184 lines
23 KiB
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184 lines
23 KiB
XML
<document ID-CLB-Dataset="276892" ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.109702" ID-GBIF-Dataset="9fe0bff4-b6c2-415d-ba49-0f1e074a9710" ID-Pensoft-Pub="1313-2970-1184-1" ID-Pensoft-UUID="C2F3B4D92BF558A0AB92E0607CB93262" ID-ZooBank="3D745E43FB6C48A8B089DDAEE1A422BB" ModsDocID="1313-2970-1184-1" checkinTime="1699693871686" checkinUser="pensoft" docAuthor="Yamada, Aiki, Nguyen, Dai Dac & Eguchi, Katsuyuki" docDate="2023" docId="ECBB48E065725887BCEC448545BBF690" docLanguage="en" docName="ZooKeys 1184: 1-17" docOrigin="ZooKeys 1184" docPubDate="2023-11-10" docSource="http://dx.doi.org/10.3897/zookeys.1184.109702" docTitle="Eburopone wroughtoni" docType="treatment" docVersion="3" id="C2F3B4D92BF558A0AB92E0607CB93262" lastPageNumber="1" masterDocId="C2F3B4D92BF558A0AB92E0607CB93262" masterDocTitle="First discovery of the ant genus Eburopone Borowiec, 2016 (Hymenoptera, Formicidae, Dorylinae) in the Oriental realm, with description of a new species from Vietnam" masterLastPageNumber="17" masterPageNumber="1" pageNumber="1" updateTime="1699701014511" updateUser="ExternalLinkService">
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<mods:titleInfo id="4F163EA92115998A510A0C80CE71D75E">
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<mods:title id="6D3741825189441CAF30E762C28AA3E1">First discovery of the ant genus Eburopone Borowiec, 2016 (Hymenoptera, Formicidae, Dorylinae) in the Oriental realm, with description of a new species from Vietnam</mods:title>
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<mods:namePart id="B76AC3E9E3F144FA73AE0E80EF56CC28">Yamada, Aiki</mods:namePart>
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<mods:nameIdentifier id="65C577439D19E10B336EEDFC0949EF24" type="ORCID">https://orcid.org/0000-0002-2141-0661</mods:nameIdentifier>
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<mods:affiliation id="7E3DB196B3C0417F84255C71E74845AA">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan</mods:affiliation>
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<mods:nameIdentifier id="578EA12B31B3EF5E0EDE44397D46001D" type="email">aiki.ymd@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F4441DA8B79F228AA23C51E7E7C0CEC2">Nguyen, Dai Dac</mods:namePart>
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<mods:nameIdentifier id="BC22149B538845CE81169D303CF05EE3" type="ORCID">https://orcid.org/0009-0006-8024-2553</mods:nameIdentifier>
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<mods:affiliation id="DF3A996C8C1BECA30A38932546ECF58F">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan & Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Cau Giay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="4E5D7158909B521AA27022A3EBFA669C">Eguchi, Katsuyuki</mods:namePart>
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<mods:nameIdentifier id="981018CF078FE32B90ADD8B1A69CF22F" type="ORCID">https://orcid.org/0000-0002-1054-1295</mods:nameIdentifier>
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<mods:affiliation id="A353CEF50ACA6718DABC9EE255CDBCD1">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan & Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, 1 - 12 - 4 Sakamoto, Nagasaki, Nagasaki, 852 - 8523, Japan</mods:affiliation>
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<mods:title id="B9C8946C29F32957CCD91A53B609AF52">ZooKeys</mods:title>
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<mods:date id="BA912A6B09F8CF5A2A7F7AF800574C94">2023</mods:date>
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<mods:number id="7BE22B949F44E14711C817EB9B32A0B4">2023-11-10</mods:number>
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<mods:number id="1D57880567D0C7D17C11B6E64CF41063">1184</mods:number>
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<mods:url id="2683977496BF5E4B25084167644913DF">http://dx.doi.org/10.3897/zookeys.1184.109702</mods:url>
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<mods:classification id="F258AA0499B1A6E6BACFF0AF38B55D7D">journal article</mods:classification>
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<mods:identifier id="1179173B6E1F5F596C95065415A77142" type="DOI">http://dx.doi.org/10.3897/zookeys.1184.109702</mods:identifier>
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<treatment id="ECBB48E065725887BCEC448545BBF690" ID-GBIF-Taxon="217559628" LSID="urn:lsid:plazi:treatment:ECBB48E065725887BCEC448545BBF690" httpUri="http://treatment.plazi.org/id/ECBB48E065725887BCEC448545BBF690" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="2684A348934785222AFB11851004168F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="27237C6E7FD318AE7831278DDA8D4869" pageId="0" pageNumber="1">
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<taxonomicName id="5AD0DE626036BDA5AC21A3092DBE1EE6" LSID="ECBB48E0-6572-5887-BCEC-448545BBF690" authority="(Forel, 1910)" baseAuthorityName="Forel" baseAuthorityYear="1910" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone wroughtoni" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wroughtoni">Eburopone wroughtoni (Forel, 1910)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="279D0E411578521CE9B7858AE8F29265" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="D2AAE7F0736A0A1B0B97FE03F1F06300" pageId="0" pageNumber="1">
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<taxonomicName id="F5362DEBDF18A20B732F522B5C190936" authorityName="Forel" authorityYear="1910" baseAuthorityName="Forel" baseAuthorityYear="1910" class="Insecta" family="Formicidae" genus="Cerapachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapachys wroughtoni" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wroughtoni">Cerapachys wroughtoni</taxonomicName>
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Forel, 1910: 422.
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<bibRefCitation id="69DC27FB9110D2360D422B1340F5CAA1" DOI="https://doi.org/10.5281/zenodo.54863" author="Emery, C" journalOrPublisher="Genera Insectorum" pageId="0" pageNumber="1" pagination="1 - 125" refId="B12" refString="Emery, C, 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1 - 125, DOI: https://doi.org/10.5281/zenodo.54863" title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae." url="https://doi.org/10.5281/zenodo.54863" volume="118" year="1911">Emery 1911</bibRefCitation>
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: 9;
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<bibRefCitation id="48A70CB8DFF2AEF924075772EA68AF15" author="Arnold, G" journalOrPublisher="Annals of the South African Museum, Cape Town" pageId="0" pageNumber="1" pagination="1 - 159" publicationUrl="https://biostor.org/reference/109829" refId="B2" refString="Arnold, G, 1915. A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae. Annals of the South African Museum, Cape Town 14: 1 - 159, https://biostor.org/reference/109829" title="A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae." url="https://biostor.org/reference/109829" volume="14" year="1915">Arnold 1915</bibRefCitation>
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: 16;
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<bibRefCitation id="AAB29C3766272C554F77A0F1B84BC26E" author="Wheeler, WM" journalOrPublisher="Bulletin of the American Museum of Natural History, New York" pageId="0" pageNumber="1" publicationUrl="http://hdl.handle.net/2246/932" refId="B25" refString="Wheeler, WM, 1922. Ants of the American Museum Congo Expedition. Bulletin of the American Museum of Natural History, New York, http://hdl.handle.net/2246/932" title="Ants of the American Museum Congo Expedition." url="http://hdl.handle.net/2246/932" year="1922">Wheeler 1922</bibRefCitation>
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: 756; Brown 1975: 24, 63;
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<bibRefCitation id="C7EF83629C83F833D0AD8E252F133D7E" author="Bolton, B" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="0" pageNumber="1" refId="B3" refString="Bolton, B, 1995. A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts" title="A New General Catalogue of the Ants of the World." year="1995">Bolton 1995</bibRefCitation>
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: 145;
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<bibRefCitation id="B8FF9F444656B8BBEA512077D0540494" author="Collingwood, CA" editor="van Harten, A" journalOrPublisher="4). Dar Al Ummah Publishers, Abu Dhabi" pageId="0" pageNumber="1" pagination="405 - 474" refId="B11" refString="Collingwood, CA, Agosti, D, Sharaf, MR, van Harten, A, 2011. Order Hymenoptera, family Formicidae. In: van Harten, A, Ed., Arthropod Fauna of the UAE (Vol. 4). Dar Al Ummah Publishers, Abu Dhabi: 405 - 474" title="Order Hymenoptera, family Formicidae." volumeTitle="Arthropod Fauna of the UAE (Vol." year="2011">Collingwood et al. 2011</bibRefCitation>
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(dubious record): 410;
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<bibRefCitation id="52881D4924FE435D3315EAB008B0E3D3" author="Borowiec, L" journalOrPublisher="Genus" pageId="0" pageNumber="1" pagination="1 - 340" refId="B5" refString="Borowiec, L, 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus 25 (1-2): 1 - 340" title="Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae)." volume="25" year="2014">Borowiec 2014</bibRefCitation>
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: 62. Combination in
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<taxonomicName id="3514CB14B428884E3726B8EA9277C8B8" authority="" family="Formicidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. (Cerapachys)" order="Hymenoptera" pageId="0" pageNumber="1" rank="subGenus" subGenus="Cerapachys">C. (Cerapachys)</taxonomicName>
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:
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<bibRefCitation id="9AAFC40DDDD37F07F860CFDDAE500628" DOI="https://doi.org/10.5281/zenodo.54863" author="Emery, C" journalOrPublisher="Genera Insectorum" pageId="0" pageNumber="1" pagination="1 - 125" refId="B12" refString="Emery, C, 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1 - 125, DOI: https://doi.org/10.5281/zenodo.54863" title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae." url="https://doi.org/10.5281/zenodo.54863" volume="118" year="1911">Emery 1911</bibRefCitation>
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: 99. Combination in
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<taxonomicName id="E2EADA468BACABCA89B8BEC45AF8EE92" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eburopone</taxonomicName>
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:
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<bibRefCitation id="EA9D4F453E58C3A0A005AB053E66F279" DOI="https://doi.org/10.3897/zookeys.608.9427" author="Borowiec, ML" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 280" refId="B6" refString="Borowiec, ML, 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280, DOI: https://doi.org/10.3897/zookeys.608.9427" title="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)." url="https://doi.org/10.3897/zookeys.608.9427" volume="608" year="2016">Borowiec 2016</bibRefCitation>
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: 129.
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</paragraph>
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<paragraph id="49264F4394CA0ADA383F4B01E04B20BB" pageId="0" pageNumber="1">
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<taxonomicName id="3DCCE91FC4A9279EE0D7E557AA4ABBC1" class="Insecta" family="Formicidae" genus="Cerapachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapachys wroughtoni subsp. var. var. rhodesiana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="wroughtoni" subSpecies="var." variety="rhodesiana">Cerapachys wroughtoni var. rhodesiana</taxonomicName>
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Forel, 1913: 212.
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<bibRefCitation id="10B91505C976FBDBE2DE1C0CA91D5933" author="Arnold, G" journalOrPublisher="Annals of the South African Museum, Cape Town" pageId="0" pageNumber="1" pagination="1 - 159" publicationUrl="https://biostor.org/reference/109829" refId="B2" refString="Arnold, G, 1915. A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae. Annals of the South African Museum, Cape Town 14: 1 - 159, https://biostor.org/reference/109829" title="A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae." url="https://biostor.org/reference/109829" volume="14" year="1915">Arnold 1915</bibRefCitation>
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: 16;
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<bibRefCitation id="B3E723A2C586AF7D8688A0F03256832F" author="Wheeler, WM" journalOrPublisher="Bulletin of the American Museum of Natural History, New York" pageId="0" pageNumber="1" publicationUrl="http://hdl.handle.net/2246/932" refId="B25" refString="Wheeler, WM, 1922. Ants of the American Museum Congo Expedition. Bulletin of the American Museum of Natural History, New York, http://hdl.handle.net/2246/932" title="Ants of the American Museum Congo Expedition." url="http://hdl.handle.net/2246/932" year="1922">Wheeler 1922</bibRefCitation>
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: 756. Junior synonym of
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<taxonomicName id="0F2346BE729F737EBD436C136F73C125" family="Formicidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. wroughtoni" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="wroughtoni">C. wroughtoni</taxonomicName>
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: Brown 1975: 24;
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<bibRefCitation id="FED6559B5B7DB55287F83E3CCF532F45" author="Bolton, B" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="0" pageNumber="1" refId="B3" refString="Bolton, B, 1995. A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts" title="A New General Catalogue of the Ants of the World." year="1995">Bolton 1995</bibRefCitation>
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: 144.
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</paragraph>
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<paragraph id="989EB294AAE3864EAC10CA87A53E49E1" pageId="0" pageNumber="1">
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<taxonomicName id="E87352AA27B567465A2589DBD2000989" class="Insecta" family="Formicidae" genus="Cerapachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapachys roberti" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roberti">Cerapachys roberti</taxonomicName>
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Forel, 1914: 212.
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<bibRefCitation id="554569E3182CD9C86CF6B223D9F29968" author="Wheeler, WM" journalOrPublisher="Bulletin of the American Museum of Natural History, New York" pageId="0" pageNumber="1" publicationUrl="http://hdl.handle.net/2246/932" refId="B25" refString="Wheeler, WM, 1922. Ants of the American Museum Congo Expedition. Bulletin of the American Museum of Natural History, New York, http://hdl.handle.net/2246/932" title="Ants of the American Museum Congo Expedition." url="http://hdl.handle.net/2246/932" year="1922">Wheeler 1922</bibRefCitation>
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: 756; Arnold 1926: 192. Junior synonym of
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<taxonomicName id="3E1E67562D90FB9053C2A5D210F8EC71" family="Formicidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. wroughtoni" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="wroughtoni">C. wroughtoni</taxonomicName>
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: Brown 1975: 24;
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<bibRefCitation id="49DC55A97DC48891F1E31DD025280246" author="Bolton, B" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="0" pageNumber="1" refId="B3" refString="Bolton, B, 1995. A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Massachusetts" title="A New General Catalogue of the Ants of the World." year="1995">Bolton 1995</bibRefCitation>
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: 144.
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</paragraph>
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</subSubSection>
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<subSubSection id="21CBD9D5A35C263F56318C2DA8EE07FF" pageId="0" pageNumber="1" type="type material">
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<paragraph id="91F6C2A5A6A0364B7CBA074AB76C6DCE" pageId="0" pageNumber="1">Type material images examined.</paragraph>
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<paragraph id="B2A4B64D9A8591FA385757B0D54935C9" pageId="0" pageNumber="1">
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The following type materials of
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<taxonomicName id="36430892AA403A5BD27622AE04C25616" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
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<emphasis id="D63B806035F4C3F0A5AF0EF4F015DD40" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
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</taxonomicName>
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and its two junior synonyms were non-physically examined based on the images available from AntWeb (https://www.antweb.org):
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<taxonomicName id="7CA2C8E8D4D7502AD947FBF19E2DC498" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
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<emphasis id="FB457D72A990B58CD7BBBA051BB38537" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
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</taxonomicName>
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(Forel, 1910), syntype, CASENT0249876 (MHNG);
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<taxonomicName id="25E371A754846A31FC87641683EEEC0A" lsidName="E. wroughtoni subsp. var. var. rhodesiana" pageId="0" pageNumber="1" rank="variety" species="wroughtoni" subSpecies="var." variety="rhodesiana">E. wroughtoni var. rhodesiana</taxonomicName>
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(Forel, 1913), syntype, CASENT0907055 (MHNG);
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<taxonomicName id="B1D49148B6F2ABE371CAB6A84BC437FA" lsidName="E. roberti" pageId="0" pageNumber="1" rank="species" species="roberti">
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<emphasis id="C18243BC8A92EB21B80EE5662E611221" italics="true" pageId="0" pageNumber="1">E. roberti</emphasis>
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</taxonomicName>
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(Forel, 1910), syntypes, CASENT0907054 (MHNG), CASENT0902707 (NHMUK).
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</paragraph>
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</subSubSection>
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<subSubSection id="44E4CAF3FD521107E1A3BC7A4592360C" pageId="0" pageNumber="1" type="notes">
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<paragraph id="DCEA60FDC4EC5D7938A99C12D14E731B" pageId="0" pageNumber="1">Notes.</paragraph>
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<paragraph id="0626F29D460ADBD547A5325D8415AEC8" pageId="0" pageNumber="1">
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Although Brown (1975) treated two southern African
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<taxonomicName id="1D678B975B904921F5B774BB48FD3F6E" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="8F515AF56216274A6DE5918061FAD7D4" italics="true" pageId="0" pageNumber="1">Eburopone</emphasis>
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|
</taxonomicName>
|
|
(sub)species,
|
|
<taxonomicName id="5A2556AFB60837573E3939E535554B92" lsidName="E. roberti" pageId="0" pageNumber="1" rank="species" species="roberti">
|
|
<emphasis id="39E110671D4E4F267E0D3E5E4D9A356C" italics="true" pageId="0" pageNumber="1">E. roberti</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="BDB7B3B4F7FB280787BBE512A1B44153" lsidName="E. w. subsp. rhodesiana" pageId="0" pageNumber="1" rank="subSpecies" species="w." subSpecies="rhodesiana">
|
|
<emphasis id="8C6B9B91135064D474682A89E15FF86C" italics="true" pageId="0" pageNumber="1">E. w. rhodesiana</emphasis>
|
|
</taxonomicName>
|
|
as junior synonyms of
|
|
<taxonomicName id="EAAF94B6105E2846BC00E02E644CB687" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
|
<emphasis id="6D9CE8694DA892E6824B20D77F15E375" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
|
</taxonomicName>
|
|
, the types of these names show conspicuous morphological differences, especially between
|
|
<taxonomicName id="9006C0110DD2F33886A052F388A35A5E" lsidName="E. w. subsp. rhodesiana" pageId="0" pageNumber="1" rank="subSpecies" species="w." subSpecies="rhodesiana">
|
|
<emphasis id="A247D3B17BA35BF29B22F6A6628DC183" italics="true" pageId="0" pageNumber="1">E. w. rhodesiana</emphasis>
|
|
</taxonomicName>
|
|
and the others.
|
|
<taxonomicName id="257C16CE3754844B745D73FFC23F3EEE" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone w. subsp. rhodesiana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="w." subSpecies="rhodesiana">
|
|
<emphasis id="0D4716AB234685D1607CD4D8EA7B19E0" italics="true" pageId="0" pageNumber="1">Eburopone w. rhodesiana</emphasis>
|
|
</taxonomicName>
|
|
is the most morphologically distinct among these by i) lateroventral flange of occipital carina less developed; ii) promesonotal suture inconspicuous (hardly recognizable in the examined image); iii) abdominal segment III distinctly wider than long.
|
|
<taxonomicName id="2586EF1A7D3FB61FF121F41A5CC1DAD0" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone roberti" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roberti">
|
|
<emphasis id="5F5711FC1490108FA3175A7B9DA80964" italics="true" pageId="0" pageNumber="1">Eburopone roberti</emphasis>
|
|
</taxonomicName>
|
|
is different from the syntype of
|
|
<taxonomicName id="06BD7511AF2E747BE49CC7BD528D44F0" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
|
<emphasis id="5D6BAE6E5FF692C4FC100C9D37D957FA" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
|
</taxonomicName>
|
|
at least by having deeper and more prominent promesonotal suture. The differences suggest a possible heterospecificity of these names, especially for
|
|
<taxonomicName id="61627C2A47721F64C1BF2B69347D2E82" lsidName="E. w. subsp. rhodesiana" pageId="0" pageNumber="1" rank="subSpecies" species="w." subSpecies="rhodesiana">
|
|
<emphasis id="8C115432F2CE9EFA3D26377E6CA42ADE" italics="true" pageId="0" pageNumber="1">E. w. rhodesiana</emphasis>
|
|
</taxonomicName>
|
|
. However, their taxonomic statuses are not addressed in the present study, since it would be appropriate to examine additional colony series; additionally, many undescribed species are known from the Afrotropical and Malagasy regions. Even if multiple species are confused under
|
|
<taxonomicName id="C84CC3710012A8BB8243B2A6ABEC029C" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
|
<emphasis id="F0328DEE8CC4A3A6D48E6DC2210E24AB" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
|
</taxonomicName>
|
|
, the independence of
|
|
<taxonomicName id="A06950838D7EE1675BC4E4CF87F8371C" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
|
<emphasis id="C4E760CACEA0E6C542FE8A8B60F676D9" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
|
</taxonomicName>
|
|
at the species level is unquestionable.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |