, 6</figureCitation>
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<figureCitation id="DDAA43B065CAC71EC03D82FF8F99B7D3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina amabilis sp. nov., paratype A median lobe in ventral view B median lobe in lateral view C right paramere in external view D right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.1007.59351.figure7" httpUri="https://binary.pensoft.net/fig/492246" pageId="0" pageNumber="129">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FE0304BF6CFCA21614C1649D6E44F70B" pageId="0" pageNumber="129" type="type locality">
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<paragraph id="4F6A2DD9B2DF9259A5F2D9C68D412506" pageId="0" pageNumber="129">Type locality.</paragraph>
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<paragraph id="86CA937F47E0C94DFC096BAA2B680A65" pageId="0" pageNumber="129">
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Indonesia: Papua Province, Pegunungan Bintang Regency, south from Ok Sibil, tributary Digul River,
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<geoCoordinate id="153A71C3186C425BEB8824BA4793D8E1" degrees="05" direction="south" minutes="03" orientation="latitude" precision="1" seconds="25.9" value="-5.0571947">05°03'25.9"S</geoCoordinate>
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,
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<geoCoordinate id="F4F5DD95BDAFE3BF13E008910F87DD8E" degrees="140" direction="east" minutes="43" orientation="longitude" precision="1" seconds="21.1" value="140.72252">140°43'21.1"E</geoCoordinate>
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, 359 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="43B574C0A2B30ABAAA314C5ED9E6A9AD" pageId="0" pageNumber="129" type="type material">
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<paragraph id="E1D995D5F0135E0A575582DCED7FF296" pageId="0" pageNumber="129">Type material.</paragraph>
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<paragraph id="5D4BEE3999346B08F8A7036DD6BD4723" pageId="0" pageNumber="129">
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<emphasis id="1C8450D64EC633C5BD0A4E5317318E64" bold="true" italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
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: male "Indonesia: Papua, S Ok Sibil, tributary Digul Riv [River], 359m, 9.vi.2015, -5,05718389 140,722535848617, Sumoked (Pap051)" (MZB).
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<emphasis id="51703CC950E1891E92023B16CDA1C093" bold="true" italics="true" pageId="0" pageNumber="129">Paratypes</emphasis>
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: 3 males, 8 females with the same label as the holotype, 2 males additionally with green text label
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<normalizedToken id="0743878EC7D3E79E763463BB51BE2402" originalValue="“6997”">"6997"</normalizedToken>
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and
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<normalizedToken id="4CB56532A154CB2A4D35D20B45AEC54A" originalValue="“6998”">"6998"</normalizedToken>
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(KSP, MZB).
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</paragraph>
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</subSubSection>
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<subSubSection id="CBCF12059FEB7764E265256EC1F4952E" pageId="0" pageNumber="129" type="description">
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<paragraph id="685A171F93809E0ABFD253366F2C4F64" pageId="0" pageNumber="129">Description.</paragraph>
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<paragraph id="2161A21465950A02922BAE39BDEAA20F" pageId="0" pageNumber="129">
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<emphasis id="93D87E10D341E5E945F7FE58E435A703" bold="true" italics="true" pageId="0" pageNumber="129">Body size and form</emphasis>
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: Beetle small: TL-H 2.65-3.45 mm, TL 3.55-3.85 mm, MW 1.75-1.9 mm (holotype: TL-H 3.25 mm, TL 3.65 mm, MW 1.8 mm), with oblong-oval habitus (Fig.
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<figureCitation id="9CA12CE8E8F65D918DD58A81F959809E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Exocelina amabilis sp. nov., holotype." figureDoi="10.3897/zookeys.1007.59351.figure6" httpUri="https://binary.pensoft.net/fig/492245" pageId="0" pageNumber="129">6</figureCitation>
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).
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</paragraph>
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<paragraph id="4F0B33EC565A0236EED8059C11A2D4B9" pageId="0" pageNumber="129">
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<emphasis id="56FB453D2C38124A248FC114050404ED" bold="true" italics="true" pageId="0" pageNumber="129">Colouration</emphasis>
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: Dorsally piceous (Fig.
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<figureCitation id="C1751D5F475733A6B872377D15CA196F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Exocelina amabilis sp. nov., holotype." figureDoi="10.3897/zookeys.1007.59351.figure6" httpUri="https://binary.pensoft.net/fig/492245" pageId="0" pageNumber="129">6</figureCitation>
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), with dark brown posterior part of head and lateral parts of pronotum, and sometimes with middle part of pronotum and elytral sutural lines; head appendages and legs yellowish-red, metathoracic legs darker distally.
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</paragraph>
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<caption id="28109D2323D614007DF1AB605AD4135F" doi="10.3897/zookeys.1007.59351.figure6" httpUri="https://binary.pensoft.net/fig/492245" pageId="0" pageNumber="129" start="Figure 6" startId="F6">
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<paragraph id="2F742CEEA73F51473BCF1DBE53CC3218" pageId="0" pageNumber="129">
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<emphasis id="AA85067FA9A7CC36885BCDBC9E25E711" bold="true" pageId="0" pageNumber="129">Figure 6.</emphasis>
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Habitus and colouration of
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<taxonomicName id="0B0C5B7956F443C73E613A5E1E260BB1" authorityName="Shaverdo & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
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<emphasis id="9E26A4251536919613A80B7ECCF84871" italics="true" pageId="0" pageNumber="129">Exocelina amabilis</emphasis>
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</taxonomicName>
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sp. nov., holotype.
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</paragraph>
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</caption>
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<paragraph id="2DD3ACD0946A93749635174D8F4A5AB6" pageId="0" pageNumber="129">
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<emphasis id="CEFF4D8DCD71BA4721D3C70A0FE887A1" bold="true" italics="true" pageId="0" pageNumber="129">Surface sculpture</emphasis>
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: Relatively shiny dorsally, with very fine, sparse punctation and distinctly impressed microreticulation. Head with dense punctation (spaces between punctures 1-3 times size of punctures), distinctly finer and sparser anteriorly and posteriorly; diameter of punctures almost equal to diameter of cells of microreticulation. Pronotum with much sparser and finer punctation than head. Elytra with very sparse and fine punctation, often inconspicuous. Pronotum and elytra with distinctly impressed microreticulation; head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine and sparse punctation.
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</paragraph>
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<paragraph id="0C6CE4F5C79665D50F52EF25A8E3432B" pageId="0" pageNumber="129">
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<emphasis id="F17C708F3F73CED67ACFD96CC6A07C25" bold="true" italics="true" pageId="0" pageNumber="129">Structures</emphasis>
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: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae.
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</paragraph>
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<paragraph id="E55C4C38F9ACE389AF85341100467DEC" pageId="0" pageNumber="129">
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<emphasis id="8D3A256890A5D6F64B8EA9037F48567D" bold="true" italics="true" pageId="0" pageNumber="129">Male</emphasis>
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: Antenna simple. Pro- and mesotarsomeres 1-3 not dilated. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 12 and posterior row of 6 short, thick, pointed setae (Fig.
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<figureCitation id="1B3CA514A3C42661D7F903FD097D2490" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina amabilis sp. nov., paratype A median lobe in ventral view B median lobe in lateral view C right paramere in external view D right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.1007.59351.figure7" httpUri="https://binary.pensoft.net/fig/492246" pageId="0" pageNumber="129">7D</figureCitation>
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). Median lobe with slightly discontinuous outline (see apically); in lateral view, more or less evenly curved, with broadly pointed apex; in ventral view, tapering to broadly pointed apex (Fig.
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<figureCitation id="717C277EC5ADE27F18B80E099B07CB0D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina amabilis sp. nov., paratype A median lobe in ventral view B median lobe in lateral view C right paramere in external view D right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.1007.59351.figure7" httpUri="https://binary.pensoft.net/fig/492246" pageId="0" pageNumber="129">7A, B</figureCitation>
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). Paramere with very deep dorsal notch, separating subdistal part; subdistal part broad, transverse, situated under apex of proximal part, with large tuft of dense, thick, flattened setae; proximal setae sparse, thin, much more inconspicuous than subdistal (Fig.
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<figureCitation id="1196D1FD6E519063016CBD48053044BA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina amabilis sp. nov., paratype A median lobe in ventral view B median lobe in lateral view C right paramere in external view D right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.1007.59351.figure7" httpUri="https://binary.pensoft.net/fig/492246" pageId="0" pageNumber="129">7C</figureCitation>
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). Abdominal ventrite 6 with extremely weak, small medial impression, visible only apically, concave apically, with 12-16 lateral striae on each side.
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</paragraph>
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<caption id="DF6DC600E39665E1A515CD3A95FFA02B" doi="10.3897/zookeys.1007.59351.figure7" httpUri="https://binary.pensoft.net/fig/492246" pageId="0" pageNumber="129" start="Figure 7" startId="F7">
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<paragraph id="399C8F253D91436676FDF01A63296B54" pageId="0" pageNumber="129">
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<emphasis id="B292F9404206B60DF07183899AB66702" bold="true" pageId="0" pageNumber="129">Figure 7.</emphasis>
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<taxonomicName id="6E1C8169C75BD1D7908BDCF956F7942E" authorityName="Shaverdo & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
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<emphasis id="EC15D0FCAE32AACA0589F9BFF12C177E" italics="true" pageId="0" pageNumber="129">Exocelina amabilis</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis id="5C5758CFDA5DF81C6EDC4F4B2D14C195" bold="true" pageId="0" pageNumber="129">A</emphasis>
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median lobe in ventral view
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<emphasis id="71D3E3F79427A4C3597FC85EDF02F615" bold="true" pageId="0" pageNumber="129">B</emphasis>
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median lobe in lateral view
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<emphasis id="F0D1F5C379143B526EE11B7F31E11BCA" bold="true" pageId="0" pageNumber="129">C</emphasis>
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right paramere in external view
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<emphasis id="527DA2BA4E7070B576D8EB445BA1AEAB" bold="true" pageId="0" pageNumber="129">D</emphasis>
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right male protarsomeres 4-5 in ventral view.
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</paragraph>
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</caption>
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<paragraph id="CDDC5B70B1873607B0A749F4EE45BF12" pageId="0" pageNumber="129">
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<emphasis id="2537A8065D759D33754A12D126131B59" bold="true" italics="true" pageId="0" pageNumber="129">Female</emphasis>
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: Pro- and mesotarsi not modified. Abdominal ventrite 6 slightly truncate or very slightly concave apically, without medial impression and lateral striae.
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</paragraph>
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</subSubSection>
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<subSubSection id="9E4425D18C5BAB0D0435FBA1B24D5492" pageId="0" pageNumber="129" type="habitat">
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<paragraph id="E7D4EECCD8280574956A55EF16E6CA68" pageId="0" pageNumber="129">Habitat.</paragraph>
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<paragraph id="A00B9F6B12BA370DA77FD1535671C836" pageId="0" pageNumber="129">The specimens were collected from the gravel banks of a primary forest stream. The beetles were mainly hidden in the coarse gravel and were only obtained after some digging.</paragraph>
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</subSubSection>
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<subSubSection id="E7278A05A521E18C8D1E36D587E37CF5" pageId="0" pageNumber="129" type="distribution">
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<paragraph id="89052D72DD1759ED2BD7BC60C7CBDA66" pageId="0" pageNumber="129">Distribution.</paragraph>
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<paragraph id="EDF33233C29D26FA9AD351AA60292685" pageId="0" pageNumber="129">
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Indonesia: Papua Province. The species is known only from the type locality (Fig.
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<figureCitation id="1A72E53A25C01ACFFC03D6A80A6AAE34" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1007.59351.figure5" httpUri="https://binary.pensoft.net/fig/492244" pageId="0" pageNumber="129">5</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="ABEFB9FC3E0622240CDE75CD99026583" pageId="0" pageNumber="129" type="etymology">
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<paragraph id="B0CB28A63F0DC156A0309BB1C46AB324" pageId="0" pageNumber="129">Etymology.</paragraph>
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<paragraph id="F24330BF58497C5F2EB2ECD6CD8CFDEC" pageId="0" pageNumber="129">
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The species name is a Latin adjective and means
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<normalizedToken id="465B56009B31BE115B4CE10860E95E1B" originalValue="“loveable”">"loveable"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C702E379B007144AF043299F25D0D47E" pageId="0" pageNumber="129" type="affinities">
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<paragraph id="CD839C800C0899C27C1927C65357B06C" pageId="0" pageNumber="129">Affinities.</paragraph>
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<paragraph id="FD7AFB17046DD1D6C233E7D6D61F677B" pageId="0" pageNumber="129">
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The species evidently belongs to the
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<taxonomicName id="1DD275C1F128EE5A1D63685E987D353D" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="ekari">
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<emphasis id="57F140BFCFB94F0D4CE9546DE91C56D2" italics="true" pageId="0" pageNumber="129">E. ekari</emphasis>
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|
</taxonomicName>
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|
group due to the discontinuous outline of its median lobe. It is similar to
|
|
<taxonomicName id="1890A8EDA638B3A4E37098AC89D20FF0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. utowaensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="utowaensis">
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<emphasis id="2728602143C45B8FFE62ABC048B81C30" italics="true" pageId="0" pageNumber="129">E. utowaensis</emphasis>
|
|
</taxonomicName>
|
|
Shaverdo, Hendrich & Balke, 2012 in modification of the abdominal ventrite 6, body size, colouration and shape but distinctly differs from it in having a pronotal bead (absent in
|
|
<taxonomicName id="DF9297C5BA29D44A9592E53A6D3E70C8" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. utowaensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="utowaensis">
|
|
<emphasis id="B1DB0DDA81AD220838FCD283DC51BBE6" italics="true" pageId="0" pageNumber="129">E. utowaensis</emphasis>
|
|
</taxonomicName>
|
|
) and different shape of the median lobe, paramere, and anterolateral hook-like seta of the male protarsomere 4 (it is also larger in
|
|
<taxonomicName id="67E7E4A6CF2DB0ACA742C7EE7FBE6D04" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. utowaensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="utowaensis">
|
|
<emphasis id="099B20549046FBB1A07FF7D0852DD9CF" italics="true" pageId="0" pageNumber="129">E. utowaensis</emphasis>
|
|
</taxonomicName>
|
|
). The species is also similar to
|
|
<taxonomicName id="8268502A2115091B5A76212B2E5B9445" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. athesphatos" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="athesphatos">
|
|
<emphasis id="C66718D470117550A8907A36762997DF" italics="true" pageId="0" pageNumber="129">E. athesphatos</emphasis>
|
|
</taxonomicName>
|
|
in modification of the abdominal ventrite 6 and paramere, but distinctly differs from it in smaller size and having simple male antennae.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |