171 lines
18 KiB
XML
171 lines
18 KiB
XML
<document id="AF4929C95E183841546E5D725BB827D4" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFF80FFD2FF0D12BF5DAC977C" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena unita Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="47" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:titleInfo id="2382B77945577BA71FD6A392B8234058">
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<mods:title id="F5692352FCD7FB86A68B1457BFAAD5D9">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F18130F4D36EDDF6907CB2C8E4C30C7B" type="personal">
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<mods:roleTerm id="07F5F8EBA36638E9FC764DEE21A1E72B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="338C78F301F6CAE5808F910C464AE943">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="15816BA7C30890EA4802348666038267">text</mods:typeOfResource>
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<mods:title id="AC29084C4FC8F211F97306CB7A2253F9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="ECCC694EF5CEE95FC8F3AD37F8FEB5B2">
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<mods:date id="01867063E0BD8E0569242F7AA2C46280">2011</mods:date>
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<mods:detail id="17A1493DFFB6ABF06F0D91BFEE4F7456" type="pubDate">
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<mods:number id="1D34585FA8C793921341CF0EE6BD4C32">2011-10-28</mods:number>
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</mods:detail>
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<mods:detail id="09641B5E8C0D7BED8FDD403B71EB17E4" type="volume">
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<mods:number id="ED4D63F3C037FAC8123D80387AA6F224">3074</mods:number>
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<mods:end id="E871BBB4212C2BB533381448C50F6347">198</mods:end>
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</mods:relatedItem>
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<mods:classification id="31A0B3254CB23A78A22006B8DA4C5836">journal article</mods:classification>
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<mods:identifier id="2F688A30B34D7FA6D418A58C5A45F89D" type="CLB-Dataset">31530</mods:identifier>
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<mods:identifier id="32A8E94152474916A822DD99FB49BABF" type="DOI">10.5281/zenodo.1050060</mods:identifier>
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<mods:identifier id="6DB6671A0605B32C16811CCEDBC02931" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A39744162C2442C6925BA6C7A6A21645" type="Zenodo-Dep">1050060</mods:identifier>
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<treatment id="C063786AFF80FFD2FF0D12BF5DAC977C" ID-DOI="http://doi.org/10.5281/zenodo.6184514" ID-GBIF-Taxon="137306980" ID-Zenodo-Dep="6184514" LSID="urn:lsid:plazi:treatment:C063786AFF80FFD2FF0D12BF5DAC977C" httpUri="http://treatment.plazi.org/id/C063786AFF80FFD2FF0D12BF5DAC977C" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<subSubSection id="00D09AF7FF80FFD3FF0D12BF5DA99686" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="4875C97CFF80FFD3FF0D12BF5ED596E5" blockId="46.[151,648,907,966]" box="[151,500,907,934]" pageId="46" pageNumber="47">
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<heading id="133D7E10FF80FFD3FF0D12BF5ED596E5" bold="true" box="[151,500,907,934]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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||
<emphasis id="7ABE156EFF80FFD3FF0D12BF5ED596E5" bold="true" box="[151,500,907,934]" pageId="46" pageNumber="47">
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||
<taxonomicName id="8FCAB2FFFF80FFD3FF0D12BF5E7496E6" ID-CoL="3N3KG" box="[151,341,908,934]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="unita" status="sp. nov.">
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<emphasis id="7ABE156EFF80FFD3FF0D12BF5E7496E6" bold="true" box="[151,341,908,934]" italics="true" pageId="46" pageNumber="47">Hydraena unita</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="618DA815FF80FFD3FEFE12B85ED596E5" box="[356,500,907,933]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4875C97CFF80FFD3FF0D129D5DA99686" blockId="46.[151,648,907,966]" box="[151,648,942,966]" pageId="46" pageNumber="47">
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<figureCitation id="D0F1D5F9FF80FFD3FF0D129D5FD09686" box="[151,241,942,966]" captionStart="FIGURE 90" captionStartId="134.[151,250,1650,1673]" captionTargetBox="[236,1357,202,1615]" captionTargetId="figure@134.[217,1370,193,1629]" captionTargetPageId="134" captionText="FIGURE 90. Hydraena unita, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050158/files/figure.png" pageId="46" pageNumber="47">Figs. 90</figureCitation>
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(habitus), 93 (aedeagus), 202 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF80FFD3FF0D12C45B49904A" pageId="46" pageNumber="47" type="materials_examined">
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<paragraph id="4875C97CFF80FFD3FF0D12C45E5991DE" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FF0D12C45E66914F" bold="true" box="[151,327,1014,1039]" pageId="46" pageNumber="47">
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<typeStatus id="977177DEFF80FFD3FF0D12C45FF1914F" box="[151,208,1015,1039]" pageId="46" pageNumber="47">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="977177DEFF80FFD3FECB12C55E98914E" box="[337,441,1014,1038]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="30DD89ECFF80FFD3FDB912C55D4D914F" box="[547,620,1014,1039]" name="Brazil" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FDB912C55D4D914F" bold="true" box="[547,620,1014,1039]" pageId="46" pageNumber="47">Brazil</emphasis>
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</collectingCountry>
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: Matto Grosso, Rio Bento Gomes (Pantanal); Flussabschnitt bei Periquito,
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<geoCoordinate id="2DFEAFBBFF80FFD3FF4815295E1D9172" box="[210,316,1050,1074]" direction="south" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="-15.9">15° 54' S</geoCoordinate>
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,
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<geoCoordinate id="2DFEAFBBFF80FFD3FED015295E9C9172" box="[330,445,1050,1074]" direction="west" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="-56.55">56° 33' W</geoCoordinate>
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,
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<date id="3C74EFBCFF80FFD3FE5015295D099172" box="[458,552,1050,1074]" pageId="46" pageNumber="47" value="1993-01-01">1 i 1993</date>
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–
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<date id="3C74EFBCFF80FFD3FDDD15295DEA9172" box="[583,715,1050,1074]" pageId="46" pageNumber="47" value="1994-12-30">30 xii 1994</date>
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, E. Stuhr (31). Deposited in the
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<collectionCode id="2EDB51B9FF80FFD3FBA715285BA79172" box="[1085,1158,1051,1074]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47">NMW</collectionCode>
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.
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<typeStatus id="977177DEFF80FFD3FB0815285A2B9173" box="[1170,1290,1051,1075]" pageId="46" pageNumber="47" type="paratype">
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<emphasis id="7ABE156EFF80FFD3FB0815285A2B9173" bold="true" box="[1170,1290,1051,1075]" pageId="46" pageNumber="47">Paratypes</emphasis>
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</typeStatus>
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(8):
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<collectingCountry id="30DD89ECFF80FFD3FAD115295AB59173" box="[1355,1428,1050,1075]" name="Brazil" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FAD115295AB59173" bold="true" box="[1355,1428,1050,1075]" pageId="46" pageNumber="47">Brazil</emphasis>
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</collectingCountry>
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: Matto Grosso, Same data as
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<typeStatus id="977177DEFF80FFD3FE7B150D5D629116" box="[481,579,1086,1110]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
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(2
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<collectionCode id="2EDB51B9FF80FFD3FDF6150D5D989116" box="[620,697,1086,1110]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47">NMW</collectionCode>
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);
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<collectingCountry id="30DD89ECFF80FFD3FD54150C5C609117" box="[718,833,1087,1111]" name="Paraguay" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FD54150C5C609117" bold="true" box="[718,833,1087,1111]" pageId="46" pageNumber="47">Paraguay</emphasis>
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</collectingCountry>
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: Dep. Alto
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<collectingCountry id="30DD89ECFF80FFD3FC4E150C5B649117" box="[980,1093,1087,1111]" name="Paraguay" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FC4E150C5B649117" bold="true" box="[980,1093,1087,1111]" pageId="46" pageNumber="47">Paraguay</emphasis>
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</collectingCountry>
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,
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<quantity id="8F326499FF80FFD3FBCF150D5BBF9116" box="[1109,1182,1086,1110]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="km" value="20.0">20 km</quantity>
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E Aqua Dulce,
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<date id="3C74EFBCFF80FFD3FAC3150D5FF3913A" pageId="46" pageNumber="47" value="1995-07-27">27 vii 1995</date>
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, Drechsel (3
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<collectionCode id="2EDB51B9FF80FFD3FEEB15515E9F913A" box="[369,446,1122,1146]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47">NMW</collectionCode>
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);
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<quantity id="8F326499FF80FFD3FE4E15515D3F913A" box="[468,542,1122,1146]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="47" unit="km" value="90.0">90 km</quantity>
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NW Bahia Negra,
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<date id="3C74EFBCFF80FFD3FC9E15515CAF913A" box="[772,910,1122,1146]" pageId="46" pageNumber="47" value="1995-07-30">30 vii 1995</date>
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, Drechsel (2
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<collectionCode id="2EDB51B9FF80FFD3FBB715515B5B913A" box="[1069,1146,1122,1146]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47">NMW</collectionCode>
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); Madrejon,
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<date id="3C74EFBCFF80FFD3FA9415515AB9913A" box="[1294,1432,1122,1146]" pageId="46" pageNumber="47" value="1995-07-27">27 vii 1995</date>
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, Drechsel (1
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<collectionCode id="2EDB51B9FF80FFD3FEBA15B55E4C91DE" box="[288,365,1158,1182]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47">NMW</collectionCode>
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).
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</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D15995B49904A" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FF5D15995EF39183" bold="true" box="[199,466,1194,1219]" pageId="46" pageNumber="47">Differential Diagnosis.</emphasis>
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This species is quite distinct in dorsal habitus, being densely punctate and microreticulate, lacking a pronotal macula, and having pronotal foveae PF2 confluent, forming a U-shaped impression (
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<figureCitation id="D0F1D5F9FF80FFD3FAF415FD5F96904A" captionStart="FIGURE 90" captionStartId="134.[151,250,1650,1673]" captionTargetBox="[236,1357,202,1615]" captionTargetId="figure@134.[217,1370,193,1629]" captionTargetPageId="134" captionText="FIGURE 90. Hydraena unita, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050158/files/figure.png" pageId="46" pageNumber="47">Fig. 90</figureCitation>
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). The aedeagus does not readily compare with any other described species (
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<figureCitation id="D0F1D5F9FF80FFD3FB9D15C15B7A904A" box="[1031,1115,1266,1290]" captionStart="FIGURES 93 – 95" captionStartId="137.[151,264,1650,1673]" captionText="FIGURES 93 – 95. Aedeagi of holotypes." pageId="46" pageNumber="47">Fig. 93</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF80FFD2FF5D14255E6E94EC" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
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<paragraph id="4875C97CFF80FFD3FF5D14255C7F90DA" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">
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<emphasis id="7ABE156EFF80FFD3FF5D14255E70906F" bold="true" box="[199,337,1302,1327]" pageId="46" pageNumber="47">Description</emphasis>
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. Size:
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<typeStatus id="977177DEFF80FFD3FE3714255D2E906E" box="[429,527,1302,1326]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.36/0.56; head 0.21/0.31; pronotum 0.34/0.43, PA 0.35, PB 0.40; elytra 0.83/0.56. Dorsum of head dark brown; pronotum, elytra and legs brown to slightly reddish brown. All specimens in
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<typeStatus id="977177DEFF80FFD3FD4E146C5C259037" box="[724,772,1375,1399]" pageId="46" pageNumber="47">type</typeStatus>
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series appear to be teneral; however, there is no indication of a pronotal macula. Maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D14945D469346" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">Frons and clypeus microreticulate, dull. Frons punctures ca. 2xef, larger and denser near eyes than medially; interstices ca. 0.5–1xpd laterally, ca. 1–2xpd medially. Mentum and postmentum microreticulate, dull. Genae raised, dull, with posterior ridge laterally.</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D17205BA293FA" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, with median area very smooth and shining, but not produced and without distinct scintilla; punctures on disc much larger and deeper than those on frons, interstices effacedly microreticulate, weakly shining, 0.5–1xpd on disc, punctures not distinctly larger and denser at anterior and posterior; PF1 absent; PF2 moderately deep, shallow confluent medially, forming U-shaped impression; PF3 deep; PF4 shallow; PF3 and PF4 shallowly confluent.</paragraph>
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<paragraph id="4875C97CFF80FFD3FF5D17F55A3B922E" blockId="46.[151,1437,1014,2011]" pageId="46" pageNumber="47">Elytra weakly arcuate laterally; summit of posterior declivity at about midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming slightly smaller and more widely spaced toward posterior. Intervals not raised, effacedly microreticulate, weakly shining; on basal 1/3 ca. 1–2xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect conjointly, rather sharply rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="4875C97CFF80FFD2FF5D16495E6E94EC" blockId="46.[151,1437,1014,2011]" lastBlockId="47.[151,1436,151,572]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/4/5. P1 laminate; median carina sinuate in profile. P2 narrow, l/w ca. 4/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques weakly raised carinate lines, parallel, located at sides of deep median depression. Metaventrite with very small point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca.
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<date id="3C74EFBCFF81FFD2FF72118F5E259594" box="[232,260,188,212]" pageId="47" pageNumber="48">4x</date>
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P2. Protibia (male) straight, slightly widened on median margin at distal 1/3. Mesotibia slender, straight. Metatibia very slender, very slightly arcuate. Metatrochanter posterior margin angulate. Abdominal apex symmetrical; last tergite concave on each side of median longitudinal ridge, free margin sharply rounded. Female abdominal apex: last tergite broadly rounded, with shallow apicomedian emargination, ca. 18 (n=1) hooked setae, in slightly irregular row, no median gap; gonocoxite not midlongitudinally divided, apical margin sharply rounded, very weak transverse ridge evident at border of microreticulation; spermatheca not examined (poor preparation on solitary female).
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</paragraph>
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</subSubSection>
|
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<subSubSection id="00D09AF7FF81FFD2FF5D108B5B2C948F" box="[199,1037,439,464]" pageId="47" pageNumber="48" type="etymology">
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<paragraph id="4875C97CFF81FFD2FF5D108B5B2C948F" blockId="47.[151,1436,151,572]" box="[199,1037,439,464]" pageId="47" pageNumber="48">
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||
<emphasis id="7ABE156EFF81FFD2FF5D108B5E6F9490" bold="true" box="[199,334,440,464]" pageId="47" pageNumber="48">Etymology.</emphasis>
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Named in reference to the united PF2 foveae of the pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D09AF7FF81FFD2FF5D10E85DAC977C" pageId="47" pageNumber="48" type="distribution">
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||
<paragraph id="4875C97CFF81FFD2FF5D10E85BB894B4" blockId="47.[151,1436,151,572]" box="[199,1177,475,501]" pageId="47" pageNumber="48">
|
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<emphasis id="7ABE156EFF81FFD2FF5D10E85E4194B4" bold="true" box="[199,352,475,500]" pageId="47" pageNumber="48">Distribution.</emphasis>
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Currently known from southern
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<collectingCountry id="30DD89ECFF81FFD2FD4B10EF5C3494B4" box="[721,789,476,500]" name="Brazil" pageId="47" pageNumber="48">Brazil</collectingCountry>
|
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and northern
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<collectingCountry id="30DD89ECFF81FFD2FC2810EE5B3B94B5" box="[946,1050,477,501]" name="Paraguay" pageId="47" pageNumber="48">Paraguay</collectingCountry>
|
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(
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<figureCitation id="D0F1D5F9FF81FFD2FBB310EF5BAC94B4" box="[1065,1165,476,500]" captionStart="FIGURE 202" captionStartId="195.[151,250,1650,1673]" captionText="FIGURE 202. Geographical distributions of Hydraena species." pageId="47" pageNumber="48">Fig. 202</figureCitation>
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||
).
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</paragraph>
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<paragraph id="4875C97CFF81FFD2FF5D13335DAC977C" blockId="47.[151,1436,151,572]" pageId="47" pageNumber="48">
|
||
<emphasis id="7ABE156EFF81FFD2FF5D13335E309758" bold="true" box="[199,273,512,536]" pageId="47" pageNumber="48">Notes.</emphasis>
|
||
The aedeagus of a specimen from
|
||
<collectingCountry id="30DD89ECFF81FFD2FD0713335C279758" box="[669,774,512,536]" name="Paraguay" pageId="47" pageNumber="48">Paraguay</collectingCountry>
|
||
,
|
||
<quantity id="8F326499FF81FFD2FC8B10CC5C789757" box="[785,857,511,535]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="48" unit="km" value="90.0">90 km</quantity>
|
||
NW Bahia Negra, is slightly wider than that of the
|
||
<typeStatus id="977177DEFF81FFD2FF0D13175FDD977C" box="[151,252,548,572]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
, and the body is also slightly wider.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |