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<document id="D9F8781C8223FD5ACCAEB6AE1A3D910F" ID-DOI="10.5281/zenodo.185153" ID-GBIF-Dataset="0b5ad8cc-9b48-4700-acc5-012d1707f720" ID-ISSN="1175-5326" ID-Zenodo-Dep="185153" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1459966635824" checkinUser="plazi" docAuthor="Nihei, Silvio Shigueo &amp; Carvalho, Claudio José Barros De" docDate="2009" docId="E60987AB2403E1619686FE42FA96F306" docLanguage="en" docName="zt01976p024.pdf" docOrigin="Zootaxa 1976" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2814bb39" docStyleName="zootaxa.2007.monograph" docTitle="Morellia Nihei &amp; Carvalho, 2009, s.s." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="1A30FFD3240EE16C9611FFBDFF97F67C" masterDocTitle="The Muscini flies of the world (Diptera, Muscidae): identification key and generic diagnoses" masterLastPageNumber="24" masterPageNumber="1" pageNumber="14" updateTime="1698231170040" updateUser="plazi">
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<mods:title id="083135F1816A4DFE5F959CA87C98523F">The Muscini flies of the world (Diptera, Muscidae): identification key and generic diagnoses</mods:title>
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<mods:namePart id="FC3BF79C1D66B64F719626FE735EEAE7">Nihei, Silvio Shigueo</mods:namePart>
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<treatment id="E60987AB2403E1619686FE42FA96F306" ID-DOI="http://doi.org/10.5281/zenodo.5658499" ID-GBIF-Taxon="119377424" ID-Zenodo-Dep="5658499" LSID="urn:lsid:plazi:treatment:E60987AB2403E1619686FE42FA96F306" httpUri="http://treatment.plazi.org/id/E60987AB2403E1619686FE42FA96F306" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="26BA65362403E1619686FE42FE3CF465" box="[151,427,511,537]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="6E1F36BD2403E1619686FE42FE3CF465" blockId="13.[151,427,511,537]" box="[151,427,511,537]" pageId="13" pageNumber="14">
<heading id="355781D12403E1619686FE42FE3CF465" bold="true" box="[151,427,511,537]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="5CD4EAAF2403E1619686FE42FE3CF465" bold="true" box="[151,427,511,537]" pageId="13" pageNumber="14">
Subgenus
<taxonomicName id="A9A04D3E2403E1619706FE42FEE8F465" ID-CoL="5V64" ID-ENA="253340" box="[279,383,511,537]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="5CD4EAAF2403E1619706FE42FEE8F465" bold="true" box="[279,383,511,537]" italics="true" pageId="13" pageNumber="14">Morellia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47E757D42403E1619797FE42FE3CF465" box="[390,427,511,537]" pageId="13" pageNumber="14" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26BA65362403E1619686FDF7FE50F4FD" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="6E1F36BD2403E1619686FDF7FE50F4FD" blockId="13.[151,1436,586,641]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="4EB011932403E1619686FDF7FE50F4FD" pageId="13" pageNumber="14">
<taxonomicName id="A9A04D3E2403E1619686FDF7FDDCF41D" ID-CoL="5V64" authority="Robineau-Desvoidy, 1830: 405" authorityName="Robineau-Desvoidy" authorityPageNumber="405" authorityYear="1830" box="[151,587,586,609]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2403E1619686FDF7FF64F41D" box="[151,243,586,609]" italics="true" pageId="13" pageNumber="14">Morellia</emphasis>
<treatmentCitation id="EF0110AC2403E16196EAFDF7FDDCF41D" author="Robineau-Desvoidy" box="[251,587,586,609]" page="405" pageId="13" pageNumber="14" year="1830">
<bibRefCitation id="0A314B4C2403E16196EAFDF7FD80F41D" author="Robineau-Desvoidy" box="[251,535,586,609]" pageId="13" pageNumber="23" refString="Robineau-Desvoidy, J. B. (1830) Essai sur les Myodaires. Memoires presentes par divers savants a l'Academie des sciences de l'Institut de France, 2, 1 - 813." type="journal article" year="1830">Robineau-Desvoidy, 1830</bibRefCitation>
: 405
</treatmentCitation>
</taxonomicName>
(as genus).
<typeStatus id="B11B881F2403E16194C0FDF7FC91F41D" box="[721,774,586,609]" pageId="13" pageNumber="14">Type</typeStatus>
species:
<taxonomicName id="A9A04D3E2403E161957DFDF7FABBF41D" ID-CoL="44CY5" authority="Robineau-Desvoidy, 1830" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[876,1324,586,609]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="5CD4EAAF2403E161957DFDF7FB9CF41D" box="[876,1035,586,609]" italics="true" pageId="13" pageNumber="14">Morellia agilis</emphasis>
<bibRefCitation id="0A314B4C2403E1619205FDF7FABBF41D" author="Robineau-Desvoidy" box="[1044,1324,586,609]" pageId="13" pageNumber="23" refString="Robineau-Desvoidy, J. B. (1830) Essai sur les Myodaires. Memoires presentes par divers savants a l'Academie des sciences de l'Institut de France, 2, 1 - 813." type="journal article" year="1830">Robineau-Desvoidy, 1830</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="A9A04D3E2403E1619344FDF7FE2AF4FD" authority="Fallen, 1817" authorityName="Fallen" authorityYear="1817" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hortorum">
<emphasis id="5CD4EAAF2403E1619344FDF7FEBBF4FD" italics="true" pageId="13" pageNumber="14">Musca hortorum</emphasis>
Fallén, 1817
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="26BA65362403E1619686FD0FFB10F5BE" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="6E1F36BD2403E1619686FD0FFB10F5BE" blockId="13.[151,1437,690,1402]" pageId="13" pageNumber="14">
<emphasis id="5CD4EAAF2403E1619686FD0FFE80F4B0" bold="true" box="[151,279,690,716]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Colouration metallic bluish-black to violaceous; wing without maculae, but with maculae in several Neotropical species. Female with proclinate fronto-orbital usually developed; parafrons setulose on upper half; interfrontal seta absent. Postsutural intra-alars 1; intrapostalar present. Prosternum setulose or bare. Katepisternals 1+2. Wing with the basal portion of stem-vein setulose ventrally (bare in
<taxonomicName id="A9A04D3E2403E16192A2FC97FA91F53F" box="[1203,1286,810,835]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="5CD4EAAF2403E16192A2FC97FA91F53F" box="[1203,1286,810,835]" italics="true" pageId="13" pageNumber="14">basalis</emphasis>
</taxonomicName>
-group); R4+5 setulose dorsally and ventrally; M bent forward towards R4+5 (
<figureCitation id="F69B2A382403E161956CFCE8FC42F513" box="[893,981,853,879]" captionStart="FIGURES 9 11. 9" captionStartId="6.[151,269,1970,1994]" captionTargetBox="[311,1269,204,1933]" captionTargetId="figure@6.[311,1269,194,1942]" captionTargetPageId="6" captionText="FIGURES 9 11. 9. Schematic presentation of Muscidae thorax, dorsal view. 10. Thorax (partial), Neomyia cornicina (Fabricius), lateral view. 11. Wing, Morellia podagrica (Loew). Figures 9 11 after Nihei and de Carvalho (2007 a)." httpUri="https://zenodo.org/record/185155/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
). Lower calypter enlarged posteriorly, extending under base of scutellum. Subcostal sclerite bare. Male mid femur of some species with a dorsal preapical protuberance bearing a dense set of setulae (
<figureCitation id="F69B2A382403E1619501FC15FCF0F5BE" box="[784,871,936,962]" captionStart="FIGURES 15 18" captionStartId="8.[151,269,1930,1954]" captionTargetBox="[217,1392,211,1878]" captionTargetId="figure@8.[194,1399,194,1902]" captionTargetPageId="8" captionText="FIGURES 15 18. Apex of mid femur and tibia: 15. Pyrellina distinct (Walker), female, posterior view. 16. Biopyrellia bipuncta, male, posterior view. 17. Morellia (Parapyrellia) maculipennis (Macquart), male, anterior view. 18. Morellia podagrica, male, posterior view. Figures 15 18 after Nihei and de Carvalho (2007 a). All figures on same scale." httpUri="https://zenodo.org/record/185157/files/figure.png" pageId="13" pageNumber="14">Fig. 18</figureCitation>
). Calcar weak or strong.
</paragraph>
</subSubSection>
<subSubSection id="26BA65362403E16196D4FC6DFE01F2CE" pageId="13" pageNumber="14" type="discussion">
<paragraph id="6E1F36BD2403E16196D4FC6DFD35F246" blockId="13.[151,1437,690,1402]" pageId="13" pageNumber="14">
<emphasis id="5CD4EAAF2403E16196D4FC6DFEC3F596" bold="true" box="[197,340,976,1002]" pageId="13" pageNumber="14">Comments.</emphasis>
The monophyly of
<taxonomicName id="A9A04D3E2403E161945AFC6CFD27F596" box="[587,688,977,1002]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="5CD4EAAF2403E161945AFC6CFD27F596" box="[587,688,977,1002]" italics="true" pageId="13" pageNumber="14">Morellia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47E757D42403E16194AAFC6DFD76F596" box="[699,737,976,1002]" pageId="13" pageNumber="14" sensu="stricto">s.s.</taxonomicNameLabel>
was not supported by Nihei and de
<bibRefCitation id="0A314B4C2403E16192B2FC6DFAEFF596" author="Carvalho" box="[1187,1400,976,1002]" pageId="13" pageNumber="22" refString="Nihei, S. S. &amp; de Carvalho, C. J. B. (2007 a) Phylogeny and classification of Muscini (Diptera, Muscidae). Zoological Journal of the Linnean Society, 149, 493 - 532." type="journal article" year="2007" yearSuffix="a">Carvalho (2007a)</bibRefCitation>
. A comprehensive phylogenetic study of
<taxonomicName id="A9A04D3E2403E161944CFC44FD54F26E" box="[605,707,1017,1042]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="5CD4EAAF2403E161944CFC44FD54F26E" box="[605,707,1017,1042]" italics="true" pageId="13" pageNumber="14">Morellia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47E757D42403E16194DDFC45FD79F26E" box="[716,750,1016,1042]" pageId="13" pageNumber="14" sensu="lato">s.l.</taxonomicNameLabel>
is required to clarify the composition of
<taxonomicName id="A9A04D3E2403E16192CFFC44FAD3F26E" box="[1246,1348,1017,1042]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="5CD4EAAF2403E16192CFFC44FAD3F26E" box="[1246,1348,1017,1042]" italics="true" pageId="13" pageNumber="14">Morellia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47E757D42403E161935CFC45FAE3F26E" box="[1357,1396,1016,1042]" pageId="13" pageNumber="14" sensu="stricto">s.s.</taxonomicNameLabel>
, as well as its relationship with other subgenera.
</paragraph>
<paragraph id="6E1F36BD2403E16196D4FBF5FE01F2CE" blockId="13.[151,1437,690,1402]" pageId="13" pageNumber="14">
Note on
<taxonomicName id="A9A04D3E2403E1619725FBF5FE10F21D" box="[308,391,1096,1121]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="5CD4EAAF2403E1619725FBF5FE10F21D" box="[308,391,1096,1121]" italics="true" pageId="13" pageNumber="14">basalis</emphasis>
</taxonomicName>
-group: This species-group was indicated as monophyletic by Nihei and de
<bibRefCitation id="0A314B4C2403E1619320FBF5FF65F2F6" author="Carvalho" pageId="13" pageNumber="22" refString="Nihei, S. S. &amp; de Carvalho, C. J. B. (2007 a) Phylogeny and classification of Muscini (Diptera, Muscidae). Zoological Journal of the Linnean Society, 149, 493 - 532." type="journal article" year="2007" yearSuffix="a">Carvalho (2007a)</bibRefCitation>
and is composed so far of two Neotropical species, namely,
<taxonomicName id="A9A04D3E2403E16195D5FBCDFB39F2F6" authority="Walker" authorityName="Walker" box="[964,1198,1136,1162]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="5CD4EAAF2403E16195D5FBCDFBA8F2F5" box="[964,1087,1136,1161]" italics="true" pageId="13" pageNumber="14">M. basalis</emphasis>
(Walker)
</taxonomicName>
and
<taxonomicName id="A9A04D3E2403E16192F8FBCDFE01F2CE" authority="Pamplona &amp; Mendes." authorityName="Pamplona &amp; Mendes." class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="paulistensis">
<emphasis id="5CD4EAAF2403E16192F8FBCDFA0BF2F5" box="[1257,1436,1136,1161]" italics="true" pageId="13" pageNumber="14">M. paulistensis</emphasis>
Pamplona &amp; Mendes.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="26BA65362403E16196D4FB7DFB52F2A6" box="[197,1221,1216,1242]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="6E1F36BD2403E16196D4FB7DFB52F2A6" blockId="13.[151,1437,690,1402]" box="[197,1221,1216,1242]" pageId="13" pageNumber="14">
<emphasis id="5CD4EAAF2403E16196D4FB7DFEC9F2A6" bold="true" box="[197,350,1216,1242]" pageId="13" pageNumber="14">Distribution</emphasis>
(50 species). Afrotropical, Nearctic, Neotropical, Oriental and Palaearctic
</paragraph>
</subSubSection>
<subSubSection id="26BA65362403E16196D4FB55FA96F306" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="6E1F36BD2403E16196D4FB55FA96F306" blockId="13.[151,1437,690,1402]" pageId="13" pageNumber="14">
<emphasis id="5CD4EAAF2403E16196D4FB55FEC2F37E" bold="true" box="[197,341,1256,1282]" pageId="13" pageNumber="14">References.</emphasis>
Afrotropical species:
<bibRefCitation id="0A314B4C2403E1619477FB55FD6AF37E" author="Peris" box="[614,765,1256,1282]" pageId="13" pageNumber="23" refString="Peris, S. V. (1961) Una nueva especie de Morellia de Camarones y synopsis de las especies etiopicas (Dipt. Muscidae). Eos, 37, 349 - 359." type="journal article" year="1961">Peris (1961)</bibRefCitation>
,
<bibRefCitation id="0A314B4C2403E161951FFB55FC22F37E" author="Zielke" box="[782,949,1256,1282]" pageId="13" pageNumber="24" refString="Zielke, E. (1971) Revision der Muscinae der athiopischen Region. Series Entomologica, 7, 1 - 199." type="journal article" year="1971">Zielke (1971)</bibRefCitation>
,
<bibRefCitation id="0A314B4C2403E16195D7FB55FB27F37E" box="[966,1200,1256,1282]" pageId="13" pageNumber="21" refString="Couri, M. S., Pont, A. C. &amp; Penny, N. D. (2006) Muscidae (Diptera) from Madagascar: identification keys, descriptions of new species, and new records. Proceedings of the California Academy of Sciences, 57, 799 - 923." type="journal article">Couri et al. (2006)</bibRefCitation>
; Australasian:
<bibRefCitation id="0A314B4C2403E1619377FB55FF7FF356" author="Pont" pageId="13" pageNumber="23" refString="Pont, A. C. (1973) Studies on Australian Muscidae (Diptera). IV. A revision of the subfamilies Muscinae and Stomoxyinae. Australian Journal of Zoology (Supplementary Series), 21, 129 - 296." type="journal article" year="1973">Pont (1973)</bibRefCitation>
; Nearctic:
<bibRefCitation id="0A314B4C2403E161977AFAADFDB4F356" author="Huckett" box="[363,547,1296,1322]" pageId="13" pageNumber="22" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1 - 370." type="journal article" year="1965">Huckett (1965)</bibRefCitation>
; Neotropical:
<bibRefCitation id="0A314B4C2403E16194DCFAADFB8AF356" author="Pamplona" box="[717,1053,1296,1322]" pageId="13" pageNumber="22" refString="Pamplona, D. M. &amp; Couri, M. S. (1995) Morellia dendropanacis, a new species, and other species with spotted wings: characterization and comparison (Diptera: Muscidae: Muscinae). Proceedings of the Biological Society of Washington, 108, 451 - 457." type="journal article" year="1995">Pamplona and Couri (1995)</bibRefCitation>
, de
<bibRefCitation id="0A314B4C2403E1619240FAADFA0FF356" author="Carvalho" box="[1105,1432,1296,1322]" pageId="13" pageNumber="21" refString="de Carvalho, C. J. B. &amp; Couri, M. S. (2002) Part I. Basal groups. In: de Carvalho, C. J. B. (Ed.), Muscidae (Diptera) of the Neotropical Region: taxonomy. Editora Universidade Federal do Parana, Curitiba, pp. 17 - 132." type="book chapter" year="2002">Carvalho and Couri (2002)</bibRefCitation>
; Oriental:
<bibRefCitation id="0A314B4C2403E1619717FA85FE27F32E" author="Emden" box="[262,432,1336,1362]" pageId="13" pageNumber="21" refString="Emden, F. I. van (1965) Diptera. Vol. 7. Muscidae, Part 1. In: Sewell, R. B. S. &amp; Roonwal, M. L. (Eds), The Fauna of India and the adjacent countries. Zoological Survey of India, Calcutta, 647 pp." type="book" year="1965">Emden (1965)</bibRefCitation>
,
<bibRefCitation id="0A314B4C2403E16197ADFA85FD2DF32E" author="Xue" box="[444,698,1336,1362]" pageId="13" pageNumber="24" refString="Xue, W. &amp; Chao, C., eds. (1998) Flies of China. Vol. 1. Liaoning Science and Technology Press, Shenyang, 1365 pp." type="book" year="1998">Xue and Chao (1998)</bibRefCitation>
; Palaearctic:
<bibRefCitation id="0A314B4C2403E1619544FA85FC63F32E" author="Zimin" box="[853,1012,1336,1362]" pageId="13" pageNumber="24" refString="Zimin, L. S. (1951) Family Muscidae (Tribes Muscini and Stomoxydini). Fauna USSR (n. s.), 45, 1 - 286." type="journal article" year="1951">Zimin (1951)</bibRefCitation>
,
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</paragraph>
</subSubSection>
</treatment>
</document>