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<document id="BDC5902DAE8958AFE7D6207344DB056C" ID-DOI="10.3897/zookeys.1218.134413" ID-ZooBank="D3F484A9-2FFB-46D3-AC9B-D43223DABE68" ID-publisher-id="134413" URI-arpha="04530E12-6D54-5F8E-9555-CE9BDFB010C0" URI-zoobank="http://zoobank.org/D3F484A9-2FFB-46D3-AC9B-D43223DABE68" article-type="research-article" checkinTime="1732212372937" checkinUser="pensoft" docAuthor="Telnov, Dmitry" docDate="2024" docId="CA80F68FD59F587592FDC5710B3F9323" docLanguage="en" docName="ZooKeys 1218: 231-250" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/134413/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Macrotomoderus hirsutus Telnov, 2024, sp. nov." docType="treatment" docUuid="1D16E28A-F43F-40E5-88C6-82CCAB013AB1" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="250" masterDocId="04530E126D545F8E9555CE9BDFB010C0" masterDocTitle="Revision of the Tomoderinae (Coleoptera, Anthicidae). Part V. Three new Macrotomoderus Pic, 1901 from continental China and an updated key to the Palaearctic species" masterLastPageNumber="250" masterPageNumber="231" pageNumber="231" updateTime="1732212759614" updateUser="ExternalLinkService">
<mods:mods id="E3755D15B0E1B3FD82D8A0FC2BFEE76E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A5314FD696691D8A862AD8ECEF571F95">Revision of the Tomoderinae (Coleoptera, Anthicidae). Part V. Three new Macrotomoderus Pic, 1901 from continental China and an updated key to the Palaearctic species</mods:title>
</mods:titleInfo>
<mods:name id="A2F36699E4BCD5976133759509D6D5B0" type="personal">
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<mods:namePart id="E545215EE5E8FA57D83A4A0ADBF9091F">Telnov, Dmitry</mods:namePart>
<mods:nameIdentifier id="1D6AE8EA4EAD8236A298E141AF031C6E" type="ORCID">0000-0003-3412-0089</mods:nameIdentifier>
<mods:affiliation id="66C2168359C3AC3A3304F974685A3846">Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV 5401, Daugavpils, Latvia &amp; Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, UK &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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<mods:typeOfResource id="73028C295A93EBDD6BC5E2FAEC30677C">text</mods:typeOfResource>
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<mods:title id="8B1ADF9AB7253D7CF8797AAE5057F040">ZooKeys</mods:title>
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<mods:part id="4921A173C05E98A4CF1C99A7ACDA3B98">
<mods:date id="B824394FFE265A1B9EBA0CA88A84DF18">2024</mods:date>
<mods:detail id="5BAFDDEF5A27492B7B2C4677B411C3CA" type="pubDate">
<mods:number id="57C64782B562DDC2736C6191A1522BD4">2024-11-21</mods:number>
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<mods:detail id="84C9999F6E55FE9730A668000687C7F1" type="volume">
<mods:number id="2609B405D53957C97170FD1F4B3B8643">1218</mods:number>
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<mods:classification id="F4BDD69A0E83B3A5FF2E6B8A68111809">journal article</mods:classification>
<mods:identifier id="0C2890A8F2E69EC858C4302A62381138" type="DOI">10.3897/zookeys.1218.134413</mods:identifier>
<mods:identifier id="93BA2B86824530E64E1D137BCEDDA5D8" type="ZooBank">D3F484A9-2FFB-46D3-AC9B-D43223DABE68</mods:identifier>
</mods:mods>
<treatment id="CA80F68FD59F587592FDC5710B3F9323" ID-DOI="http://doi.org/10.5281/zenodo.14199417" ID-Zenodo-Dep="14199417" ID-arpha="CA80F68F-D59F-5875-92FD-C5710B3F9323" ID-zoobank="https://zoobank.org/1D16E28A-F43F-40E5-88C6-82CCAB013AB1" LSID="urn:lsid:zoobank.org:act:1D16E28A-F43F-40E5-88C6-82CCAB013AB1" LSID-ZBK="urn:lsid:zoobank.org:act:1D16E28A-F43F-40E5-88C6-82CCAB013AB1" httpUri="http://treatment.plazi.org/id/CA80F68FD59F587592FDC5710B3F9323">
<subSubSection id="05D647887F9D85EFC0B13AC9FC3EAB65" type="nomenclature">
<paragraph id="C62B047FB177F9A7C8E128227200C087">
<taxonomicName id="C904F6DB1D2D85928D86F6B556A65797" authorityName="Telnov" authorityYear="2024" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="3CF8D48B9DA18E50E1D03F1144F8B24E" italics="true">Macrotomoderus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7798F50AB0271B90A19E2CDAE23E8718" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5EEAB6DEA2B29EA774D267BBEF14E888" type="description">
<paragraph id="8585EE49763A680C9DA81064D220DE35">
<figureCitation id="42F00535911063611CCDE26EAB8C6DED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.134413.figure3" httpUri="https://binary.pensoft.net/fig/1180020">Figs 3</figureCitation>
,
<figureCitation id="8F7CB751F315D8FE808EE5C805B75604" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrotomoderus hirsutus sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, median portion, endophallic armature E ditto, basal portion, endophallic armature F ditto, endophallic armature, lateral view of basal portion G ditto, ventral view of apex [not to scale]." figureDoi="10.3897/zookeys.1218.134413.figure4" httpUri="https://binary.pensoft.net/fig/1180021">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EODAE" type="type material">
<paragraph id="324043F399FE7ECD2BCAAC27003890C6">
<heading id="2B862CEF5E805A6284053652A2AA5F70" reason="title">Type material designated.</heading>
</paragraph>
<paragraph id="A505F2F799243B47421F6625FC6388B0">
<materialsCitation id="92A7725EB5EF025B76E1B76C0A921773" collectingDate="2002-07-16" collectionCode="NME" collectorName="Stary" country="China" elevation="1900" location="Dabashan" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="58CB39627E0208916DFB61DBA9C7A6CC" bold="true" italics="true">
<typeStatus id="9562A96B2095E8BBDFCABA9AD160639C" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="39B8BD3FC967C6E733907C73C26BFB9A" count="1" type="male"></specimenCount>
<named-content id="0BF25D0E18B5BA19A7DB353069721901" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturkundemuseum-erfurt" xlink_title="Naturkundemuseum Erfurt, Erfurt, Germany">
<collectionCode id="92D9CC5A795098AD6AEE06027B2490E7" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" type="">NME</collectionCode>
</named-content>
: “
<collectingCountry id="24A10663032100FC522A7BE769E88784" name="China">CHINA</collectingCountry>
:
<collectingRegion id="B3267A0D0576DD3C0AC588DCCBAD3E83" country="China" name="Hubei">Hubei</collectingRegion>
:
<location id="4549604B2F32B5B11EC6867A46918B9E" LSID="urn:lsid:plazi:treatment:CA80F68FD59F587592FDC5710B3F9323:4549604B2F32B5B11EC6867A46918B9E" country="China" name="Dabashan" stateProvince="Hubei">Dabashan</location>
;
<locationDeviation id="ED537491BD04970B706C01FA6FBA8187" location="Muyuping">
<quantity id="669306ECC349D2BC8462ECD994A26A00" metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
NW Muyuping
</locationDeviation>
<quantity id="E8AA0ECCF36E0A9409F3D5B794C35966" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">
<elevation id="2D2EAE33C0A805FA4E5353CD9423A311" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</elevation>
</quantity>
leg.
<collectorName id="16D68BCF3A0FF6D437455B4F4CBB3756">Stary</collectorName>
<date id="AB505F6EA47E2A7AC8C141EB186C7067" value="2002-07-16">
<collectingDate id="A91E7FD0CCC20CD3A3C3126F77E1B22F" value="2002-07-16">16. 7. 2002</collectingDate>
</date>
” [printed].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5DAE" type="description">
<paragraph id="22B5E6A55FF40D16A8544474D88E0FFD">
<heading id="6603D01CC58D82984720C82DD2F06015" reason="title">Measurements.</heading>
</paragraph>
<paragraph id="08A252F8F04337143A367725C04797B5">
<typeStatus id="CAC0B22706C6BB3B62CECC617F6D5BBA" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="0E267F1DB28C612AA8A5E1C0750DA613" metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
; head including exposed part of cranial neck
<quantity id="1129958E9E1123B4A1E4982052A62E24" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
long, across eyes
<quantity id="EDDF23BEE0BFD68D33BD80900891F463" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
wide, pronotum
<quantity id="53FF820F46213B25333B649936A413EC" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
long, maximum width
<quantity id="C9CBF45743791A65E7A6ACBFA2310FA9" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
, minimum width
<quantity id="2DB50F8A10641BF42B94729AAC16D958" metricMagnitude="-4" metricUnit="m" metricValue="3.5" unit="mm" value="0.35">0.35 mm</quantity>
, elytra
<quantity id="15B9984E9B41D0618F72797B5450D60B" metricMagnitude="-3" metricUnit="m" metricValue="2.3" unit="mm" value="2.3">2.3 mm</quantity>
long,
<quantity id="0EAE49C9FFF98AAFE718BCD6D61ABCB3" metricMagnitude="-3" metricUnit="m" metricValue="1.7" unit="mm" value="1.7">1.7 mm</quantity>
combined wide.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDEAE" type="description">
<paragraph id="86D841B9DF446884E9A6E43B83DB5856">
<heading id="809E34928CC85AED0EA7F613369E86C5" reason="title">Description.</heading>
</paragraph>
<paragraph id="718B4EBB1293D6AC7BDDFFE0920E4524">
<typeStatus id="ABFC469FE910B14DADE9F37E5A0D79A6" type="holotype">Holotype</typeStatus>
, male. Head and pronotum brown, elytra pale brown. Mouthparts, antennae, palps, and legs pale brownish testaceous. Head ovoid, moderately glossy dorsally and ventrally, with moderate, nearly circular compound eyes which are slightly protruding beyond lateral outline of head. Head rounded in broad arc posterior to eyes. Head dorsal punctures minute but rather deep, denser on occiput and vertex. Intervening spaces 24 × as wide as punctures. Head dorsal setae yellowish, inconspicuous on most of head dorsum but long and dense, forming a subconical bristle (in dorsal view) at head base and therefore concealing median part of anterior pronotal margin. Apical portions of these long head base setae are curled and in part tangled (Fig.
<figureCitation id="39156AD79FEFD92C86F0F5DD1B02CC16" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.134413.figure3" httpUri="https://binary.pensoft.net/fig/1180020">3 B, C</figureCitation>
). Antenna exceeds slightly beyond base of elytra when directed posteriad. Antennomere 3 subequal in length to antennomere 2, antennomeres 610 transverse, of which 810 strongly so. Terminal antennomere asymmetrically triangular with rounded apex, ~ 1.6 × as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum stout, moderately glossy dorsally and laterally, approximately as wide as head across compound eyes, with broad, medially strongly notched (in dorsal view) postmedian lateral constriction. Anterior lobe much wider than posterior, its front margin broadly rounded, dorsally without modifications and anterior rim. Anterior lobe nearly flat in lateral view (Fig.
<figureCitation id="6BE0357E4796C9A64221F6F59128A2EE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.134413.figure3" httpUri="https://binary.pensoft.net/fig/1180020">3 B</figureCitation>
). Lateral constriction does not continue onto disc in lateral view (Fig.
<figureCitation id="F64B997B631427D550351AAD37D49965" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.134413.figure3" httpUri="https://binary.pensoft.net/fig/1180020">3 B</figureCitation>
). Lateral pronotal fovea wide at lower (lateroventral) extent of laterally strongly declivous pronotal disc, not widens upwards towards pronotal disc in lateral view, lateral edges of fovea carinate, moderately separated and densely setose (all in lateral view), in dorsal view denticle-like. In lateral view anterior and posterior edge of pronotal fovea completely covered with bristle of short golden setae. Cavity in lateral wall of pronotum moderate. In dorsal view, lateral pronotal fovea wide, anterior and posterior denticle clearly visible, dense brush-like setose (Fig.
<figureCitation id="C6CB1176336CE0A81F78C1F900355CC3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1218.134413.figure3" httpUri="https://binary.pensoft.net/fig/1180020">3 A, C</figureCitation>
). Pronotal punctures on disc similar to those on head dorsum, intervening spaces 37 × as wide as punctures; lateral constriction dorsally with irregularly shaped and variably dense and deep, generally moderately to strongly elongated punctures; median ones particularly large and elongate, groove-like. Dorsal pronotal setae moderately dense, but much denser on lateral sides of pronotum, especially in constriction area where setation effectively conceals shape of lateral pronotal denticles. Several longer erect tactile setae on lateral sides of anterior lobe. Scutellar shield minute, apically rounded, glabrous, and glossy. Elytra moderately glossy, dorsally elongate elliptical, slightly convex in lateral view, widened laterally around midlength, lateral margins broadly rounded, humerus obsolete (apterous species). Elytral punctures stronger and larger than those on dorsal forebody, more or less regularly circular, smaller than dorsal punctures on lateral pronotal constriction. Punctures becoming more flat and somewhat sparser on apical third of elytra. Intervening spaces ~ 35 × as wide as diameter of punctures. Elytral setae long and sparse, suberect, yellowish. Male tergite and sternite VII broadly rounded at posterior margin. Sternite IX rod like, slightly bisinuate (Fig.
<figureCitation id="8D3E60AA9F20FEA44A960419D8025B4D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrotomoderus hirsutus sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, median portion, endophallic armature E ditto, basal portion, endophallic armature F ditto, endophallic armature, lateral view of basal portion G ditto, ventral view of apex [not to scale]." figureDoi="10.3897/zookeys.1218.134413.figure4" httpUri="https://binary.pensoft.net/fig/1180021">4 A</figureCitation>
). Aedeagus as in Fig.
<figureCitation id="7D4004503BFAD166A6940C2B5F9B63D6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrotomoderus hirsutus sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, median portion, endophallic armature E ditto, basal portion, endophallic armature F ditto, endophallic armature, lateral view of basal portion G ditto, ventral view of apex [not to scale]." figureDoi="10.3897/zookeys.1218.134413.figure4" httpUri="https://binary.pensoft.net/fig/1180021">4 B G</figureCitation>
, elongate, narrows apically, apex somewhat subspatulate, widened and rounded. Endophallic armature of numerous large peculiarly spike / nail-shaped spines (Fig.
<figureCitation id="AB6E471F06369A7802621BBEEBF7C161" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrotomoderus hirsutus sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, median portion, endophallic armature E ditto, basal portion, endophallic armature F ditto, endophallic armature, lateral view of basal portion G ditto, ventral view of apex [not to scale]." figureDoi="10.3897/zookeys.1218.134413.figure4" httpUri="https://binary.pensoft.net/fig/1180021">4 B F</figureCitation>
).
</paragraph>
<caption id="7CDB1E99D922E6350B1A684CA70A4BF2" ID-DOI="10.3897/zookeys.1218.134413.figure3" ID-arpha="DC55F809-C8D1-5774-BBAE-7CA57FEB4604" httpUri="https://binary.pensoft.net/fig/1180020" startId="F3">
<paragraph id="AEF477C33DA47E711A94038DEA81876A">
<label id="281C205E9E1587879CAC4B6CF81CB44C">Figure 3.</label>
</paragraph>
<paragraph id="9480382ECBB73E102DF9D079EDC7519B">
<taxonomicName id="92B76402ECF3025ADAE6A09B13630465" authorityName="Telnov" authorityYear="2024" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="129482BA1060F388821ACC49BAA5532A" italics="true">Macrotomoderus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE035BA02452BEA5C47D508C89FA4DC0" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂
<emphasis id="70A605D4DBBC08D5B74C0D6B86611AD5" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="5563FB98BECED6A34B368CDF438567D1" bold="true">B</emphasis>
ditto, lateral view
<emphasis id="C0F0764567D87D5764C18B7B04703F5B" bold="true">C</emphasis>
dorsal forebody. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="8147C757A8016E522283CC38D2D02B1E" ID-DOI="10.3897/zookeys.1218.134413.figure4" ID-arpha="132EDE69-9FE4-5402-B0CC-10154EEB100D" httpUri="https://binary.pensoft.net/fig/1180021" startId="F4">
<paragraph id="2FBAEE9DB4FACC1D793C424990FEB821">
<label id="484CEE0B4F353E4D40D95F228538DF39">Figure 4.</label>
</paragraph>
<paragraph id="D4D086A1EEC4D471324FB2951C1AF4AC">
<taxonomicName id="C998572192C3F733F9AD0425766743E2" authorityName="Telnov" authorityYear="2024" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="749553B1F766530CC3F4B12625F93F7D" italics="true">Macrotomoderus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B096C055CD7246C8EF5C4DA5AEADCD2" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂, terminalia and aedeagus
<emphasis id="F29768998F083F6DD731180D60933A91" bold="true">A</emphasis>
sternite IX, lateral view
<emphasis id="BAFEBAFCF89A70CEFE5112F0FCA4FEB0" bold="true">B</emphasis>
aedeagus, ventral view
<emphasis id="98DF7DD82666CEEED5D49CDE5F3286EB" bold="true">C</emphasis>
ditto, lateral view
<emphasis id="F37159A30411EAC463F39247E6D1F75E" bold="true">D</emphasis>
ditto, median portion, endophallic armature
<emphasis id="38959E2622D41BECE232C1F346836ECA" bold="true">E</emphasis>
ditto, basal portion, endophallic armature
<emphasis id="AF4218841E272F243FFE0FF16672C123" bold="true">F</emphasis>
ditto, endophallic armature, lateral view of basal portion
<emphasis id="2A4E9DF597545355097926E87173C0B7" bold="true">G</emphasis>
ditto, ventral view of apex [not to scale].
</paragraph>
</caption>
<paragraph id="DCA56E37CD71FF014EA8323DE51D9F80">
<emphasis id="62D7F27A112D4FB3F0A590ACB2CAAD48" bold="true">
<emphasis id="6B76E068DA5D5E5A7A91984233858EFC" bold="true" italics="true">Sexual dimorphism</emphasis>
.
</emphasis>
Female is unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQHAE" type="Differential diagnosis">
<paragraph id="FC66C0C097A947561B75B8AFAF4CDC39">
<heading id="2E13E38223CA0151FAF2F248586EBE0E" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="E7BE2276968E207E38E7817552182639">
This species is readily recognized due to the presence of the dense clump of setae on male head base in the combination with the peculiar, spike / nail-like endophallic armature of the aedeagus. The shape of the aedeagus somewhat resembles that of
<taxonomicName id="7E82677FBEAD7335F65D596B80D6D1DA" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dali">
<emphasis id="07BB1E142F1AC6CEB0A673FB21E93F2E" italics="true">M. dali</emphasis>
<bibRefCitation id="12441B178DED5D54862097D095E8988F" DOI="10.5852/ejt.2022.797.1667" author="Telnov D" journalOrPublisher="European Journal of Taxonomy" pagination="1-100" refId="B11" refString="Telnov D (2022) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species. European Journal of Taxonomy 797: 1 - 100. https: // doi. org / 10.5852 / ejt. 2022.797. 1667" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species." volume="797" year="2022">Telnov, 2022</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="555A46C059B32ADF1B199E2BD8F7D761" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingCountry id="B7DEAF48AE99106A8993AFAB185C4583" name="China">China</collectingCountry>
),
<taxonomicName id="0E3D85A6EE11D7390938ADE12D279DB3" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muli">
<emphasis id="F570F818393845F514B0232750356338" italics="true">M. muli</emphasis>
<bibRefCitation id="B50EBFD2EA6626F7A18A26B0ACF1F9F4" DOI="10.5852/ejt.2022.797.1667" author="Telnov D" journalOrPublisher="European Journal of Taxonomy" pagination="1-100" refId="B11" refString="Telnov D (2022) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species. European Journal of Taxonomy 797: 1 - 100. https: // doi. org / 10.5852 / ejt. 2022.797. 1667" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species." volume="797" year="2022">Telnov, 2022</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="3ED19BF8A1796283FC66B9CBC2E9F2EC" country="China" name="Sichuan">Sichuan</collectingRegion>
,
<collectingCountry id="2B8FBFB325050C906FD85ABDBCBFC07E" name="China">China</collectingCountry>
), and
<taxonomicName id="DEC0C252057519A4E0FF49FD74907915" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wudu">
<emphasis id="8C2EB1E9325D3A770EEBC4325B64A1B0" italics="true">M. wudu</emphasis>
<bibRefCitation id="216465CE88DD4F83E69997587826BEDB" DOI="10.5852/ejt.2022.797.1667" author="Telnov D" journalOrPublisher="European Journal of Taxonomy" pagination="1-100" refId="B11" refString="Telnov D (2022) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species. European Journal of Taxonomy 797: 1 - 100. https: // doi. org / 10.5852 / ejt. 2022.797. 1667" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species." volume="797" year="2022">Telnov, 2022</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="6539E2302EA6659290750F30134579CF" country="China" name="Gansu">Gansu</collectingRegion>
,
<collectingCountry id="778E6D53B57F7FBAA257D0D06FC4C08F" name="China">China</collectingCountry>
), but all three have entirely different endophallic armatures. The head base with more or less dense setation is present in
<taxonomicName id="52B367526D57C3F74F7EEB8A82170492" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="conus">
<emphasis id="ED30D8ED40B62752530729EE9D24532D" italics="true">M. conus</emphasis>
<bibRefCitation id="B4C560E36387E6E8E54B4B4C1A5FEC45" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FFA3F04763A745E0C98796E9004B0453" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B9F48B1AE71CF9C52BF640B155253ACF" italics="true">M. gracilis</emphasis>
<bibRefCitation id="21C9A56C858208C197F3478EB7A1C6D7" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="6597F96E3F2A4541F5D5EFA141AF2596" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="microscopicus">
<emphasis id="AE133D1367F064D1B1C7DCD46EFC770C" italics="true">M. microscopicus</emphasis>
<bibRefCitation id="0603E6D00AC8F8150BA47955AF198FA3" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AE2CF18E2DF145E5AB39553946DCDC69" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monstrificabilis">
<emphasis id="7190BE25B35EC6BF41200630C98BA9FB" italics="true">M. monstrificabilis</emphasis>
<bibRefCitation id="3697DF66E655BC6D1D2378DF43B52B0F" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="892966B1156A426252DC7F60AFE074BC" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="perforatus">
<emphasis id="AF4643452AEB23D57908D8A6FCAC8268" italics="true">M. perforatus</emphasis>
<bibRefCitation id="D1AE23C513838BDAE906C669C778E8CA" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="1530AAF1077DB1019C3AC449935DB444" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="900FA719EC71FA5F671A783405250B56" italics="true">M. schuelkei</emphasis>
<bibRefCitation id="781AB8E07C1845592F8440D4E84BE5FD" DOI="10.3161/00034541ANZ2018.68.3.008" author="Telnov D" issue="3" journalOrPublisher="Annales zoologici" pagination="463-492" refId="B9" refString="Telnov D (2018) Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68. 3.008" title="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan." volume="68" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
(all from
<collectingRegion id="AD68A60A234DA8046F3232EE7D94CEE6" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingCountry id="4F244DBFABDC6017EEC65A59724A7357" name="China">China</collectingCountry>
), but other morphological features and the endophallic armature are quite different.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZKAE" type="ecology">
<paragraph id="88CFA3E27ED199983F4091757D61A99E">
<heading id="013DBAE41F824862BAF8D7E59975DDB6" reason="title">Ecology.</heading>
</paragraph>
<paragraph id="13323488B65D2C9048A013C8476BF5FB">
Collected at
<quantity id="BF1E08628208C104BF53B06F9B78CA05" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</quantity>
a. s. l.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5KAE" type="distribution">
<paragraph id="E9AF7CEAAB42ED7FAD55CC12B675C587">
<heading id="675F508375E49C0C8C09ADE02B0A3FF0" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="AD624DEF362B023A0CEB73F677E0F4AB">
Known from Daba Mountains in western part of
<collectingRegion id="4D3A78F164137D7765CB61CA0BF46024" country="China" name="Hubei">Hubei Province</collectingRegion>
, central
<collectingCountry id="433A72E8284589AE32FC91F200B4F10B" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDLAE" type="etymology">
<paragraph id="A03BEE921CC09E4231A184AD391A0206">
<heading id="0D8C9AD18D96478C45D0064E374847E8" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="97C0DB2014B8FB2B9543E28E6AD99C2D">
From Latin
<emphasis id="712762898C62A7F5ABF93506B5F7B83F" italics="true">hirsutus</emphasis>
shaggy, hairy, bristly, referring to the bristle of setae on the head base of this species.
</paragraph>
</subSubSection>
</treatment>
</document>