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<document id="797159713DCDD5CB80BFC10BF1927253" ID-DOI="10.5281/zenodo.182603" ID-GBIF-Dataset="5d95a7c8-6bcf-4fbc-93ff-646498a5a96d" ID-ISSN="1175-5326" ID-Zenodo-Dep="182603" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459871429437" checkinUser="plazi" docAuthor="Mondaca, José &amp; Smith, Andrew B. T." docDate="2008" docId="5F60A91F742C76353DD8FD34FD3571D6" docLanguage="en" docName="zt01794p048.pdf" docOrigin="Zootaxa 1794" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Bolborhinum trilobulicorne Mondaca &amp; Smith, sp. nov." docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="A359D167743C76273D4FFFD6FF92710C" masterDocTitle="A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="17" updateTime="1698226675509" updateUser="plazi">
<mods:mods id="E138675A0E9DB8543C489A487C562885" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF9AAEF6603FA3CDE460FCE4AB3BC50C">
<mods:title id="21914A863B9D5BAB9459525FF621806E">A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
</mods:titleInfo>
<mods:name id="941786151A984FDEA3513AD3ABB90A39" type="personal">
<mods:role id="9E1DA76A8F73FAE885EC05B715514672">
<mods:roleTerm id="DBAE35E6869B690A81002F5CEECF7E6B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1269903463138B3D7B01521E2DAF545">Mondaca, José</mods:namePart>
</mods:name>
<mods:name id="0E832CA59204869134B59E8CF27ED698" type="personal">
<mods:role id="A440FA79C5E261E48BCC6DB58ED1040C">
<mods:roleTerm id="8C421FF64B6897FA957A45B9B26D5CDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02BA88E58BC4DFEF54B4BA2F77C0B88C">Smith, Andrew B. T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2C180E656105B218CB11E6BC807BF454">text</mods:typeOfResource>
<mods:relatedItem id="EE623AD615F260B91D8AE25201B820F9" type="host">
<mods:titleInfo id="1589EB30368759EAF104DDB46F8B1525">
<mods:title id="83A3C69ACE85D4CAB3156145921A666F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AF110D5636C38E64D8C7EA49458211A3">
<mods:date id="19F690503D24A4E7DA4E3C02450BE908">2008</mods:date>
<mods:detail id="BF9DBAE546621216D7ED1579297C359E" type="volume">
<mods:number id="AD9D78B6040FC00D6181A0BC319FD6A1">1794</mods:number>
</mods:detail>
<mods:extent id="1FDA69907C83B5C1CF71DCE6832C90D0" unit="page">
<mods:start id="7FC145C56D9762D8F8FD785051BE2D35">1</mods:start>
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</mods:extent>
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<mods:classification id="29DC9E078F0FB941E06172B962B559E6">journal article</mods:classification>
<mods:identifier id="C96B0F8F5A93F47D47DBF08BF66033FB" type="DOI">10.5281/zenodo.182603</mods:identifier>
<mods:identifier id="A174CFB74633BBCAB57257895380E5A9" type="GBIF-Dataset">5d95a7c8-6bcf-4fbc-93ff-646498a5a96d</mods:identifier>
<mods:identifier id="3A1EC3702214F774E876C62191C6FCEB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E930DCA695F6E90DF192A36215C4E090" type="Zenodo-Dep">182603</mods:identifier>
</mods:mods>
<treatment id="5F60A91F742C76353DD8FD34FD3571D6" ID-DOI="http://doi.org/10.5281/zenodo.6230995" ID-GBIF-Taxon="119369639" ID-Zenodo-Dep="6230995" LSID="urn:lsid:zoobank.org:act:447729D6-E944-4E88-A224-16C23EE0449B" httpUri="http://treatment.plazi.org/id/5F60A91F742C76353DD8FD34FD3571D6" lastPageId="18" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="9FD34B82742C76373DD8FD34FE1E7213" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="D7761809742C76373DD8FD34FCD073F0" blockId="16.[151,834,738,799]" box="[151,834,738,764]" pageId="16" pageNumber="17">
<heading id="8C3EAF65742C76373DD8FD34FCD073F0" bold="true" box="[151,834,738,764]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="E5BDC41B742C76373DD8FD34FCD073F0" bold="true" box="[151,834,738,764]" pageId="16" pageNumber="17">
<taxonomicName id="10C9638A742C76373DD8FD34FD4A73F0" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[151,728,738,764]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne" status="sp. nov.">
<emphasis id="E5BDC41B742C76373DD8FD34FE7573F0" bold="true" box="[151,487,738,764]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="FE8E7960742C76373FABFD34FCD073F0" box="[740,834,738,764]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D7761809742C76373DD8FCD3FE1E7213" blockId="16.[151,834,738,799]" box="[151,396,773,799]" pageId="16" pageNumber="17">
(
<figureCitation id="4FF2048C742C76373DD0FCD3FF6D7213" box="[159,255,773,799]" captionStart="FIGURE 11" captionStartId="27.[151,255,1362,1386]" captionText="FIGURE 11. Bolborhinum trilobulicorne male in lateral view." pageId="16" pageNumber="17">Figs. 11</figureCitation>
,
<figureCitation id="4FF2048C742C76373C43FCD3FEBE7213" box="[268,300,773,799]" captionStart="FIGURE 12" captionStartId="27.[151,255,1394,1418]" captionText="FIGURE 12. Bolborhinum trilobulicorne female in lateral view." pageId="16" pageNumber="17">12</figureCitation>
,
<figureCitation id="4FF2048C742C76373C77FCD3FECA7213" box="[312,344,773,799]" captionStart="FIGURE 34" captionStartId="34.[151,255,1878,1902]" captionTargetBox="[285,1318,211,1842]" captionTargetId="figure@34.[264,1323,194,1857]" captionTargetPageId="34" captionText="FIGURE 34. Bolborhinum trilobulicorne male in dorsal view. FIGURE 35. Bolborhinum trilobulicorne female in dorsal view. FIGURE 36. Bolborhinum tubericeps male in dorsal view." httpUri="https://zenodo.org/record/182609/files/figure.png" pageId="16" pageNumber="17">34</figureCitation>
, 35)
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742C76363DD8FC99FA90723E" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="D7761809742C76373DD8FC99FD5B76AE" blockId="16.[151,1437,847,2034]" pageId="16" pageNumber="17">
<emphasis id="E5BDC41B742C76373DD8FC99FEF07265" bold="true" box="[151,354,847,873]" pageId="16" pageNumber="17">
<typeStatus id="0872A6AB742C76373DD8FC99FF467265" box="[151,212,847,873]" pageId="16" pageNumber="17">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="0872A6AB742C76373C24FC86FE4A7266" box="[363,472,848,874]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
male and 43
<typeStatus id="0872A6AB742C76373F36FC86FD7B7266" box="[633,745,848,874]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
(
<specimenCount id="C1CFD380742C76373FB5FC86FCFA7266" box="[762,872,848,874]" pageId="16" pageNumber="17" type="male">41 males</specimenCount>
,
<specimenCount id="C1CFD380742C76373E3AFC86FC7A7266" box="[885,1000,848,874]" pageId="16" pageNumber="17" type="female">2 females</specimenCount>
) deposited in 14 institutions and private collections.
<typeStatus id="0872A6AB742C76373C23FCAEFE4B729E" box="[364,473,888,914]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
male at
<collectionCode id="B1D880CC742C76373F1CFCAEFD39729E" box="[595,683,888,914]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
labeled: a) “
<collectingCountry id="AFDE5899742C76373E19FCAEFC09729E" box="[854,923,888,914]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
, VII Reg., Maule / R.N. Los Queules /
<geoCoordinate id="B2FD7ECE742C76373DD8FC76FEB172B6" box="[151,291,928,954]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-35.987778">35º59’16’’S</geoCoordinate>
<geoCoordinate id="B2FD7ECE742C76373C64FC76FE4B72B6" box="[299,473,928,954]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="-72.695724">72º41’44.6’’W</geoCoordinate>
/
<date id="A3773EC9742C76373CBFFC76FDEC72B6" box="[496,638,928,954]" pageId="16" pageNumber="17" value="2004-11-30">30-XI-2004</date>
, J. Mondaca &amp; M.J. Paulsen col.” (typeset), b) “Southern Neotropical Scarabs/ database #JM2000486 ɗ /
<taxonomicName id="10C9638A742C76373F28FC1EFB0272EE" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[615,1168,968,994]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373F28FC1EFC3F72ED" box="[615,941,968,993]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/ DET: J. Mondaca E. 2007” (typeset), c) “
<taxonomicName id="10C9638A742C76373CC7FC26FC3F7506" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[392,941,1008,1034]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373CC7FC26FD5F7505" box="[392,717,1008,1033]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/
<typeStatus id="0872A6AB742C76373E8AFC26FBF27506" box="[965,1120,1008,1034]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
ɗ (red label, typeset). One male
<typeStatus id="0872A6AB742C76373D99FBCEFEA9753E" box="[214,315,1048,1074]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
at
<collectionCode id="B1D880CC742C76373C11FBCEFE24753E" box="[350,438,1048,1074]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
labeled: a) “
<collectingCountry id="AFDE5899742C76373F07FBCEFD1F753E" box="[584,653,1048,1074]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
, VII Reg., Curanipe / R.N. Los Queules / 13/
<date id="A3773EC9742C763739D7FBCEFAB4753E" box="[1176,1318,1048,1074]" pageId="16" pageNumber="17" value="2001-11-25">25-XI-2001</date>
, A. Fierro col.” (typeset), b) “Southern Neotropical Scarabs / database #JM2000490 ɗ /
<taxonomicName id="10C9638A742C7637391CFB96FEFD758E" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C7637391CFB96FA0E7555" box="[1107,1436,1088,1113]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/ DET: J. Mondaca E. 2007” (typeset), c) “
<taxonomicName id="10C9638A742C76373E26FBBEFA1E758E" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[873,1420,1128,1154]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373E26FBBEFB3C758D" box="[873,1198,1128,1153]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/
<typeStatus id="0872A6AB742C76373DD8FB46FEBB75A6" box="[151,297,1168,1194]" pageId="16" pageNumber="17" type="paratype">PARATYPE</typeStatus>
ɗ (red label, typeset). Eleven male
<typeStatus id="0872A6AB742C76373F85FB46FCA875A6" box="[714,826,1168,1194]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
(two at
<collectionCode id="B1D880CC742C76373ED3FB46FC6675A6" LSID="urn:lsid:biocol.org:col:34795" box="[924,1012,1168,1194]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="17">FMNH</collectionCode>
, two at
<collectionCode id="B1D880CC742C7637391FFB46FB3A75A6" box="[1104,1192,1168,1194]" pageId="16" pageNumber="17">UMCE</collectionCode>
, four at
<collectionCode id="B1D880CC742C76373846FB46FAF675A6" box="[1289,1380,1168,1194]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
, two at
<collectionCode id="B1D880CC742C76373DFCFB6EFE9475DE" box="[179,262,1208,1234]" httpUri="http://grbio.org/cool/jgvc-yfsb" name="Universidad de Concepcion, Museo de Zoologia" pageId="16" pageNumber="17">UCCC</collectionCode>
, one at
<collectionCode id="B1D880CC742C76373C11FB6EFE2775DE" LSID="urn:lsid:biocol.org:col:34868" box="[350,437,1208,1234]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="16" pageNumber="17">UNSM</collectionCode>
) labeled: a) “
<collectingCountry id="AFDE5899742C76373F00FB6EFD0675DE" box="[591,660,1208,1234]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
, VII Reg., Maule / R.N. Los Queules /
<geoCoordinate id="B2FD7ECE742C76373914FB6EFB7A75DE" box="[1115,1256,1208,1234]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-35.987778">35º59’16’’S</geoCoordinate>
<geoCoordinate id="B2FD7ECE742C763739A1FB6EFA0E75DE" box="[1262,1436,1208,1234]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="-72.695724">72º41’44.6’’W</geoCoordinate>
/
<date id="A3773EC9742C76373DE3FB36FEA875F6" box="[172,314,1248,1274]" pageId="16" pageNumber="17" value="2004-11-30">30-XI-2004</date>
, J. Mondaca &amp; M.J. Paulsen col.” (typeset), b) “Southern Neotropical Scarabs / database [#JM2000470-#JM2000472 (
<collectionCode id="B1D880CC742C76373CB6FADEFDC7742E" LSID="urn:lsid:biocol.org:col:34795" box="[505,597,1288,1314]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="17">FMNH</collectionCode>
); #JM2000473-#JM2000474 (
<collectionCode id="B1D880CC742C76373E84FADEFBB5742E" box="[971,1063,1288,1314]" pageId="16" pageNumber="17">UMCE</collectionCode>
); #JM2000475, #JM2000477- #JM2000478-#JM2000479 (
<collectionCode id="B1D880CC742C76373CA5FAE6FDDB7446" box="[490,585,1328,1354]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
); JM2000487-#JM2000488 (
<collectionCode id="B1D880CC742C76373EE9FAE6FC6F7446" box="[934,1021,1328,1354]" httpUri="http://grbio.org/cool/jgvc-yfsb" name="Universidad de Concepcion, Museo de Zoologia" pageId="16" pageNumber="17">UCCC</collectionCode>
); #JM2000489 (
<collectionCode id="B1D880CC742C7637398BFAE6FA8D7446" LSID="urn:lsid:biocol.org:col:34868" box="[1220,1311,1328,1354]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="16" pageNumber="17">UNSM</collectionCode>
)] ɗ /
<taxonomicName id="10C9638A742C76373826FAE6FD01747E" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373826FAE6FE21747D" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/ DET: J. Mondaca E. 2007” (typeset), c) “
<taxonomicName id="10C9638A742C763739DFFA8EFE2A7496" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C763739DFFA8EFF487495" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/
<typeStatus id="0872A6AB742C76373C80FA56FDF37496" box="[463,609,1408,1434]" pageId="16" pageNumber="17" type="paratype">PARATYPE</typeStatus>
ɗ (red label, typeset). Fourteen male and one female
<typeStatus id="0872A6AB742C7637399EFA56FAD37496" box="[1233,1345,1408,1434]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
(five at
<collectionCode id="B1D880CC742C76373DD8FA7EFF6374CE" box="[151,241,1448,1474]" pageId="16" pageNumber="17">CMNC</collectionCode>
, two at
<collectionCode id="B1D880CC742C76373C02FA7EFE0474CE" box="[333,406,1448,1474]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="16" pageNumber="17">CNCI</collectionCode>
, two at
<collectionCode id="B1D880CC742C76373CBCFA7EFDC974CE" box="[499,603,1448,1474]" pageId="16" pageNumber="17">IADIZA</collectionCode>
, two at
<collectionCode id="B1D880CC742C76373FF8FA7EFC9D74CE" LSID="urn:lsid:biocol.org:col:34802" box="[695,783,1448,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="17">LACM</collectionCode>
, two at
<collectionCode id="B1D880CC742C76373E23FA7EFC5574CE" box="[876,967,1448,1474]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="17">MACN</collectionCode>
, one at
<collectionCode id="B1D880CC742C7637396DFA7EFBE874CE" LSID="urn:lsid:biocol.org:col:34868" box="[1058,1146,1448,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="16" pageNumber="17">UNSM</collectionCode>
, one
<collectionCode id="B1D880CC742C763739F8FA7EFA8374CE" box="[1207,1297,1448,1474]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
) labeled: a) “
<collectingCountry id="AFDE5899742C76373DEEFA06FF7474E6" box="[161,230,1488,1514]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
, VII Reg., Maule / R.N. Los Queules /
<geoCoordinate id="B2FD7ECE742C76373F85FA06FCC474E6" box="[714,854,1488,1514]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-35.987778">35º59’16’’S</geoCoordinate>
<geoCoordinate id="B2FD7ECE742C76373E2FFA06FB9C74E6" box="[864,1038,1488,1514]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="-72.695724">72º41’44.6’’W</geoCoordinate>
/
<date id="A3773EC9742C76373965FA06FB2574E6" box="[1066,1207,1488,1514]" pageId="16" pageNumber="17" value="2004-11-30">30-XI-2004</date>
, M.J. Paulsen &amp; J. Mondaca col.” (typeset), b) “Southern Neotropical Scarabs/ database [#JM2000466-#JM2000467,
<collectingCountry id="AFDE5899742C76373DD8F9F6FF2E7736" box="[151,188,1568,1594]" name="American Samoa" pageId="16" pageNumber="17">AS</collectingCountry>
2609509, #
<collectingCountry id="AFDE5899742C76373C1DF9F6FEE47736" box="[338,374,1568,1594]" name="American Samoa" pageId="16" pageNumber="17">AS</collectingCountry>
2610458-#
<collectingCountry id="AFDE5899742C76373CBAF9F6FD8B7736" box="[501,537,1568,1594]" name="American Samoa" pageId="16" pageNumber="17">AS</collectingCountry>
2610459 (
<collectionCode id="B1D880CC742C76373FEBF9F6FC937736" box="[676,769,1568,1594]" pageId="16" pageNumber="17">CMNC</collectionCode>
); #JM2000468-#JM2000469 (
<collectionCode id="B1D880CC742C763739C7F9F6FB467736" box="[1160,1236,1568,1594]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="16" pageNumber="17">CNCI</collectionCode>
); #JM2000471, #JM2000480 (
<collectionCode id="B1D880CC742C76373C1AF99EFE53776E" box="[341,449,1608,1634]" pageId="16" pageNumber="17">IADIZA</collectionCode>
); #JM2000481-#JM2000482 (
<collectionCode id="B1D880CC742C76373E04F99EFC35776E" LSID="urn:lsid:biocol.org:col:34802" box="[843,935,1608,1634]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="17">LACM</collectionCode>
); #JM2000483-#JM2000484 (
<collectionCode id="B1D880CC742C7637387EF99EFA02776E" box="[1329,1424,1608,1634]" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="17">MACN</collectionCode>
); #JM2000485 (
<collectionCode id="B1D880CC742C76373C0BF9A6FE0D7786" LSID="urn:lsid:biocol.org:col:34868" box="[324,415,1648,1674]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="16" pageNumber="17">UNSM</collectionCode>
); #JM2000491 (
<collectionCode id="B1D880CC742C76373F2BF9A6FD507786" box="[612,706,1648,1674]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="16" pageNumber="17">MNNC</collectionCode>
)] [ɗ or Ψ] /
<taxonomicName id="10C9638A742C76373E16F9A6FA1B7786" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[857,1417,1648,1674]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373E16F9A6FB337785" box="[857,1185,1648,1673]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/ DET: J. Mondaca E. 2007” (typeset), c) “
<taxonomicName id="10C9638A742C76373FCAF94EFB3C77BE" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" box="[645,1198,1688,1714]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373FCAF94EFC5877BD" box="[645,970,1688,1713]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/
<typeStatus id="0872A6AB742C76373988F94EFACB77BE" box="[1223,1369,1688,1714]" pageId="16" pageNumber="17" type="paratype">PARATYPE</typeStatus>
[ɗ or Ψ] (red label, typeset). Fifteen male and one female
<typeStatus id="0872A6AB742C76373E4EF916FCE377D6" box="[769,881,1728,1754]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
(two at
<collectionCode id="B1D880CC742C76373E99F916FBB677D6" box="[982,1060,1728,1754]" pageId="16" pageNumber="17">ABTS</collectionCode>
, one at
<collectionCode id="B1D880CC742C763739CEF916FB5B77D6" box="[1153,1225,1728,1754]" pageId="16" pageNumber="17">JEBC</collectionCode>
, twelve at
<collectionCode id="B1D880CC742C76373805F916FA0A77D6" box="[1354,1432,1728,1754]" pageId="16" pageNumber="17">JMEC</collectionCode>
, one at
<collectionCode id="B1D880CC742C76373DABF93EFEA9760E" box="[228,315,1768,1794]" httpUri="http://grbio.org/cool/mqd5-c2qe" name="Universidad de Chile" pageId="16" pageNumber="17">MEUC</collectionCode>
) labeled: a) “
<collectingCountry id="AFDE5899742C76373C99F93EFD89760E" box="[470,539,1768,1794]" name="Chile" pageId="16" pageNumber="17">Chile</collectingCountry>
, VII Reg., Maule / R.N. Los Queules/ 1/
<date id="A3773EC9742C76373EBEF93EFBEF760E" box="[1009,1149,1768,1794]" pageId="16" pageNumber="17" value="2003-12-02">2-XII-2003</date>
, J. Mondaca col.” (typeset), b) “Southern Neotropical Scarabs / database [#JM2000492-#JM2000493 (
<collectionCode id="B1D880CC742C7637397BF8C6FB177626" box="[1076,1157,1808,1834]" pageId="16" pageNumber="17">ABTS</collectionCode>
); #JM2000494 (
<collectionCode id="B1D880CC742C7637380BF8C6FA1D7626" box="[1348,1423,1808,1834]" pageId="16" pageNumber="17">JEBC</collectionCode>
); #JM2000495-#JM2000505, #JM2000507 (
<collectionCode id="B1D880CC742C76373FC2F8EEFD4C765E" box="[653,734,1848,1874]" pageId="16" pageNumber="17">JMEC</collectionCode>
); #JM2000506 ɗ (
<collectionCode id="B1D880CC742C76373EFDF8EEFB9F765E" box="[946,1037,1848,1874]" httpUri="http://grbio.org/cool/mqd5-c2qe" name="Universidad de Chile" pageId="16" pageNumber="17">MEUC</collectionCode>
)] [ɗ or Ψ] /
<taxonomicName id="10C9638A742C763739DEF8EEFE2F7676" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C763739DEF8EEFF487675" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/ DET: J. Mondaca E. 2007” (typeset), c) “
<taxonomicName id="10C9638A742C76373E8DF8B6FF4E76AE" authority="Mondaca &amp; Smith" authorityName="Mondaca &amp; Smith" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742C76373E8DF8B6FA9A7675" box="[962,1288,1888,1913]" italics="true" pageId="16" pageNumber="17">Bolborhinum trilobulicorne</emphasis>
Mondaca &amp; Smith
</taxonomicName>
/
<typeStatus id="0872A6AB742C76373DA3F85EFEEC76AE" box="[236,382,1928,1954]" pageId="16" pageNumber="17" type="paratype">PARATYPE</typeStatus>
[ɗ or Ψ] (red label, typeset).
</paragraph>
<paragraph id="D7761809742C76363D89F879FA90723E" blockId="16.[151,1437,847,2034]" lastBlockId="17.[151,1437,152,2018]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="E5BDC41B742C76373D89F879FEAA76C5" bold="true" box="[198,312,1967,1993]" pageId="16" pageNumber="17">
<typeStatus id="0872A6AB742C76373D89F879FEAA76C5" box="[198,312,1967,1993]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
. Male.
<emphasis id="E5BDC41B742C76373CD6F866FE7876C5" box="[409,490,1968,1993]" italics="true" pageId="16" pageNumber="17">Length</emphasis>
: 15.0 mm.
<emphasis id="E5BDC41B742C76373F34F866FD2C76C5" box="[635,702,1968,1993]" italics="true" pageId="16" pageNumber="17">Width</emphasis>
: 8.0 mm.
<emphasis id="E5BDC41B742C76373E0DF866FC1476C5" box="[834,902,1968,1993]" italics="true" pageId="16" pageNumber="17">Color</emphasis>
: Dorsally shiny black, legs and venter dark brown.
<emphasis id="E5BDC41B742C76373DBBF80EFEA676FD" box="[244,308,2008,2033]" italics="true" pageId="16" pageNumber="17">Head</emphasis>
: With two horns; clypeal horn long, very robust, inclined anteriorly; apex widened, trilobed, with basal face excavated; frontal horn vertical, slightly curved basally at apex; apex bifurcate in “Y” shape. Frons and vertex smooth, impunctate. Labrum widely emarginated; dorsal surface punctate to rugose.
<emphasis id="E5BDC41B742D76363805FF16FF51700D" italics="true" pageId="17" pageNumber="18">Pronotum</emphasis>
: With deep, narrow anterior excavation that surrounds apical margin of the pronotum. Behind this excavation are a pair of prominent protuberances on each side of midline divided vertically by a furrow in the pronotal declivity; lateral edge of pronotum projected anteriorly in well-developed individuals; posterosuperior ridge transverse, arcuate posteriorly, dorsomedially with longitudinal furrow that reaches the basal margin; dorsal surface with sparse punctures, more concentrated behind pronotal protuberances. Laterally with large, dense punctures concentrated around the lateral fovea, at the apical and basal angles, and along the lateral margin.
<taxonomicName id="10C9638A742D76363C60FE0EFDBD70FE" authority="Triangular" authorityName="Triangular" box="[303,559,472,498]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="E5BDC41B742D76363C60FE0EFE3170FD" box="[303,419,472,497]" italics="true" pageId="17" pageNumber="18">Scutellum</emphasis>
: Triangular
</taxonomicName>
in shape, wider than long, surface with fine punctures.
<emphasis id="E5BDC41B742D76363982FE0EFABC70FD" box="[1229,1326,472,497]" italics="true" pageId="17" pageNumber="18">Elytron:</emphasis>
Apically with impressed striae; punctures small, separated by one puncture diameter; sutural interstriae wide, dorsally smooth, impunctate.
<emphasis id="E5BDC41B742D76363CDEFDFEFE4B734D" box="[401,473,552,577]" italics="true" pageId="17" pageNumber="18">Ve nt e r</emphasis>
: Surface covered by long, yellowish-brown, moderately dense setae; abdominal sternites moderately setose, covered with dense punctures. Metasternum densely punctate, with median longitudinal carina; metasternal process anteriorly truncate, sparsely setose.
<emphasis id="E5BDC41B742D76363EAFFDAEFB85739D" box="[992,1047,632,657]" italics="true" pageId="17" pageNumber="18">Legs</emphasis>
: Protibia wide, with six teeth on outer margin; protibial spur long, subparallel, acuminate at apex, protruding slightly past the second protarsomere. Mesocoxae contiguous. Mesotibia strongly thickened; subapical transverse carina well developed, surrounded by acute spines; apex with two apical spurs; spurs long, asymmetrical. Metatibia with subapical carina as in mesotibia, surrounded by long, acute spines.
<emphasis id="E5BDC41B742D76363E7BFCCEFC4D723D" box="[820,991,792,817]" italics="true" pageId="17" pageNumber="18">Male genitalia</emphasis>
: Aedeagus as in
<figureCitation id="4FF2048C742D763639E8FCCEFB6C723E" box="[1191,1278,792,818]" captionStart="FIGURE 53" captionStartId="39.[151,255,1059,1083]" captionText="FIGURE 53. Bolborhinum male genitalia. FIGURE 54. Pereirabolbus male genitalia. FIGURE 55. Zefevazia male genitalia." pageId="17" pageNumber="18">Fig. 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76363D89FCE9FE5075CE" pageId="17" pageNumber="18" type="description">
<paragraph id="D7761809742D76363D89FCE9FCBB72DE" blockId="17.[151,1437,152,2018]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89FCE9FEAB7255" bold="true" box="[198,313,831,857]" pageId="17" pageNumber="18">Variation</emphasis>
.
<specimenCount id="C1CFD380742D76363C02FC96FE3F7256" box="[333,429,832,858]" pageId="17" pageNumber="18" type="male">41 male</specimenCount>
and
<specimenCount id="C1CFD380742D76363CA5FC96FDC07256" box="[490,594,832,858]" pageId="17" pageNumber="18" type="female">2 female</specimenCount>
<typeStatus id="0872A6AB742D76363F14FC96FD5D7256" box="[603,719,832,858]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
. Length 12.0–14.8 mm, width 6.9–8.0 mm. Variation in the specimens examined is mainly in size. Some male
<typeStatus id="0872A6AB742D76363FBEFCBEFCF3728E" box="[753,865,872,898]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
differ from the
<typeStatus id="0872A6AB742D76363950FCBEFB17728E" box="[1055,1157,872,898]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
in the weaker development of the cephalic horns, pronotal protuberances, and pronotal excavation, all of which are sexually dimorphic characters present only in the males of this species.
</paragraph>
<paragraph id="D7761809742D76363D89FC36FE5075CE" blockId="17.[151,1437,152,2018]" pageId="17" pageNumber="18">
Female.
<emphasis id="E5BDC41B742D76363C60FC36FE1872F5" box="[303,394,992,1017]" italics="true" pageId="17" pageNumber="18">Length:</emphasis>
14.5 mm.
<emphasis id="E5BDC41B742D76363F5CFC36FDC472F5" box="[531,598,992,1017]" italics="true" pageId="17" pageNumber="18">Width</emphasis>
: 8.0 mm. Similar to male
<typeStatus id="0872A6AB742D76363EEFFC36FB9472F6" box="[928,1030,992,1018]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
except in the following respects:
<emphasis id="E5BDC41B742D76363DD8FBDEFF44752D" box="[151,214,1032,1057]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
: Surface rugulose and rugopunctate, anteriorly on the clypeal region with a pair of short carinae that converge distally. Clypeus truncate, elevated apically. Frons with an obsolete, bifid tubercle.
<emphasis id="E5BDC41B742D7636399AFBE6FADB7545" box="[1237,1353,1072,1097]" italics="true" pageId="17" pageNumber="18">Pronotum</emphasis>
: Without anteromedian excavation, only with small depression located immediately behind the apical margin; surface with larger punctures concentrated laterally, at medial furrow, and on pronotal depression.
<emphasis id="E5BDC41B742D7636380BFB56FF6D75CD" italics="true" pageId="17" pageNumber="18">Female genitalia</emphasis>
: Not diagnostic.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76363D89FB19FCBD741E" pageId="17" pageNumber="18" type="etymology">
<paragraph id="D7761809742D76363D89FB19FCBD741E" blockId="17.[151,1437,152,2018]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89FB19FED975E5" bold="true" box="[198,331,1231,1257]" pageId="17" pageNumber="18">Etymology</emphasis>
. The name &quot;
<taxonomicName id="10C9638A742D76363CAFFB06FD1175E5" box="[480,643,1232,1257]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742D76363CAFFB06FD1175E5" box="[480,643,1232,1257]" italics="true" pageId="17" pageNumber="18">trilobulicorne</emphasis>
</taxonomicName>
&quot; refers to the three lobes on the apex of the clypeal horn in the male of this species, which is a unique character in this genus.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76363D89FAC9FD4C770E" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="D7761809742D76363D89FAC9FD4C770E" blockId="17.[151,1437,152,2018]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89FAC9FEAD7435" bold="true" box="[198,319,1311,1337]" pageId="17" pageNumber="18">Diagnosis</emphasis>
.
<taxonomicName id="10C9638A742D76363C02FAF6FD037435" box="[333,657,1312,1337]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trilobulicorne">
<emphasis id="E5BDC41B742D76363C02FAF6FD037435" box="[333,657,1312,1337]" italics="true" pageId="17" pageNumber="18">Bolborhinum trilobulicorne</emphasis>
</taxonomicName>
is distinguished from other species of
<taxonomicName id="10C9638A742D7636391AFAF6FB7D7435" box="[1109,1263,1312,1337]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="E5BDC41B742D7636391AFAF6FB7D7435" box="[1109,1263,1312,1337]" italics="true" pageId="17" pageNumber="18">Bolborhinum</emphasis>
</taxonomicName>
by the following combination of characters: Head with two robust cephalic horns; clypeal horn directed anteriorly, with apex trilobed; frontal horn vertical, bifurcate distally, slightly curved posteriorly. Pronotum with deep, narrow, apical excavation expanded laterally to the apical angle; excavation with a pair of prominent protuberances and the lateral border projected apically; posterosuperior ridge transverse, medially with weakly depressed longitudinal furrow that reaches the basal margin.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76363D89F9D9FB0C775E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="D7761809742D76363D89F9D9FA137726" blockId="17.[151,1437,152,2018]" box="[198,1409,1551,1578]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89F9D9FECD7725" bold="true" box="[198,351,1551,1577]" pageId="17" pageNumber="18">Distribution</emphasis>
(
<figureCitation id="4FF2048C742D76363C20F9C6FE557726" box="[367,455,1552,1578]" captionStart="FIGURE 62" captionStartId="46.[151,255,1747,1771]" captionTargetBox="[152,1424,199,1713]" captionTargetId="figure@46.[151,1436,194,1726]" captionTargetPageId="46" captionText="FIGURE 62. Bolborhinum trilobulicorne distribution." httpUri="https://zenodo.org/record/182617/files/figure.png" pageId="17" pageNumber="18">Fig. 62</figureCitation>
).
<emphasis id="E5BDC41B742D76363C93F9D9FC5A7725" bold="true" box="[476,968,1551,1577]" pageId="17" pageNumber="18">
<collectingCountry id="AFDE5899742D76363C93F9D9FDA87725" box="[476,570,1551,1577]" name="Chile" pageId="17" pageNumber="18">CHILE</collectingCountry>
(44). VII Región del Maule (44)
</emphasis>
: Reserva Nacional Los Queules (44).
</paragraph>
<paragraph id="D7761809742D76363D89F9E1FB0C775E" blockId="17.[151,1437,152,2018]" box="[198,1182,1591,1618]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89F9E1FE43775D" bold="true" box="[198,465,1591,1617]" pageId="17" pageNumber="18">Specimens examined.</emphasis>
<specimenCount id="C1CFD380742D76363C90F9EEFDEC775E" box="[479,638,1592,1618]" pageId="17" pageNumber="18" type="generic">44 specimens</specimenCount>
were examined from CMNC, JMEC, MNNC.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76363D89F989FB01764E" pageId="17" pageNumber="18" type="description">
<paragraph id="D7761809742D76363D89F989FC9C7776" blockId="17.[151,1437,152,2018]" box="[198,782,1631,1658]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89F989FEEF7775" bold="true" box="[198,381,1631,1657]" pageId="17" pageNumber="18">Temporal data</emphasis>
. November (28), December (16).
</paragraph>
<paragraph id="D7761809742D76363D89F951FB01764E" blockId="17.[151,1437,152,2018]" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89F951FE1E77AD" bold="true" box="[198,396,1671,1697]" pageId="17" pageNumber="18">Natural History</emphasis>
. Specimens were collected at dusk (20:30 and 21:00 hrs) flying near or excavated from their burrows, which ranged in depth from
<quantity id="1031B5EC742D76363FC2F966FC8577C6" box="[653,791,1712,1738]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="18" unit="cm" value="50.0" valueMax="60.0" valueMin="40.0">40 to 60 cm</quantity>
and were located in forest gaps such as trails and roadsides. The forest that surrounds the open areas with burrows consists mainly of non-native pine plantations (
<taxonomicName id="10C9638A742D76363DEEF8D6FE357616" authority="D. Don" authorityName="D. Don" box="[161,423,1792,1818]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="radiata">
<emphasis id="E5BDC41B742D76363DEEF8D6FED07615" box="[161,322,1792,1817]" italics="true" pageId="17" pageNumber="18">Pinus radiata</emphasis>
D. Don
</taxonomicName>
,
<taxonomicName id="10C9638A742D76363CFAF8D6FDB07616" box="[437,546,1792,1818]" class="Pinopsida" family="Pinaceae" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Pinaceae</taxonomicName>
), mixed with fragments of relictual coastal Maulino forest restricted to the coastal zone of Maule in Region VII. This new species is sympatric with
<taxonomicName id="10C9638A742D76363EBDF8FEFB1F764D" box="[1010,1165,1832,1858]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tubericeps">
<emphasis id="E5BDC41B742D76363EBDF8FEFB96764D" box="[1010,1028,1832,1857]" italics="true" pageId="17" pageNumber="18">B</emphasis>
.
<emphasis id="E5BDC41B742D7636395CF8FEFB1F764D" box="[1043,1165,1832,1857]" italics="true" pageId="17" pageNumber="18">tubericeps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9FD34B82742D76353D89F899FD3571D6" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="D7761809742D76353D89F899FD3571D6" blockId="17.[151,1437,152,2018]" lastBlockId="18.[151,1436,152,218]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="E5BDC41B742D76363D89F899FED27665" bold="true" box="[198,320,1871,1897]" pageId="17" pageNumber="18">Remarks.</emphasis>
The known distribution of this species is restricted to the
<typeStatus id="0872A6AB742D76363EAFF886FB817666" box="[992,1043,1872,1898]" pageId="17" pageNumber="18">type</typeStatus>
locality, which is in the mountain range near the coast of Maule Region. This geographic distribution corresponds to the biogeographic region of Central
<collectingCountry id="AFDE5899742D76363C54F876FEC876B6" box="[283,346,1952,1978]" name="Chile" pageId="17" pageNumber="18">Chile</collectingCountry>
and Maule used by
<bibRefCitation id="B35865F8742D76363F17F876FCF276B6" box="[600,864,1952,1978]" pageId="17" pageNumber="23" refString="Morrone, J. J., Katinas, L., &amp; Crisci, J. V. (1997) A cladistic biogeographis analysis of Central Chile. Journal of Comparative Biology, 2, 25 - 42." type="journal article">
Morrone
<emphasis id="E5BDC41B742D76363F86F876FC9276B5" box="[713,768,1952,1977]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1997)
</bibRefCitation>
and
<bibRefCitation id="B35865F8742D76363ED0F876FBF076B6" author="Morrone" box="[927,1122,1952,1978]" pageId="17" pageNumber="23" refString="Morrone, J. J. (1999) Presentacion preliminar de un nuevo esquema biogeografico de America del Sur. Biogeografica, 75, 1 - 16." type="journal article" year="1999">Morrone (1999)</bibRefCitation>
. This new species is only known from specimens that were collected after the year 2000. This highlights the need for continued survey work in poorly collected coastal areas of central
<collectingCountry id="AFDE5899742E76353F98FF4EFC8B71BE" box="[727,793,152,178]" name="Chile" pageId="18" pageNumber="19">Chile</collectingCountry>
, especially since natural habitats these areas are under intensive pressure from human development.
</paragraph>
</subSubSection>
</treatment>
</document>
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