278 lines
40 KiB
XML
278 lines
40 KiB
XML
<document id="99E36921A1E408AC10D98E984468E526" ID-CLB-Dataset="45422" ID-DOI="10.11646/zootaxa.3821.1.11" ID-GBIF-Dataset="d71e28bf-f1bf-404f-8475-970828c0b492" ID-ISSN="1175-5326" ID-Zenodo-Dep="226219" ID-ZooBank="50C1CE30-6F91-4B08-B93A-A749F6579807" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460440634606" checkinUser="plazi" docAuthor="Molineri, C." docDate="2014" docId="6E1987B9FFCAFF8CFF35E3C86858FCDE" docLanguage="en" docName="zt03821p145.pdf" docOrigin="Zootaxa 3821 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloretochus peruanicus Soldan" docType="treatment" docVersion="9" lastPageNumber="142" masterDocId="9220FFC1FFCBFF8FFFA2E2376A3BFFE6" masterDocTitle="Description of Alloretochus sigillatus new species with comments and new distributional records for Alloretochus peruanicus (Ephemeroptera, Caenidae, Brachycercinae)" masterLastPageNumber="145" masterPageNumber="139" pageNumber="140" updateTime="1698332865549" updateUser="ExternalLinkService">
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<mods:titleInfo id="50FA43BC074BC5E58EAB1FF6DC580EBA">
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<mods:title id="DB86140E686BC06D3E5DF0637E59AEFE">Description of Alloretochus sigillatus new species with comments and new distributional records for Alloretochus peruanicus (Ephemeroptera, Caenidae, Brachycercinae)</mods:title>
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<mods:name id="F38B64C2B90C6C08616C6A9A0B2D225F" type="personal">
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<mods:namePart id="F8F6D86F9487397D47023B9310AAE460">Molineri, C.</mods:namePart>
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<mods:typeOfResource id="DE9A06BFCF4C8F31244C0BF374FEBCA2">text</mods:typeOfResource>
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<mods:title id="00277E10B9B648785BABE4A74BA92492">Zootaxa</mods:title>
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<mods:part id="8D29532C034F2963D0695D43B32F0255">
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<mods:date id="0C15D4C57C51390633C1255A99CC8D84">2014</mods:date>
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<mods:detail id="AD8A500DF2556C5A989611C898C2CEF0" type="volume">
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<mods:number id="DDF52E1D10292DFD55E9DC73E751D2FF">3821</mods:number>
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<mods:number id="C3F0932D8EB24049E1DAD7FEA44C34D0">1</mods:number>
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<mods:identifier id="58AC9A36D00AA09AF65103BB261D45DE" type="CLB-Dataset">45422</mods:identifier>
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<mods:identifier id="84CD596AFD349879A71B4B5D9E7E8E82" type="DOI">10.11646/zootaxa.3821.1.11</mods:identifier>
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<treatment id="6E1987B9FFCAFF8CFF35E3C86858FCDE" ID-DOI="http://doi.org/10.5281/zenodo.6137792" ID-GBIF-Taxon="119507887" ID-Zenodo-Dep="6137792" LSID="urn:lsid:plazi:treatment:6E1987B9FFCAFF8CFF35E3C86858FCDE" httpUri="http://treatment.plazi.org/id/6E1987B9FFCAFF8CFF35E3C86858FCDE" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140">
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<subSubSection id="AEAA6524FFCAFF8EFF35E3C86B49FDDD" pageId="1" pageNumber="140" type="nomenclature">
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<paragraph id="E60F36AFFFCAFF8EFF35E3C8680AFDFC" blockId="1.[151,561,511,571]" box="[151,561,511,538]" pageId="1" pageNumber="140">
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<heading id="BD4781C3FFCAFF8EFF35E3C8680AFDFC" bold="true" box="[151,561,511,538]" fontSize="11" level="1" pageId="1" pageNumber="140" reason="1">
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<taxonomicName id="21B04D2CFFCAFF8EFF35E3C8680AFDFC" ID-CoL="BZ37" authority="Soldan" authorityName="Soldan" box="[151,561,511,538]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="peruanicus">
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<emphasis id="D4C4EABDFFCAFF8EFF35E3C8680AFDFC" bold="true" box="[151,561,511,538]" pageId="1" pageNumber="140">
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<emphasis id="D4C4EABDFFCAFF8EFF35E3C86BFBFDFF" bold="true" box="[151,448,511,537]" italics="true" pageId="1" pageNumber="140">Alloretochus peruanicus</emphasis>
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(Soldán)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="E60F36AFFFCAFF8EFF35E0156B49FDDD" blockId="1.[151,561,511,571]" box="[151,370,546,571]" pageId="1" pageNumber="140">Figs. (1A–I, 3A–B)</paragraph>
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</subSubSection>
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<subSubSection id="AEAA6524FFCAFF8EFF35E05C691AFD79" pageId="1" pageNumber="140" type="reference_group">
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<paragraph id="E60F36AFFFCAFF8EFF35E05C691AFD79" blockId="1.[151,1212,618,671]" pageId="1" pageNumber="140">
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<treatmentCitationGroup id="C6A01181FFCAFF8EFF35E05C691AFD79" pageId="1" pageNumber="140">
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<taxonomicName id="21B04D2CFFCAFF8EFF35E05C6866FD66" authority="Soldan, 1986: 343" authorityName="Soldan" authorityPageNumber="343" authorityYear="1986" box="[151,605,618,640]" class="Insecta" family="Caenidae" genus="Cercobrachys" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="peruanicus">
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<emphasis id="D4C4EABDFFCAFF8EFF35E05C6BA1FD66" box="[151,410,619,640]" italics="true" pageId="1" pageNumber="140">Cercobrachys peruanicus</emphasis>
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<treatmentCitation id="671110BEFFCAFF8EFE03E05D6866FD66" author="Soldan" box="[417,605,618,640]" page="343" pageId="1" pageNumber="140" year="1986">
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<bibRefCitation id="82214B5EFFCAFF8EFE03E05D6811FD66" author="Soldan" box="[417,554,618,640]" pageId="1" pageNumber="140" refString="Soldan, T. (1986) A revision of the Caenidae with ocellar tubercles in the nymphal stage (Ephemeroptera). Acta Universitatis Carolinae-Biologica, 1982 - 1984, 289 - 362." type="book chapter" year="1986">Soldán, 1986</bibRefCitation>
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: 343
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="671110BEFFCAFF8EFDC5E05D69B3FD66" author="Molineri" box="[615,904,618,640]" page="64" pageId="1" pageNumber="140" year="2006">
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<bibRefCitation id="82214B5EFFCAFF8EFDC5E05D6959FD66" author="Molineri" box="[615,866,618,640]" pageId="1" pageNumber="140" refString="Molineri, C. & Goitia, E. (2006) Description of the adult stage of Cercobrachys peruanicus (Ephemeroptera: Caenidae). Revista de la Sociedad Entomologica Argentina, 65, 63 - 67." type="journal article" year="2006">Molineri & Goitía, 2006</bibRefCitation>
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: 64
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</treatmentCitation>
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; Domínguez
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<emphasis id="D4C4EABDFFCAFF8EFBADE05C6E79FD66" box="[1039,1090,619,640]" italics="true" pageId="1" pageNumber="140">et al.</emphasis>
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, 2006: 223.
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<taxonomicName id="21B04D2CFFCAFF8EFF35E0BE6BD6FD79" ID-CoL="BZ37" authority="Soldan" authorityName="Soldan" box="[151,493,649,671]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="peruanicus">
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<emphasis id="D4C4EABDFFCAFF8EFF35E0BE6BB5FD79" box="[151,398,649,671]" italics="true" pageId="1" pageNumber="140">Alloretochus peruanicus</emphasis>
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(Soldán)
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</taxonomicName>
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,
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<treatmentCitation id="671110BEFFCAFF8EFE5AE0BE6927FD79" author="Sun" box="[504,796,649,671]" page="78" pageId="1" pageNumber="140" year="2008">
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<bibRefCitation id="82214B5EFFCAFF8EFE5AE0BE68CDFD79" author="Sun" box="[504,758,649,671]" pageId="1" pageNumber="140" refString="Sun, L. & MCCafferty. W. P. (2008) Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa, 1801, 1 - 239." type="journal article" year="2008">Sun & McCafferty, 2008</bibRefCitation>
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: 78
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="AEAA6524FFCAFF8EFF35E0FB681EFC92" pageId="1" pageNumber="140" type="diagnosis">
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<paragraph id="E60F36AFFFCAFF8EFF35E0FB681EFC92" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
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<emphasis id="D4C4EABDFFCAFF8EFF35E0FB6B2AFD03" bold="true" box="[151,273,716,741]" pageId="1" pageNumber="140">Diagnosis.</emphasis>
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Adult size variable (body length
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<quantity id="21489B4AFFCAFF8EFD0DE0FB692DFD02" box="[687,790,716,740]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="140" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
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); body with gray shading widely extended mainly on abdomen (darker on terga I–II and VII–VIII in male) and thoracic sterna (
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<figureCitation id="7E8B2A2AFFCAFF8EFC74E0D86EA2FCEE" box="[982,1177,751,776]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Figs. 1B–C, E–F</figureCitation>
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); male subgenital plate broadly emarginated posteriorly as in
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<figureCitation id="7E8B2A2AFFCAFF8EFD9CE12368FFFCCA" box="[574,708,788,813]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Figs. 1G–H</figureCitation>
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; ratios forceps length/subbasal width 6.8–8.0; female sternum IX produced distally but not reaching apex of segment X, distally truncated with small median notch; egg with 7–8 costae in a lateral half (
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<figureCitation id="7E8B2A2AFFCAFF8EFE3EE16B6BD2FC92" box="[412,489,860,885]" captionStart="FIGURE 2. A – G" captionStartId="4.[151,250,1702,1724]" captionTargetBox="[191,1395,193,1680]" captionTargetId="figure@4.[191,1395,193,1681]" captionTargetPageId="4" captionText="FIGURE 2. A – G: Alloretochus sigillatus sp. nov.: A, male imago, head and thorax, lateral view; B, same, dorsal view of abdomen; C, same, detail of head; D, male genitalia, ventral view; E, female imago, dorsal view; F – G, eggs. H – I, A. peruanicus eggs." httpUri="https://zenodo.org/record/226221/files/figure.png" pageId="1" pageNumber="140">Figs. 2</figureCitation>
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H–I).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="AEAA6524FFCAFF8EFF65E1486AC5F91E" pageId="1" pageNumber="140" type="materials_examined">
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<paragraph id="E60F36AFFFCAFF8EFF65E14868BAFB96" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
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<emphasis id="D4C4EABDFFCAFF8EFF65E1486B0DFC7E" bold="true" box="[199,310,895,920]" pageId="1" pageNumber="140">Material.</emphasis>
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<specimenCount id="F0B6FD26FFCAFF8EFE9FE1B76BB3FC7E" box="[317,392,895,920]" pageId="1" pageNumber="140" type="male">7 male</specimenCount>
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and
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<specimenCount id="F0B6FD26FFCAFF8EFE1DE1B7681AFC7E" box="[447,545,895,920]" pageId="1" pageNumber="140" type="female">4 female</specimenCount>
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imagos (parts of a male on slide IBN636CM) from
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<collectingCountry id="9EA7763FFFCAFF8EFBCBE1486EE0FC7E" box="[1129,1243,895,920]" name="Argentina" pageId="1" pageNumber="140">Argentina</collectingCountry>
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, Jujuy, Ledesma, río San Francisco, puente a Aguas Calientes,
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<geoCoordinate id="83845068FFCAFF8EFD31E1936919FC5A" box="[659,802,932,956]" direction="south" orientation="latitude" pageId="1" pageNumber="140" precision="15" value="-23.711668">S 23º 42' 42"</geoCoordinate>
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—
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<geoCoordinate id="83845068FFCAFF8EFCE2E19369EAFC5A" box="[832,977,932,957]" direction="west" orientation="longitude" pageId="1" pageNumber="140" precision="15" value="-64.53445">W 64º 32' 4"</geoCoordinate>
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, m,
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<date id="920E106FFFCAFF8EFBA0E1936EA1FC5A" box="[1026,1178,932,956]" pageId="1" pageNumber="140" value="2013-01-17" valueMax="2013-01-18" valueMin="2013-01-17">17–18.I.2013</date>
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, light trap, C. Molineri col.;
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<specimenCount id="F0B6FD26FFCAFF8EFF72E1FF6B27FC06" box="[208,284,967,992]" pageId="1" pageNumber="140" type="male">1 male</specimenCount>
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imago from
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<collectingCountry id="9EA7763FFFCAFF8EFE10E1FF6813FC06" box="[434,552,968,992]" name="Bolivia" pageId="1" pageNumber="140">BOLIVIA</collectingCountry>
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, río Blanco, ca. de Once por Ciento, camino entre Sta. Cruz y
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<collectingCountry id="9EA7763FFFCAFF8EFB53E1F06F68FC06" box="[1265,1363,967,992]" name="Trinidad and Tobago" pageId="1" pageNumber="140">Trinidad</collectingCountry>
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,
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<geoCoordinate id="83845068FFCAFF8EFAC2E1FF6AC1FBE2" direction="south" orientation="latitude" pageId="1" pageNumber="140" precision="1" value="-15.361028">S 15º 21' 39.7"</geoCoordinate>
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—
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<geoCoordinate id="83845068FFCAFF8EFEBBE1DB6BEDFBE2" box="[281,470,1004,1029]" direction="west" orientation="longitude" pageId="1" pageNumber="140" precision="1" value="-63.291332">W 63º 17' 28.8"</geoCoordinate>
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,
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<quantity id="21489B4AFFCAFF8EFE46E1DB6814FBE2" box="[484,559,1004,1029]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="140" unit="m" value="250.0">250 m</quantity>
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,
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<date id="920E106FFFCAFF8EFD9CE1DB68F9FBE2" box="[574,706,1004,1028]" pageId="1" pageNumber="140" value="2000-06-14">14/VI/2000</date>
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, E. Domínguez col.;
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<specimenCount id="F0B6FD26FFCAFF8EFC1AE1DB6E3DFBE3" box="[952,1030,1004,1029]" pageId="1" pageNumber="140" type="male">1 male</specimenCount>
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||
imago (parts on slide IBN621CM) from
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<collectingCountry id="9EA7763FFFCAFF8EFF76E6276B57FBCE" box="[212,364,1040,1064]" name="Colombia" pageId="1" pageNumber="140">COLOMBIA</collectingCountry>
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, Dpto Amazonas, P.N. Amacayacu, Qda. Mata-Mata,
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<geoCoordinate id="83845068FFCAFF8EFC6BE6276E71FBCE" box="[969,1098,1040,1064]" direction="south" orientation="latitude" pageId="1" pageNumber="140" precision="15" value="-3.8077776">S 3º 48' 28"</geoCoordinate>
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||
—
|
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<geoCoordinate id="83845068FFCAFF8EFBC5E6276F3DFBCE" box="[1127,1286,1039,1064]" direction="west" orientation="longitude" pageId="1" pageNumber="140" precision="15" value="-70.25584">W 70º 15' 21"</geoCoordinate>
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,
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<date id="920E106FFFCAFF8EFAB1E6386FA3FBCE" box="[1299,1432,1039,1064]" pageId="1" pageNumber="140" value="1999-02-02" valueMax="1999-02-05" valueMin="1999-02-02">2–5.II.1999</date>
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||
, at light 4–6 h AM, E. Domínguez, M.C. Zúñiga & C. Molineri cols. All material deposited in IBN. Material listed in
|
||
<bibRefCitation id="82214B5EFFCAFF8EFF16E6606BEAFB96" author="Molineri" box="[180,465,1111,1136]" pageId="1" pageNumber="140" refString="Molineri, C. & Goitia, E. (2006) Description of the adult stage of Cercobrachys peruanicus (Ephemeroptera: Caenidae). Revista de la Sociedad Entomologica Argentina, 65, 63 - 67." type="journal article" year="2006">Molineri & Goitia (2006)</bibRefCitation>
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||
also examined.
|
||
</paragraph>
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<paragraph id="E60F36AFFFCAFF8EFF65E64B68D1FA8B" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
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Male. Length (mm): body, 2.3–3.8; forewing, 2.0–2.7; caudal filaments, 5.8–9.0. Ratio pedicel/scape, 1.7–2.0. Metanotum with posteromedian finger-like tubercle (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFD50E6A86970FB5E" box="[754,843,1183,1208]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Fig. 1B</figureCitation>
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). Foreleg with black dot on apex of tibia and tarsus segments 2–3 (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFEE7E6F36B9AFB3A" box="[325,417,1220,1245]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Fig. 1D</figureCitation>
|
||
). Ratios of length of body: foreleg: midleg: hindleg - 1: 0.6–0.8: 0.4: 0.4–0.5. Ratios of length of forefemur: tibia: tarsus - 1: 1.8–2.3: 1.5–1.7. Ratios of length of foretarsus segment I: II: III: IV:
|
||
<date id="920E106FFFCAFF8EFAFAE6DE6FADFAE6" box="[1368,1430,1256,1280]" bridgedPair="-" pageId="1" pageNumber="140">V - 1</date>
|
||
: 3.6–8.4: 3.0–3.4: 2.6–2.8: 1.8–2.0. Wing length/max width 1.8–2.3. Abdomen with thin and long vestiges of posterolateral projections on segments IV–VI; gray shading darker on terga I–II and VII–VIII. Genitalia (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFAC6E7186AE4FA8A" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Figs. 1G–H</figureCitation>
|
||
): ratios forceps length/subbasal width 6.8–8.0.
|
||
</paragraph>
|
||
<paragraph id="E60F36AFFFCAFF8EFF65E7406BC3F9C6" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
|
||
Female. Length (mm): body, 4.0–4.5; forewing, 3.4–3.7; caudal filaments, 1.5–1.7. Metanotum with dorsal finger-like tubercle (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFEDCE7AB6BEEFA52" box="[382,469,1436,1461]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Fig. 1E</figureCitation>
|
||
). Wing length/max width 2.3. Abdomen with vestiges of posterolateral projections on segments IV–VI; abdominal sternum IX produced distally, surpassing half of length of segment X but not reaching its apex; apex of sternum IX truncated with small median notch. Abdominal terga and sterna yellowish, widely washed with gray (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFECEE4306BD0F9C6" box="[364,491,1543,1568]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Figs. 1E–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E60F36AFFFCAFF8EFF65E41A6AC5F91E" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
|
||
Egg. Shape elongate-ovate (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFDA8E41B6861F9A2" box="[522,602,1580,1605]" captionStart="FIGURE 2. A – G" captionStartId="4.[151,250,1702,1724]" captionTargetBox="[191,1395,193,1680]" captionTargetId="figure@4.[191,1395,193,1681]" captionTargetPageId="4" captionText="FIGURE 2. A – G: Alloretochus sigillatus sp. nov.: A, male imago, head and thorax, lateral view; B, same, dorsal view of abdomen; C, same, detail of head; D, male genitalia, ventral view; E, female imago, dorsal view; F – G, eggs. H – I, A. peruanicus eggs." httpUri="https://zenodo.org/record/226221/files/figure.png" pageId="1" pageNumber="140">Figs. 2</figureCitation>
|
||
H–I), ratios total length/maximum width: 3.0–3.3. Polar cap 1/10–1/5 of whole egg in length, without tubercles. Chorion with 7–8 costae in lateral half. Costa asymmetrical in cross-section (overlapping adjacent inter-costal groove on one side only), costae covered with subcircular wart-like microsculpture (detail in
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFE11E4A06BCCF956" box="[435,503,1687,1712]" captionStart="FIGURE 2. A – G" captionStartId="4.[151,250,1702,1724]" captionTargetBox="[191,1395,193,1680]" captionTargetId="figure@4.[191,1395,193,1681]" captionTargetPageId="4" captionText="FIGURE 2. A – G: Alloretochus sigillatus sp. nov.: A, male imago, head and thorax, lateral view; B, same, dorsal view of abdomen; C, same, detail of head; D, male genitalia, ventral view; E, female imago, dorsal view; F – G, eggs. H – I, A. peruanicus eggs." httpUri="https://zenodo.org/record/226221/files/figure.png" pageId="1" pageNumber="140">
|
||
Fig.
|
||
<date id="920E106FFFCAFF8EFE45E4AF6BCCF956" box="[487,503,1688,1712]" pageId="1" pageNumber="140">2</date>
|
||
</figureCitation>
|
||
I). A thin layer that covers the egg (probably not part of the egg, but part of female body cavity) appeared partially detached in most of them, but always presented small circular tubercles (arrow in
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFF35E4E86AE2F91E" box="[151,217,1759,1784]" captionStart="FIGURE 2. A – G" captionStartId="4.[151,250,1702,1724]" captionTargetBox="[191,1395,193,1680]" captionTargetId="figure@4.[191,1395,193,1681]" captionTargetPageId="4" captionText="FIGURE 2. A – G: Alloretochus sigillatus sp. nov.: A, male imago, head and thorax, lateral view; B, same, dorsal view of abdomen; C, same, detail of head; D, male genitalia, ventral view; E, female imago, dorsal view; F – G, eggs. H – I, A. peruanicus eggs." httpUri="https://zenodo.org/record/226221/files/figure.png" pageId="1" pageNumber="140">Fig. 2</figureCitation>
|
||
H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEAA6524FFCAFF8CFF65E5336853FD2A" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140" type="discussion">
|
||
<paragraph id="E60F36AFFFCAFF8EFF65E5336E10F836" blockId="1.[151,1437,716,2001]" pageId="1" pageNumber="140">
|
||
<emphasis id="D4C4EABDFFCAFF8EFF65E5336B77F8FB" bold="true" box="[199,332,1796,1821]" pageId="1" pageNumber="140">Discussion.</emphasis>
|
||
<bibRefCitation id="82214B5EFFCAFF8EFEF6E5336841F8FA" author="Sun" box="[340,634,1796,1821]" pageId="1" pageNumber="140" refString="Sun, L. & MCCafferty. W. P. (2008) Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa, 1801, 1 - 239." type="journal article" year="2008">Sun & McCafferty (2008)</bibRefCitation>
|
||
differentiated the larvae of
|
||
<taxonomicName id="21B04D2CFFCAFF8EFC17E5336E78F8FA" box="[949,1091,1796,1820]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4C4EABDFFCAFF8EFC17E5336E78F8FA" box="[949,1091,1796,1820]" italics="true" pageId="1" pageNumber="140">Alloretochus</emphasis>
|
||
</taxonomicName>
|
||
from those of all other genera of
|
||
<taxonomicName id="21B04D2CFFCAFF8EFF16E5106B59F8A6" box="[180,354,1831,1856]" class="Insecta" family="Caenidae" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="subFamily" subFamily="Brachycercinae">Brachycercinae</taxonomicName>
|
||
by "having a maxillary palp segment 2 that is as long as about 2.3 X the length of the segment 1" (vs. no longer than 2.0 X in the others). In the only larva studied here (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFC66E57B6E25F882" box="[964,1054,1868,1893]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">Fig. 1A</figureCitation>
|
||
), a ready to molt male, maxillary palps roughly coincide with this ratio, with one having a longer segment 2 (2.56X the length of segment 1) than the other (2.25X the length of segment 1) (
|
||
<figureCitation id="7E8B2A2AFFCAFF8EFDFFE5A36890F84A" box="[605,683,1940,1965]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="1" pageNumber="140">
|
||
Fig.
|
||
<date id="920E106FFFCAFF8EFD36E5A36890F84A" box="[660,683,1940,1964]" pageId="1" pageNumber="140">1I</date>
|
||
</figureCitation>
|
||
). Other characters of this larva, including the diagnostic patch of setae on labial palp, are identical to those described by
|
||
<bibRefCitation id="82214B5EFFCAFF8EFCA3E58F6E1CF836" author="Sun" box="[769,1063,1976,2001]" pageId="1" pageNumber="140" refString="Sun, L. & MCCafferty. W. P. (2008) Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa, 1801, 1 - 239." type="journal article" year="2008">Sun & McCafferty (2008)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="B2CF6627FFC9FF8DFF35E5586B11F807" httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="2" pageNumber="141" targetBox="[208,1372,195,1864]" targetPageId="2">
|
||
<paragraph id="E60F36AFFFC9FF8DFF35E5586B11F807" blockId="2.[151,1436,1903,2017]" pageId="2" pageNumber="141">
|
||
<emphasis id="D4C4EABDFFC9FF8DFF35E5586B2FF862" bold="true" box="[151,276,1903,1925]" pageId="2" pageNumber="141">FIGURE 1.</emphasis>
|
||
A–I.
|
||
<taxonomicName id="21B04D2CFFC9FF8DFEF2E547687CF863" box="[336,583,1904,1925]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC9FF8DFEF2E547687CF863" box="[336,583,1904,1925]" italics="true" pageId="2" pageNumber="141">Alloretochus peruanicus</emphasis>
|
||
</taxonomicName>
|
||
: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E60F36AFFFC8FF8CFF65E2AF6B0BFEA6" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="142">
|
||
Among the characters listed for
|
||
<taxonomicName id="21B04D2CFFC8FF8CFD8CE2AF6887FF56" box="[558,700,152,176]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFD8CE2AF6887FF56" box="[558,700,152,176]" italics="true" pageId="3" pageNumber="142">Alloretochus</emphasis>
|
||
</taxonomicName>
|
||
adults (
|
||
<bibRefCitation id="82214B5EFFC8FF8CFCB6E2AF6E12FF56" author="Sun" box="[788,1065,151,176]" pageId="3" pageNumber="142" refString="Sun, L. & MCCafferty. W. P. (2008) Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa, 1801, 1 - 239." type="journal article" year="2008">Sun & McCafferty 2008</bibRefCitation>
|
||
), the following were also present in the males studied here: prosternum conically produced ventrally (slightly marked in some specimens), pronotum with slightly convex to nearly straight lateral margin, vestiges of posterolateral projections on abdominal segments IV–VI (in segments I–III poorly developed, not projected), and subgenital plate broadly emarginate and distally membranous.
|
||
</paragraph>
|
||
<paragraph id="E60F36AFFFC8FF8CFF65E37B6E3EFDFE" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="142">
|
||
The small male (body,
|
||
<quantity id="21489B4AFFC8FF8CFE46E37B6878FE82" box="[484,579,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="142" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, forewing, 2.0 mm; caudal filaments,
|
||
<quantity id="21489B4AFFC8FF8CFBB7E37B6E48FE82" box="[1045,1139,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="142" unit="mm" value="5.8">5.8 mm</quantity>
|
||
) collected in Colombian Amazonas and attributed here to
|
||
<taxonomicName id="21B04D2CFFC8FF8CFDB1E347688AFE61" box="[531,689,368,391]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFDB1E347688AFE61" box="[531,689,368,391]" italics="true" pageId="3" pageNumber="142">A. peruanicus</emphasis>
|
||
</taxonomicName>
|
||
presents certain differences in relation to males of
|
||
<taxonomicName id="21B04D2CFFC8FF8CFB5CE3476FA7FE61" box="[1278,1436,368,391]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFB5CE3476FA7FE61" box="[1278,1436,368,391]" italics="true" pageId="3" pageNumber="142">A. peruanicus</emphasis>
|
||
</taxonomicName>
|
||
from other localities, including the absence of gray shading or other marks on body (except the small black dots at apex of foretibia and tarsal segments,
|
||
<figureCitation id="7E8B2A2AFFC8FF8CFD9DE38068ADFE36" box="[575,662,439,464]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="3" pageNumber="142">Fig. 1D</figureCitation>
|
||
), absence of posteromedian projection on metanotum and vestiges of posterolateral projections on abdominal segments. However, the morphology of male genitalia of this individual (
|
||
<figureCitation id="7E8B2A2AFFC8FF8CFF3DE3C86ACDFDFE" box="[159,246,511,536]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="3" pageNumber="142">Fig. 1G</figureCitation>
|
||
) is very similar to that of other specimens of
|
||
<taxonomicName id="21B04D2CFFC8FF8CFD56E03769ABFDF1" box="[756,912,512,535]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFD56E03769ABFDF1" box="[756,912,512,535]" italics="true" pageId="3" pageNumber="142">A. peruanicus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="7E8B2A2AFFC8FF8CFC3DE3C869C3FDFE" box="[927,1016,511,536]" captionStart="FIGURE 1. A – I" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[208,1372,195,1864]" captionTargetId="figure@2.[208,1372,195,1865]" captionTargetPageId="2" captionText="FIGURE 1. A – I. Alloretochus peruanicus: A, larva, ventrolateral view; B male from Blanco river in Bolivia, lateral view; C, male from San Francisco river, Argentina, ventrolateral view; D, foreleg of male from Colombia; E, female (S. Francisco river, Argentina) dorsal view; F, same, ventral; G, male genitalia (Colombian specimen); H, male genitalia (Argentinean specimen); I, larval maxilla." httpUri="https://zenodo.org/record/226220/files/figure.png" pageId="3" pageNumber="142">Fig. 1H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E60F36AFFFC8FF8CFF65E0136E03FD63" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="142">
|
||
<taxonomicName id="21B04D2CFFC8FF8CFF65E0136BE7FDDA" box="[199,476,548,572]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFF65E0136BE7FDDA" box="[199,476,548,572]" italics="true" pageId="3" pageNumber="142">Alloretochus peruanicus</emphasis>
|
||
</taxonomicName>
|
||
shows a relatively wide range of variation in size and some ratios, as seen in the description above (for example the length of the second fore tarsal segment). This is not rare in the group; and other species may present similar intraspecific variations where specimens are abundant.
|
||
</paragraph>
|
||
<paragraph id="E60F36AFFFC8FF8CFF65E0A76853FD2A" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="142">
|
||
In general eggs of
|
||
<taxonomicName id="21B04D2CFFC8FF8CFE16E0A7686CFD41" box="[436,599,656,679]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFE16E0A7686CFD41" box="[436,599,656,679]" italics="true" pageId="3" pageNumber="142">A. peruanicus</emphasis>
|
||
</taxonomicName>
|
||
are similar to
|
||
<taxonomicName id="21B04D2CFFC8FF8CFCB7E0A769DAFD4E" box="[789,993,656,680]" class="Insecta" family="Caenidae" genus="Cercobrachys" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="fox">
|
||
<emphasis id="D4C4EABDFFC8FF8CFCB7E0A769DAFD4E" box="[789,993,656,680]" italics="true" pageId="3" pageNumber="142">Cercobrachys fox</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="21B04D2CFFC8FF8CFB84E0A76EB2FD4E" box="[1062,1161,656,680]" class="Insecta" family="Caenidae" genus="Cercobrachys" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="lilliei">
|
||
<emphasis id="D4C4EABDFFC8FF8CFB84E0A76EB2FD4E" box="[1062,1161,656,680]" italics="true" pageId="3" pageNumber="142">C. lilliei</emphasis>
|
||
</taxonomicName>
|
||
(figures 570 and 572, respectively, in
|
||
<bibRefCitation id="82214B5EFFC8FF8CFEE5E0836867FD2A" author="Sun" box="[327,604,692,717]" pageId="3" pageNumber="142" refString="Sun, L. & MCCafferty. W. P. (2008) Cladistics, classification and identification of the brachycercine mayflies (Insecta: Ephemeroptera: Caenidae). Zootaxa, 1801, 1 - 239." type="journal article" year="2008">Sun & McCafferty 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEAA6524FFC8FF8CFF65E0E06858FCDE" pageId="3" pageNumber="142" type="distribution">
|
||
<paragraph id="E60F36AFFFC8FF8CFF65E0E06858FCDE" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="142">
|
||
<emphasis id="D4C4EABDFFC8FF8CFF65E0E06B5BFD16" bold="true" box="[199,352,727,752]" pageId="3" pageNumber="142">Distribution.</emphasis>
|
||
<collectingCountry id="9EA7763FFFC8FF8CFED0E0E06BD9FD16" box="[370,482,727,752]" name="Argentina" pageId="3" pageNumber="142">Argentina</collectingCountry>
|
||
(Jujuy),
|
||
<collectingCountry id="9EA7763FFFC8FF8CFDFFE0E0688BFD16" box="[605,688,727,752]" name="Bolivia" pageId="3" pageNumber="142">Bolivia</collectingCountry>
|
||
(Tarija, Santa Cruz),
|
||
<collectingCountry id="9EA7763FFFC8FF8CFC6DE0E06E06FD16" box="[975,1085,727,752]" name="Colombia" pageId="3" pageNumber="142">Colombia</collectingCountry>
|
||
(Amazonas),
|
||
<collectingCountry id="9EA7763FFFC8FF8CFB50E0EF6F1EFD09" box="[1266,1317,728,751]" name="Peru" pageId="3" pageNumber="142">Peru</collectingCountry>
|
||
(Loreto).
|
||
<taxonomicName id="21B04D2CFFC8FF8CFF35E0CB6B8BFCF2" box="[151,432,764,788]" class="Insecta" family="Caenidae" genus="Alloretochus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="peruanicus">
|
||
<emphasis id="D4C4EABDFFC8FF8CFF35E0CB6B8BFCF2" box="[151,432,764,788]" italics="true" pageId="3" pageNumber="142">Alloretochus peruanicus</emphasis>
|
||
</taxonomicName>
|
||
shows a wide distributional range including medium sized piedmont rivers as the Pilcomayo to larger ones as the Amazon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |