135 lines
11 KiB
XML
135 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.72.744" ID-GBIF-Dataset="9b029a3f-f31b-421f-a62e-fb4dd8ff6731" ID-PMC="PMC3083004" ID-Pensoft-Pub="1313-2970-72-23" ID-PubMed="21594103" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-72-23" ModsDocOrigin="ZooKeys 72" ModsDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451250861919" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak" docDate="2010" docId="E3E602972AA21466696255867FC27CDA" docLanguage="en" docName="ZooKeys 72: 23-68" docOrigin="ZooKeys 72" docSource="http://dx.doi.org/10.3897/zookeys.72.744" docTitle="Tylopus extremus Likhitrakarn, Golovatch, Prateepasen & Panha, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="30" masterDocId="D7146C3215665021FFDFFFACFF8CFFD0" masterDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="68" masterPageNumber="23" pageNumber="28" updateTime="1668165214761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prateepasen, Rujiporn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>72</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.72.744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.72.744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-72-23</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360927" LSID="urn:lsid:zoobank.org:act:188F2E10-CAC7-406A-B176-6818DF526D0B" httpUri="http://treatment.plazi.org/id/E3E602972AA21466696255867FC27CDA" lastPageId="7" lastPageNumber="30" pageId="5" pageNumber="28">
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<subSubSection pageId="5" pageNumber="28" type="nomenclature">
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<paragraph pageId="5" pageNumber="28">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:188F2E10-CAC7-406A-B176-6818DF526D0B" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus extremus" order="Polydesmida" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="extremus">Tylopus extremus</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="28">sp. n.</taxonomicNameLabel>
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Figs 79
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="holotype">
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<paragraph pageId="5" pageNumber="28">Holotype</paragraph>
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<paragraph pageId="5" pageNumber="28">♂ (CUMZ), Thailand, Chiang Mai Province, Fang District, Doi Phahom Pok National Park, 6.07.2009, leg. A. Pansook.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="paratypes">
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<paragraph pageId="5" pageNumber="28">Paratypes:</paragraph>
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<paragraph pageId="5" pageNumber="28">1 ♂, 1 ♀ (CUMZ), 1 ♂ (ZMUC), 1 ♂ (ZMUM), same locality, together with holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="name">
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<paragraph pageId="5" pageNumber="28">Name:</paragraph>
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<paragraph pageId="5" pageNumber="28">To emphasize the extremely long spines h and m of the gonopod.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="diagnosis">
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<paragraph pageId="5" pageNumber="28">Diagnosis:</paragraph>
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<paragraph pageId="5" pageNumber="28">Differs from congeners in process h being subflagelliform while process m extremely long and prominent.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="30" pageId="5" pageNumber="28" type="description">
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<paragraph pageId="5" pageNumber="28">Description:</paragraph>
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<paragraph lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28">
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Length ca 30 mm (holotype), 27-30 mm (♂), 32.5 mm (♀), width of midbody pro- and metazona 2.0 and 2.9 mm (holotype), 1.9-2.4 and 2.8-3.3 mm (♂), 2.5 and 3.0 mm (♀), respectively. Coloration of live animals, as well as of alcohol material black-brown (Fig. 7
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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): calluses of paraterga and antennae only slightly
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<pageBreakToken pageId="6" pageNumber="29" start="start">lighter</pageBreakToken>
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, light brown to brown, venter and legs contrastingly light, yellow (Fig. 7
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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), tip of antennae pallid.
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</paragraph>
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<paragraph pageId="6" pageNumber="29">
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All characters as in
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., except as follows.
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</paragraph>
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<paragraph pageId="6" pageNumber="29">Antennae rather short and slender, reaching behind to end of segment 3 (♂) dorsally. Collum with paraterga like rudimentary flaps, especially poorly developed in ♀. In width, head <collum = segments 3-4 <2 <5-16 (♂) (Fig. 7B), or head <collum <segment 3 <2 and 4 <5-16(♀); thereafter body gradually and gently tapering towards telson.</paragraph>
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<paragraph pageId="7" pageNumber="30">
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<pageBreakToken pageId="7" pageNumber="30" start="start">Metaterga</pageBreakToken>
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with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points on low longitudinal ridges or tubercles (Fig. 7
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<normalizedToken originalValue="B–G">B-G</normalizedToken>
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). Axial line thin, visible on both halves of metaterga. Paraterga strongly developed (Fig. 7
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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), lying rather low (at 1/2-1/3 midbody height), slightly inclined laterally, pointed caudally and acutangular already from segment 2, especially strongly so on caudal segments, very clearly surpassing rear contour only on segments 16-19; anterior 1/3 of poreless calluses with two barely visible, lateral, setigerous incisions, but with only a single, likewise poorly developed incision anteriorly on pore-bearing calluses (Fig. 7
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<normalizedToken originalValue="B–F">B-F</normalizedToken>
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); paraterga slightly less strongly developed in ♀. Transverse sulcus evident on metaterga 5-18, reaching bases of paraterga, evident and rather deep, faintly rugulose at bottom. Stricture between pro- and metazona weakly striolate (Fig. 7
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<normalizedToken originalValue="B–G">B-G</normalizedToken>
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). Epiproct emarginate at tip, pre-apical papillae evident (Fig. 7G). Hypoproct subtrapeziform, caudal setae widely separated (Fig. 7H). Pleurosternal carinae as compete ridges with a caudal tooth on segments 2-4 (♂) or 2 and 3 (♀), like separated anterior bulges and increasingly poorly developed caudal denticles until segment 16 (Fig. 7C, E, F).
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</paragraph>
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<paragraph pageId="7" pageNumber="30">Sterna moderately setose, without modifications; an entire, linguiform, sternal lobe between ♂ coxae 4 (Fig. 7I, J). Legs long, in ♂ very distinctly incrassate, 1.7-2.0 or ca 1.3 times as long as midbody height in ♂ and ♀, respectively (Fig. 7B, C, H), ♂ prefemora very distinctly bulged laterally and clothed with dense and adpressed pilosity ventrally (Fig. 7K), acropodites also with similarly dense and ventrally adpressed pilosity, including tarsal brushes. All ♂ postfemora and tibiae except for a few posteriormost ones with a small, but evident adenostyle (= tubercle) at midway on ventral side (Fig. 7K).</paragraph>
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<paragraph pageId="7" pageNumber="30">Gonopods (Figs 8, 9) with lobe l well-demarcated, but unusually prominent, high and elongated; spine h long, extremely slender and subflagelliform; spine z rather short and simple; spine m very prominent, straight and long.</paragraph>
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<caption pageId="7" pageNumber="30">
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<paragraph pageId="7" pageNumber="30">
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Figure 7.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus extremus" order="Polydesmida" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="extremus">Tylopus extremus</taxonomicName>
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sp. n., ♂ paratype (
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<normalizedToken originalValue="A–K">A-K</normalizedToken>
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). A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively K midbody leg.
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</paragraph>
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</caption>
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<caption pageId="7" pageNumber="30">
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<paragraph pageId="7" pageNumber="30">
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Figure 8.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus extremus" order="Polydesmida" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="extremus">Tylopus extremus</taxonomicName>
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sp. n., ♂ paratype. A, B right gonopod, mesal and lateral views, respectively
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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distal part of right gonopod, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="7" pageNumber="30">
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<paragraph pageId="7" pageNumber="30">
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Figure 9.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus extremus" order="Polydesmida" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="extremus">Tylopus extremus</taxonomicName>
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sp. n., ♂ paratype. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |