treatments-xml/data/4A/7A/AF/4A7AAFCCD8BB5522A03FE374AFA2EB71.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.64.6864" ID-PMC="PMC4956916" ID-Pensoft-Pub="1314-2003-64-1" ID-Pensoft-UUID="FF986A234A327324FFD9B756FF91FFB9" ID-PubMed="27489483" ID-Zenodo-Dep="576339" ModsDocID="1314-2003-64-1" checkinTime="1465845097482" checkinUser="pensoft" docAuthor="Hensold, Nancy" docDate="2016" docId="4A7AAFCCD8BB5522A03FE374AFA2EB71" docLanguage="en" docName="PhytoKeys 64: 1-57" docOrigin="PhytoKeys 64" docPubDate="2016-06-13" docSource="http://dx.doi.org/10.3897/phytokeys.64.6864" docTitle="Paepalanthus pilosus subsp. var. var. leoniae Hensold 2016, var. nov." docType="treatment" docVersion="3" id="FF986A234A327324FFD9B756FF91FFB9" lastPageNumber="43" masterDocId="FF986A234A327324FFD9B756FF91FFB9" masterDocTitle="The Andean Paepalanthus pilosus complex (Eriocaulaceae): a revision with three new taxa" masterLastPageNumber="57" masterPageNumber="1" pageNumber="40" updateTime="1668141705855" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The Andean Paepalanthus pilosus complex (Eriocaulaceae): a revision with three new taxa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hensold, Nancy</mods:namePart>
<mods:affiliation>Keller Science Action Center, The Field Museum of Natural History, 1400 S. Lakeshore Drive, Chicago, IL 60605 - 2496, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-06-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>64</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>57</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.64.6864</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.64.6864</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-64-1</mods:identifier>
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<treatment ID-GBIF-Taxon="182227051" LSID="urn:lsid:plazi:treatment:4A7AAFCCD8BB5522A03FE374AFA2EB71" httpUri="http://treatment.plazi.org/id/4A7AAFCCD8BB5522A03FE374AFA2EB71" lastPageId="42" lastPageNumber="43" pageId="39" pageNumber="40">
<subSubSection pageId="39" pageNumber="40" type="nomenclature">
<paragraph pageId="39" pageNumber="40">
5b. 
<taxonomicName LSID="4A7AAFCC-D8BB-5522-A03F-E374AFA2EB71" authority="Hensold" authorityName="Hensold" authorityYear="2016" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paepalanthus pilosus subsp. var. var. leoniae" order="Poales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="variety" species="pilosus" status="var. nov." subSpecies="var." variety="leoniae">Paepalanthus pilosus var. leoniae Hensold</taxonomicName>
<taxonomicNameLabel pageId="39" pageNumber="40">var. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habits of Paepalanthus pilosus varieties and &quot; Paepalanthus sp. A. &quot; A-D Paepalanthus pilosus var. pilosus. (A Soderstrom 1346 B Sagastegui 12242 C Cano 16840 D Larsen 237.) E Paepalanthus pilosus var. leoniae (Leon 1597) F-G &quot; Paepalanthus sp. A &quot; (Cuadros 3732) F Habit G Capitulum. In D arrows indicate flowering (fl) and fruiting (fr) capitula." figureDoi="10.3897/phytokeys.64.6864.figure11" httpUri="https://binary.pensoft.net/fig/92253" pageId="39" pageNumber="40">Figs 11E</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pistillate flowers and fruits of Paepalanthus pilosus varieties. A-K Paepalanthus pilosus var. pilosus (A-B Jorgensen 2366 C-F Cano 16840 G Cuatrecasas 25574 H MacDougal 4463 I Wurdack 1616 J Ollgaard 9557 K Boeke 2133) A-B Flower in anthesis with detail of gynoecial nectaries C Flower in anthesis D Gynoecium E Petal F Seed G Diaspore with detached calyx H Sepal (fr) I Diaspore releasing seed J-K Seeds after wetting L Paepalanthus pilosus var. leoniae. Mature diaspore, sepals and petals intact. (Leon 1597)." figureDoi="10.3897/phytokeys.64.6864.figure12" httpUri="https://binary.pensoft.net/fig/92254" pageId="39" pageNumber="40">, 12L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="diagnosis">
<paragraph pageId="40" pageNumber="41">
<pageBreakToken pageId="40" pageNumber="41" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="40" pageNumber="41">Differs from the typical variety by the capitula 2-3 mm in diameter, flowers 5-10 per capitulum, the pistillate flowers with sepals elliptic, uniformly thickened in fruit and dispersed with the fruit; petals of pistillate flowers oblong to oblong-spatulate with broadly rounded to truncate apex.</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="materials_examined">
<paragraph pageId="40" pageNumber="41">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="40" pageNumber="41">
<materialsCitation collectingDate="1988-03-16" collectorName="Prov. Mariscal Caceres, NW, B. Leon, K. Young" country="PERU" elevation="3600" latitude="-7.671314" location="Paredones" longLatPrecision="1" longitude="-77.483826" municipality="Rio Abiseo National Park" specimenCount="1" stateProvince="San Martin" typeStatus="Holotype">
<collectingCountry name="Peru">PERU</collectingCountry>
. 
<collectingRegion country="Peru" name="San Martin">
San 
<normalizedToken originalValue="Martín">Martin</normalizedToken>
</collectingRegion>
: 
<collectorName>Prov. Mariscal Caceres, NW</collectorName>
sector 
<collectingMunicipality>Rio Abiseo National Park</collectingMunicipality>
, grassland in 
<location LSID="urn:lsid:plazi:treatment:4A7AAFCCD8BB5522A03FE374AFA2EB71:5BC31ABE2E8B23330FE3AAE63F67CABF" country="PERU" latitude="-7.671314" longLatPrecision="1" longitude="-77.483826" municipality="Rio Abiseo National Park" name="Paredones" stateProvince="San Martin">Paredones</location>
, [
<geoCoordinate degrees="07" direction="south" minutes="40" orientation="latitude" precision="1" seconds="16.73" value="-7.671314">07°40'16.73&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="29" orientation="longitude" precision="1" seconds="1.78" value="-77.483826">77°29'1.78&quot;W</geoCoordinate>
], 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">3600 m</elevation>
</quantity>
, 
<collectingDate value="1988-03-16">16 March 1988</collectingDate>
, 
<emphasis italics="true" pageId="40" pageNumber="41">
<collectorName>
B. 
<normalizedToken originalValue="León">Leon</normalizedToken>
</collectorName>
&amp; 
<collectorName>K. Young</collectorName>
1597
</emphasis>
(
<typeStatus>Holotype</typeStatus>
: MO!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph pageId="40" pageNumber="41">Description.</paragraph>
<paragraph lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis italics="true" pageId="40" pageNumber="41">Leaves</emphasis>
5.5-12 mm long 
<normalizedToken originalValue="×">x</normalizedToken>
0.6-1.4 mm wide at midpoint, sharply cuspidate, finely appressed-ciliate near apex when young, early glabrate. Peduncles 3-6.5 mm long, fruiting peduncles the same length in specimens observed, the sheaths 5-7 mm. 
<emphasis italics="true" pageId="40" pageNumber="41">Capitula</emphasis>
2-3 mm in diameter, the involucres obconic or cupulate in flower and fruit, not opening broadly, the bracts similar to typical variety. Flowers 5-10 per capitulum, the staminate equal in number to pistillate or in fewer-flowered capitula the proportion of staminate flowers may be reduced. 
<emphasis italics="true" pageId="40" pageNumber="41">Pistillate flowers</emphasis>
: Pedicels 0.25-0.35 mm. 
<emphasis italics="true" pageId="40" pageNumber="41">Sepals</emphasis>
elliptic, obtuse to rounded, 1.35-2.4 mm long 
<normalizedToken originalValue="×">x</normalizedToken>
0.65-0.8 mm wide at middle, 0.25-0.35 mm wide at base, pale brownish, tinged dusky-brown on shoulders, becoming uniformly thickened and cymbiform at maturity, enclosing the fruit and dispersed with it. 
<emphasis italics="true" pageId="40" pageNumber="41">Petals</emphasis>
oblong to oblong-spatulate, with apex broadly rounded to truncate, retuse or apiculate, 1.5-2.0 mm long 
<normalizedToken originalValue="×">x</normalizedToken>
0.65-0.85 mm wide, 2.2-2.5 times longer than wide, pilose as in the typical variety or with trichomes restricted to subapical tufts, uniformly thickening in fruit, as the sepals. 
<emphasis italics="true" pageId="40" pageNumber="41">Gynoecium</emphasis>
with style base 0.5-0.65 mm long, nectaries 0.65-1.0 mm long, papillae at upper margin colorless, stiff, globose (
<emphasis italics="true" pageId="40" pageNumber="41">Leon 1597</emphasis>
) to linear (
<emphasis italics="true" pageId="40" pageNumber="41">Young 4368</emphasis>
); style branches 0.75-1.7 mm. Seeds 
<pageBreakToken pageId="41" pageNumber="42" start="start">not</pageBreakToken>
seen but mature ovary locules 0.65 mm (
<emphasis italics="true" pageId="41" pageNumber="42">Young 4368</emphasis>
), 0.75 mm (
<emphasis italics="true" pageId="41" pageNumber="42">Leon 1597</emphasis>
). 
<emphasis italics="true" pageId="41" pageNumber="42">Staminate flowers</emphasis>
: Pedicels 0.25-0.35 mm. Sepals 1.35-2.1 mm long, fused 35-55 % of their length. Corolla 1.55-2.05 mm long, with the anthophore 0.85-1.15 mm long, comprising 45-75% the length of the whole corolla, ca. 0.1-0.25 mm in diameter at base, the tube and lobes 0.4-1.05 mm long. Filaments brownish toward apex; nectaries reaching corolla sinuses.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="etymology">
<paragraph pageId="41" pageNumber="42">Etymology.</paragraph>
<paragraph pageId="41" pageNumber="42">
Named in honor of Dr. Blanca 
<normalizedToken originalValue="León">Leon</normalizedToken>
, of the Universidad Nacional Mayor de San Marcos and the University of Texas, a generous and accomplished botanist whose many contributions include ecological and taxonomic study of Rio Abiseo National Park.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="phenology">
<paragraph pageId="41" pageNumber="42">Phenology.</paragraph>
<paragraph pageId="41" pageNumber="42">
Collected in March and July. A mild dry season occurs June to August (
<bibRefCitation author="Young, KR" journalOrPublisher="Revista Forestal del Peru" pageId="48" pageNumber="49" pagination="3 - 20" refId="B77" refString="Young, KR, Leon, B, 1988. Vegetacion de la zona alta del Parque Nacional Rio Abiseo, San Martin. Revista Forestal del Peru 15: 3 - 20" title="Vegetacion de la zona alta del Parque Nacional Rio Abiseo, San Martin." volume="15" year="1988">Young and Leon 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="distribution">
<paragraph pageId="41" pageNumber="42">Distribution.</paragraph>
<paragraph pageId="41" pageNumber="42">
Endemic to Peru, San 
<normalizedToken originalValue="Martín">Martin</normalizedToken>
, Prov. Mariscal Caceres, Rio Abiseo National Park. (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Paepalanthus caryonauta and suspected hybrids, Paepalanthus pilosus var. pilosus, Paepalanthus pilosus var. leoniae, and Paepalanthus &quot; Species A. &quot;" figureDoi="10.3897/phytokeys.64.6864.figure6" httpUri="https://binary.pensoft.net/fig/92248" pageId="41" pageNumber="42">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="habitat">
<paragraph pageId="41" pageNumber="42">Habitat.</paragraph>
<paragraph pageId="41" pageNumber="42">
From high-elevation grasslands, at 3450-3800 m. 
<bibRefCitation author="Young, KR" journalOrPublisher="Revista Forestal del Peru" pageId="48" pageNumber="49" pagination="3 - 20" refId="B77" refString="Young, KR, Leon, B, 1988. Vegetacion de la zona alta del Parque Nacional Rio Abiseo, San Martin. Revista Forestal del Peru 15: 3 - 20" title="Vegetacion de la zona alta del Parque Nacional Rio Abiseo, San Martin." volume="15" year="1988">
Young and 
<normalizedToken originalValue="León">Leon</normalizedToken>
(1988)
</bibRefCitation>
compare the habitat to the wet 
<normalizedToken originalValue="páramos">paramos</normalizedToken>
of southern Ecuador, and note that the U-shaped valleys at high elevations where this variety was collected were glaciated as recently as 12,000 years ago.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="conservation status">
<paragraph pageId="41" pageNumber="42">Conservation status.</paragraph>
<paragraph pageId="41" pageNumber="42">
Endangered, Criteria B1ab(iii) (
<bibRefCitation author="IUCN Standards and Petitions Subcommittee" journalOrPublisher="Prepared by the Standards and Petitions Subcommittee" pageId="46" pageNumber="47" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B24" refString="IUCN Standards and Petitions Subcommittee, 2014. Guidelines for using the IUCN Red List Categories and Criteria. Version 11. Prepared by the Standards and Petitions Subcommittee, http://www.iucnredlist.org/documents/RedListGuidelines.pdf [on 20 July 2015]" title="Guidelines for using the IUCN Red List Categories and Criteria. Version 11" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2014">IUCN 2014</bibRefCitation>
). This variety is known from only two sites 10 km apart in the protected area of Rio Abiseo National Park In recent years, cattle-grazing, which may have been a threat, has ceased (B. 
<normalizedToken originalValue="León">Leon</normalizedToken>
, pers. comm.) However, the very small distribution, and the fact that this variety represents the southernmost occurrence of a wetland 
<normalizedToken originalValue="páramo">paramo</normalizedToken>
species suggests vulnerability to climate fluctuations.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="discussion">
<paragraph pageId="41" pageNumber="42">Discussion.</paragraph>
<paragraph pageId="41" pageNumber="42">
This variety is found at the southernmost and highest elevation station for the species in Peru. It occurs above 3400 m at the very south end of the Amotape-Huancabamba floristic zone as characterized by 
<bibRefCitation author="Weigend, M" journalOrPublisher="Revista Peruana de Biologia" pageId="48" pageNumber="49" pagination="127 - 134" publicationUrl="10.15381/rpb.v11i2.2447" refId="B75" refString="Weigend, M, 2004. Additional observations on the biogeography of the Amotape-Huancabamba zone in Northern Peru: defining the south-eastern limits. Revista Peruana de Biologia 11 (2): 127 - 134, DOI: 10.15381/rpb.v11i2.2447" title="Additional observations on the biogeography of the Amotape-Huancabamba zone in Northern Peru: defining the south-eastern limits." url="10.15381/rpb.v11i2.2447" volume="11" year="2004">Weigend (2004)</bibRefCitation>
, in the drainage of the Rio Huallaga. This compares to the typical variety collected at up to 3400 m, in the drainage of the Rio 
<normalizedToken originalValue="Marañón">Maranon</normalizedToken>
.
</paragraph>
<paragraph pageId="41" pageNumber="42">
The uniformly thickened fruiting perianth of this variety is similar to that of 
<taxonomicName authorityName="Hensold" authorityYear="2016" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paepalanthus caryonauta" order="Poales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="caryonauta">
<emphasis italics="true" pageId="41" pageNumber="42">Paepalanthus caryonauta</emphasis>
</taxonomicName>
, complicating the otherwise tidy distinction between the two species. However the size and shape of the leaves, the dwarf peduncles, the pale greenish involucres, and the large rigid nectary papillae, all suggest 
<taxonomicName class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paepalanthus pilosus" order="Poales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="pilosus">
<emphasis italics="true" pageId="41" pageNumber="42">Paepalanthus pilosus</emphasis>
</taxonomicName>
, and the type specimen in particular looks nearly identical to populations of typical 
<taxonomicName class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paepalanthus pilosus" order="Poales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="pilosus">
<emphasis italics="true" pageId="41" pageNumber="42">Paepalanthus pilosus</emphasis>
</taxonomicName>
from adjacent southern Amazonas. In fact, these populations are partly intermediate in floral morphology between the two varieties, as discussed under the typical variety. Given the variation in 
<taxonomicName class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paepalanthus pilosus" order="Poales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="pilosus">
<emphasis italics="true" pageId="41" pageNumber="42">Paepalanthus pilosus</emphasis>
</taxonomicName>
in Peru and the contiguity of distribution, it seems best for now to treat this taxon as a variety of that species.
</paragraph>
<paragraph pageId="41" pageNumber="42">
The specimen 
<emphasis italics="true" pageId="41" pageNumber="42">
Young &amp; 
<normalizedToken originalValue="León">Leon</normalizedToken>
4368
</emphasis>
, collected ca. 10 km from the other two, is marked by the very small size (5.5-7 mm) of the dark green leaves. The flowers and seeds are also smaller. The habitats of the two localities differed, with the type locality a typical patchy wet 
<normalizedToken originalValue="páramo">paramo</normalizedToken>
among outcrops, while the small-leaved plant was in a broad boggy area bordered by small trees, where cattle once pastured (B. 
<normalizedToken originalValue="León">Leon</normalizedToken>
, pers. comm.)
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="materials_examined">
<paragraph pageId="42" pageNumber="43">
<pageBreakToken pageId="42" pageNumber="43" start="start">Additional</pageBreakToken>
specimens examined.
</paragraph>
<paragraph pageId="42" pageNumber="43">
<materialsCitation collectingDate="1987-07-10" collectingDateMax="2000-07-27" collectingDateMin="1987-07-10" collectorName="K. Young, B. Leon, de Empedrada" country="Peru" county="Prov. Mariscal Caceres. Puerta del Monte" elevation="3450" latitude="-7.6713886" location="Pastizales de Empedrada" longLatPrecision="21" longitude="-77.48389" municipality="Rio Abiseo National Park" specimenCount="1" stateProvince="San Martin">
<normalizedToken originalValue="Perú">
<collectingCountry name="Peru">Peru</collectingCountry>
</normalizedToken>
. 
<collectingRegion country="Peru" name="San Martin">
San 
<normalizedToken originalValue="Martín">Martin</normalizedToken>
</collectingRegion>
: 
<collectingCounty>Prov. Mariscal Caceres. Puerta del Monte</collectingCounty>
, northwest corner of 
<collectingMunicipality>Rio Abiseo National Park</collectingMunicipality>
; high elevation grassland on bottom of u-shaped valley; [
<geoCoordinate degrees="7" direction="south" minutes="39" orientation="latitude" precision="1" seconds="30.20" value="-7.658389">7°39'30.20&quot;S</geoCoordinate>
; 
<geoCoordinate degrees="77" direction="west" minutes="28" orientation="longitude" precision="1" seconds="13.67" value="-77.47047">77°28'13.67&quot;W</geoCoordinate>
], 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.45" unit="m" value="3450.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.45" unit="m" value="3450.0">3450 m</elevation>
</quantity>
, 
<collectingDate value="1987-07-10">10 Jul 1987</collectingDate>
, 
<emphasis italics="true" pageId="42" pageNumber="43">
<collectorName>K. Young</collectorName>
&amp; 
<collectorName>
B. 
<normalizedToken originalValue="León">Leon</normalizedToken>
</collectorName>
4368
</emphasis>
(F); 
<location LSID="urn:lsid:plazi:treatment:4A7AAFCCD8BB5522A03FE374AFA2EB71:9D5E528F2F0D307FD3B7E8A2CA957BFE" country="Peru" county="Prov. Mariscal Caceres. Puerta del Monte" latitude="-7.6713886" longLatPrecision="21" longitude="-77.48389" municipality="Rio Abiseo National Park" name="Pastizales de Empedrada" stateProvince="San Martin">
Pastizales 
<collectorName>de Empedrada</collectorName>
</location>
, entre manojos de 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="42" pageNumber="43" phylum="Magnoliophyta" rank="genus">Calamagrostis</taxonomicName>
, [
<geoCoordinate degrees="07" direction="south" minutes="40" orientation="latitude" precision="15" seconds="17" value="-7.6713886">07°40'17&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="29" orientation="longitude" precision="15" seconds="2" value="-77.48389">77°29'2&quot; W</geoCoordinate>
], 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.7674999999999996" metricValueMax="3.785" metricValueMin="3.75" unit="m" value="3767.5" valueMax="3785.0" valueMin="3750.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.7674999999999996" metricValueMax="3.785" metricValueMin="3.75" unit="m" value="3767.5" valueMax="3785.0" valueMin="3750.0">3750-3785 m</elevation>
</quantity>
, 
<collectingDate value="2000-07-27">27 Jul 2000</collectingDate>
, 
<emphasis italics="true" pageId="42" pageNumber="43">
<collectorName>
B. 
<normalizedToken originalValue="León">Leon</normalizedToken>
</collectorName>
&amp; 
<collectorName>K. Young</collectorName>
4579
</emphasis>
(USM [photo!])
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>