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<document id="B21DDF18185838085BA147D722EBD8C6" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFF85FFD6FF0D11A45CD1908A" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena dariensis Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="44" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:title id="30462809DA77A25C60A9B38A222BE928">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="44412ED49B5F7FF75711F82B85CC86D4" type="personal">
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<mods:namePart id="AF4D241EC2C4909678C8F786B9E82C49">Perkins, Philip D.</mods:namePart>
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<mods:title id="19F6CEFEAA15F03A9FCEF695A303AF46">Zootaxa</mods:title>
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<mods:part id="76FC4A231790352D46F6C16D7AED5DF5">
<mods:date id="ED117631AB45C78E5396C7CBAA7DD6FC">2011</mods:date>
<mods:detail id="D45B681F1A8E93E76D8F6F937302E4CE" type="pubDate">
<mods:number id="FF4318BC7D32F44303A09F42B76FEF11">2011-10-28</mods:number>
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<mods:number id="C7C7913D3CD0E303F90B942307E20917">3074</mods:number>
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<mods:classification id="2A52E913DDA4F464430F65CB8444A68A">journal article</mods:classification>
<mods:identifier id="DDE51DE4C0C181D75B9BE5B2A29C6769" type="CLB-Dataset">31530</mods:identifier>
<mods:identifier id="64EE26C42FC1D5E5716F07D976F18B52" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<treatment id="C063786AFF85FFD6FF0D11A45CD1908A" ID-DOI="http://doi.org/10.5281/zenodo.6184496" ID-GBIF-Taxon="137306991" ID-Zenodo-Dep="6184496" LSID="urn:lsid:plazi:treatment:C063786AFF85FFD6FF0D11A45CD1908A" httpUri="http://treatment.plazi.org/id/C063786AFF85FFD6FF0D11A45CD1908A" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="00D09AF7FF85FFD6FF0D11A45B1B9595" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="4875C97CFF85FFD6FF0D11A45D0395F1" blockId="43.[151,1082,151,213]" box="[151,546,151,177]" pageId="43" pageNumber="44">
<heading id="133D7E10FF85FFD6FF0D11A45D0395F1" bold="true" box="[151,546,151,177]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="7ABE156EFF85FFD6FF0D11A45D0395F1" bold="true" box="[151,546,151,177]" pageId="43" pageNumber="44">
<taxonomicName id="8FCAB2FFFF85FFD6FF0D11A45EA295F1" ID-CoL="3N38R" box="[151,387,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="dariensis" status="sp. nov.">
<emphasis id="7ABE156EFF85FFD6FF0D11A45EA295F1" bold="true" box="[151,387,151,177]" italics="true" pageId="43" pageNumber="44">Hydraena dariensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618DA815FF85FFD6FE0811A45D0395F1" box="[402,546,151,177]" pageId="43" pageNumber="44" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4875C97CFF85FFD6FF0D118E5B1B9595" blockId="43.[151,1082,151,213]" box="[151,1082,189,213]" pageId="43" pageNumber="44">
<figureCitation id="D0F1D5F9FF85FFD6FF0D118E5FD09595" box="[151,241,189,213]" captionStart="FIGURE 76" captionStartId="126.[151,250,1650,1673]" captionTargetBox="[268,1319,193,1626]" captionTargetId="figure@126.[268,1320,193,1629]" captionTargetPageId="126" captionText="FIGURE 76. Hydraena dariensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050146/files/figure.png" pageId="43" pageNumber="44">Figs. 76</figureCitation>
(habitus), 80 (aedeagus), 162 (female abd.), 178 (spermatheca), 203 (map)
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF85FFD6FF0D103B5ED494BF" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="4875C97CFF85FFD6FF0D103B5E4694CF" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FF0D103B5E699460" bold="true" box="[151,328,263,288]" pageId="43" pageNumber="44">
<typeStatus id="977177DEFF85FFD6FF0D103B5FF39460" box="[151,210,264,288]" pageId="43" pageNumber="44">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="977177DEFF85FFD6FED410345E97945F" box="[334,438,263,287]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="30DD89ECFF85FFD6FD8610345DB19460" box="[540,656,263,288]" name="Colombia" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FD8610345DB19460" bold="true" box="[540,656,263,288]" pageId="43" pageNumber="44">Colombia</emphasis>
</collectingCountry>
: Rio Atrato, Sautata, muddy puddles in marsh,
<date id="3C74EFBCFF85FFD6FB3610345A70945F" box="[1196,1361,263,287]" pageId="43" pageNumber="44" value="1972-03-27" valueMax="1972-03-31" valueMin="1972-03-27">2731 iii 1972</date>
, M. E. Bacchus. Deposited in the
<collectionCode id="2EDB51B9FF85FFD6FE58101D5D3B9405" box="[450,538,302,325]" pageId="43" pageNumber="44">BMNH</collectionCode>
.
<typeStatus id="977177DEFF85FFD6FDBE101E5DBD9405" box="[548,668,301,325]" pageId="43" pageNumber="44" type="paratype">
<emphasis id="7ABE156EFF85FFD6FDBE101E5DBD9405" bold="true" box="[548,668,301,325]" pageId="43" pageNumber="44">Paratypes</emphasis>
</typeStatus>
(21):
<collectingCountry id="30DD89ECFF85FFD6FD7D101E5C7D9406" box="[743,860,301,326]" name="Colombia" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FD7D101E5C7D9406" bold="true" box="[743,860,301,326]" pageId="43" pageNumber="44">Colombia</emphasis>
</collectingCountry>
: Same data as
<typeStatus id="977177DEFF85FFD6FB90101E5B4D9405" box="[1034,1132,301,325]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(5
<collectionCode id="2EDB51B9FF85FFD6FB08101E5BCA9405" box="[1170,1259,301,325]" pageId="43" pageNumber="44">BMNH</collectionCode>
); Rio Parencho, La Lomas, small puddle in clay,
<date id="3C74EFBCFF85FFD6FD9610615D94942A" box="[524,693,338,362]" pageId="43" pageNumber="44" value="1972-03-23" valueMax="1972-03-26" valueMin="1972-03-23">2326 iii 1972</date>
, M. E. Bacchus (4
<collectionCode id="2EDB51B9FF85FFD6FC0B10615CCB942A" box="[913,1002,338,362]" pageId="43" pageNumber="44">BMNH</collectionCode>
); Magd., Rio Frio [no date], P. J. Darlington (12
<collectionCode id="2EDB51B9FF85FFD6FE8010445E7D94CF" LSID="urn:lsid:biocol.org:col:33791" box="[282,348,375,399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="43" pageNumber="44">MCZ</collectionCode>
).
</paragraph>
<paragraph id="4875C97CFF85FFD6FF5D10AE5ED494BF" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FF5D10AE5EF294F6" bold="true" box="[199,467,413,438]" pageId="43" pageNumber="44">Differential Diagnosis.</emphasis>
A relatively densely punctate species, especially on the elytra, with a distinctive rectangular pronotal macula (
<figureCitation id="D0F1D5F9FF85FFD6FE2210F15D2C949A" box="[440,525,450,474]" captionStart="FIGURE 76" captionStartId="126.[151,250,1650,1673]" captionTargetBox="[268,1319,193,1626]" captionTargetId="figure@126.[268,1320,193,1629]" captionTargetPageId="126" captionText="FIGURE 76. Hydraena dariensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050146/files/figure.png" pageId="43" pageNumber="44">Fig. 76</figureCitation>
). The male genitalia suggest a relationship with
<taxonomicName id="8FCAB2FFFF85FFD6FBB410F75B98949B" box="[1070,1209,451,475]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="beniensis">
<emphasis id="7ABE156EFF85FFD6FBB410F75B98949B" box="[1070,1209,451,475]" italics="true" pageId="43" pageNumber="44">H. beniensis</emphasis>
</taxonomicName>
(
<figureCitation id="D0F1D5F9FF85FFD6FB5210F15A72949A" box="[1224,1363,450,474]" captionStart="FIGURES 78 81" captionStartId="128.[151,264,1650,1673]" captionText="FIGURES 78 81. Aedeagi of holotypes." pageId="43" pageNumber="44">Figs. 7980</figureCitation>
); refer to the diagnosis of that species.
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF85FFD6FF5D133E5D25903F" pageId="43" pageNumber="44" type="description">
<paragraph id="4875C97CFF85FFD6FF5D133E5C7297D5" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FF5D133E5E709766" bold="true" box="[199,337,525,550]" pageId="43" pageNumber="44">Description</emphasis>
. Size:
<typeStatus id="977177DEFF85FFD6FE37133E5D2E9765" box="[429,527,525,549]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.11/0.49; head 0.19/0.30; pronotum 0.30/0.39, PA 0.33, PB 0.35; elytra 0.67/0.49. Dorsum of head piceous; pronotum testaceous around rectangular piceous macula, ratios of color bands, as measured in midline, ca.
<date id="3C74EFBCFF85FFD6FB9D13645B70972F" box="[1031,1105,599,623]" pageId="43" pageNumber="44" value="2011-04-05">4/11/5</date>
; elytra dark brown; legs light brown to testaceous; maxillary palpi testaceous, tip not darker.
</paragraph>
<paragraph id="4875C97CFF85FFD6FF5D13905A639645" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">Frons punctures ca. 1.52xef, slightly larger and denser near eyes than medially; interstices shining, ca. 1xpd laterally, 24xpd medially. Clypeus microreticulate laterally, very finely sparsely punctulate medially. Mentum and postmentum finely, moderately densely punctulate, shining. Genae raised, shining, without posterior ridge.</paragraph>
<paragraph id="4875C97CFF85FFD6FF5D12205A249635" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; punctures on disc slightly larger and deeper than largest punctures on frons, interstices shining, ca. 12xpd on disc, punctures larger and denser at anterior and posterior; PF1, PF2 and PF4 absent; PF3 shallow.</paragraph>
<paragraph id="4875C97CFF85FFD6FF5D12B15DB4916F" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">Elytra weakly arcuate laterally; summit of posterior declivity at ca. midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming slightly smaller and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 1xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very shallow angle with one another.</paragraph>
<paragraph id="4875C97CFF85FFD6FF5D150E5D25903F" blockId="43.[151,1437,263,1483]" pageId="43" pageNumber="44">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2/1/4/3. P1 slightly wider than laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, parallel or nearly so, not raised, located at sides of moderately deep median longitudinal depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1.5x P2. Legs simple.
<collectingCountry id="30DD89ECFF85FFD6FAF815E15ABD91AA" box="[1378,1436,1234,1258]" name="Malta" pageId="43" pageNumber="44">Male</collectingCountry>
abdominal apex symmetrical. Female (microslide mount, n=1): last tergite broadly rounded, without apicomedian notch, ca. 22 (n=1) strongly hooked setae, no median gap; gonocoxite not midlongitudinally divided, apical margin sharply rounded, low transverse ridge evident at border of microreticulation (
<figureCitation id="D0F1D5F9FF85FFD6FB9614715B50901A" box="[1036,1137,1346,1370]" captionStart="FIGURES 160 162" captionStartId="179.[151,264,1650,1673]" captionText="FIGURES 160 162. Female tergite X and gonocoxite of Hydraena species." pageId="43" pageNumber="44">Fig. 162</figureCitation>
); spermatheca not C-type, with concave capsule (
<figureCitation id="D0F1D5F9FF85FFD6FE0F14545ED9903F" box="[405,504,1383,1407]" captionStart="FIGURES 174 178" captionStartId="183.[151,264,1600,1623]" captionTargetBox="[251,1323,222,1568]" captionTargetId="figure@183.[226,1360,193,1579]" captionTargetPageId="183" captionText="FIGURES 174 178. Spermathecae of Hydraena species." httpUri="https://zenodo.org/record/1050230/files/figure.png" pageId="43" pageNumber="44">Fig. 178</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF85FFD6FF5D14BE5C2E90E5" box="[199,783,1421,1446]" pageId="43" pageNumber="44" type="etymology">
<paragraph id="4875C97CFF85FFD6FF5D14BE5C2E90E5" blockId="43.[151,1437,263,1483]" box="[199,783,1421,1446]" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FF5D14BE5E6F90E5" bold="true" box="[199,334,1421,1445]" pageId="43" pageNumber="44">Etymology.</emphasis>
Named in reference to the
<typeStatus id="977177DEFF85FFD6FDE414BD5D8F90E6" box="[638,686,1422,1446]" pageId="43" pageNumber="44">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF85FFD6FF5D14815CD1908A" box="[199,1008,1458,1483]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="4875C97CFF85FFD6FF5D14815CD1908A" blockId="43.[151,1437,263,1483]" box="[199,1008,1458,1483]" pageId="43" pageNumber="44">
<emphasis id="7ABE156EFF85FFD6FF5D14815E41908B" bold="true" box="[199,352,1458,1483]" pageId="43" pageNumber="44">Distribution.</emphasis>
Currently known from northwestern
<collectingCountry id="30DD89ECFF85FFD6FC9914815C50908A" box="[771,881,1458,1482]" name="Colombia" pageId="43" pageNumber="44">Colombia</collectingCountry>
(
<figureCitation id="D0F1D5F9FF85FFD6FC1B14815CC2908A" box="[897,995,1458,1482]" captionStart="FIGURE 203" captionStartId="196.[151,250,1650,1673]" captionText="FIGURE 203. Geographical distributions of Hydraena species." pageId="43" pageNumber="44">Fig. 203</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>