treatments-xml/data/30/4F/87/304F87936817FFDAFF3BFDF9FD81AB4E.xml

<document id="8292B268C39666CA3CA53F70FB4C068F" ID-DOI="10.11646/zootaxa.4006.1.9" ID-GBIF-Dataset="0c519915-9ecb-427d-998a-5429a90132e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="237087" ID-ZooBank="E987B43A-E54A-4F64-9829-0B8EBE457E03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461203959611" checkinUser="plazi" docAuthor="Dózsa-Farkas, Klára, Felföldi, Tamás &amp; Hong, Yong" docDate="2015" docId="304F87936817FFDAFF3BFDF9FD81AB4E" docLanguage="en" docName="zt04006p197.pdf" docOrigin="Zootaxa 4006 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Henlea magnaampullacea Dózsa-Farkas, Felföldi &amp; Hong, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="177" masterDocId="CC76FFEB6814FFDDFFACFFECFFB3AA53" masterDocTitle="New enchytraeid species (Enchytraeidae, Oligochaeta) from Korea" masterLastPageNumber="197" masterPageNumber="171" pageNumber="174" updateTime="1698601274417" updateUser="plazi">
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<mods:title id="F8F38BB6BD977F684C3C102AED3C1F19">New enchytraeid species (Enchytraeidae, Oligochaeta) from Korea</mods:title>
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<mods:namePart id="93C40FFA2CCAF8594118E548E56F8575">Dózsa-Farkas, Klára</mods:namePart>
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<mods:namePart id="1246E726237C99228858DBAE02CE0F86">Felföldi, Tamás</mods:namePart>
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<mods:namePart id="0D0115B6A12C12AD2E500070CE6D1A1E">Hong, Yong</mods:namePart>
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<treatment id="304F87936817FFDAFF3BFDF9FD81AB4E" ID-DOI="http://doi.org/10.5281/zenodo.6100530" ID-GBIF-Taxon="119630007" ID-Zenodo-Dep="6100530" LSID="urn:lsid:plazi:treatment:304F87936817FFDAFF3BFDF9FD81AB4E" httpUri="http://treatment.plazi.org/id/304F87936817FFDAFF3BFDF9FD81AB4E" lastPageId="7" lastPageNumber="177" pageId="3" pageNumber="174">
<subSubSection id="F0FC650E6817FFDEFF3BFDF9FE03A803" pageId="3" pageNumber="174" type="nomenclature">
<paragraph id="B85936856817FFDEFF3BFDF9FDACA87C" blockId="3.[151,543,533,592]" box="[151,543,533,559]" pageId="3" pageNumber="174">
<heading id="E31181E96817FFDEFF3BFDF9FDACA87C" bold="true" box="[151,543,533,559]" fontSize="11" level="1" pageId="3" pageNumber="174" reason="1">
<emphasis id="8A92EA976817FFDEFF3BFDF9FDACA87C" bold="true" box="[151,543,533,559]" pageId="3" pageNumber="174">
<taxonomicName id="7FE64D066817FFDEFF3BFDF9FE66A87C" ID-CoL="85SQB" box="[151,469,533,559]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="3" pageNumber="174" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976817FFDEFF3BFDF9FE66A87C" bold="true" box="[151,469,533,559]" italics="true" pageId="3" pageNumber="174">Henlea magnaampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6817FFDEFE70FDF9FDACA87C" box="[476,543,533,559]" pageId="3" pageNumber="174" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="B85936856817FFDEFF3BFDDBFE03A803" blockId="3.[151,543,533,592]" box="[151,432,567,592]" pageId="3" pageNumber="174">
(
<figureCitation id="20DD2A006817FFDEFF32FDDBFEB7A803" box="[158,260,567,592]" captionStart="FIGURE 1" captionStartId="4.[151,250,1886,1908]" captionTargetBox="[186,1403,208,1851]" captionTargetId="figure@4.[175,1411,195,1865]" captionTargetPageId="4" captionText="FIGURE 1. Henlea magnaampullacea sp. n. (A – D) and Fridericia sphaerica sp. n. (E – G). H. magnaampullacea: A. Spermathecae; B. Sperm funnel; C. Nephridium; D. Intestinal diverticulum (ventral view). F. sphaerica: E. Oesophageal appendage. F. Sperm funnel; G. Spermatheca." httpUri="https://zenodo.org/record/237088/files/figure.png" pageId="3" pageNumber="174">Figures 1</figureCitation>
A–D, 2A–F, 3)
</paragraph>
</subSubSection>
<subSubSection id="F0FC650E6817FFDEFF3BFD93FBDCA9EB" pageId="3" pageNumber="174" type="materials_examined">
<paragraph id="B85936856817FFDEFF3BFD93FA8FA8B3" blockId="3.[151,1437,639,2033]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF3BFD93FD0FA8CB" bold="true" box="[151,700,639,664]" pageId="3" pageNumber="174">
<typeStatus id="675D88276817FFDEFF3BFD93FF61A8CB" box="[151,210,639,664]" pageId="3" pageNumber="174">Type</typeStatus>
material. 
<typeStatus id="675D88276817FFDEFEE3FD93FE0DA8CB" box="[335,446,639,664]" pageId="3" pageNumber="174" type="holotype">Holotype</typeStatus>
. NIBRIV0000320515
</emphasis>
, slide 
<emphasis id="8A92EA976817FFDEFCA5FD93FCCBA8CB" bold="true" box="[777,888,639,664]" pageId="3" pageNumber="174">No. 1126,</emphasis>
adult whole mount stained with borax-carmine. 
<typeStatus id="675D88276817FFDEFF3BFD49FF7CA8EE" box="[151,207,677,701]" pageId="3" pageNumber="174">Type</typeStatus>
locality: Site 1: College of Agriculture &amp; Life Science, Chonbuk National University, Jeonju-si, Jeollabuk-do, 
<collectingCountry id="C0F176156817FFDEFF3BFD24FF6DA88C" box="[151,222,712,735]" name="South Korea" pageId="3" pageNumber="174">Korea</collectingCountry>
, 
<geoCoordinate id="DDD250426817FFDEFF45FD2BFECCA8B3" box="[233,383,711,736]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="1" value="35.84972">35º50'59.0&quot;N</geoCoordinate>
<geoCoordinate id="DDD250426817FFDEFE2AFD2BFD99A8B3" box="[390,554,711,736]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="1" value="127.13233">127º07'56.4&quot;E</geoCoordinate>
, 
<quantity id="7F1E9B606817FFDEFD9AFD2BFDDCA88C" box="[566,623,711,736]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="174" unit="m" value="55.0">55 m</quantity>
asl, soil and litter layers of woodland, leg.Y. Hong, 
<date id="CC5810456817FFDEFB1AFD2BFA8BA8B3" box="[1206,1336,711,736]" pageId="3" pageNumber="174" value="2014-05-19">19.05.2014</date>
.
</paragraph>
<paragraph id="B85936856817FFDEFF6BFD00FA3BA9C6" blockId="3.[151,1437,639,2033]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF6BFD00FCE0A956" bold="true" box="[199,851,748,773]" pageId="3" pageNumber="174">
<typeStatus id="675D88276817FFDEFF6BFD00FEF2A956" box="[199,321,748,773]" pageId="3" pageNumber="174" type="paratype">Paratypes</typeStatus>
. NIBRIV0000320516, NIBRIV0000320517,
</emphasis>
<specimenCount id="AEE0FD0C6817FFDEFCCEFD00FC04A956" box="[866,951,748,773]" pageId="3" pageNumber="174" type="adult">2 adult</specimenCount>
specimens from 
<typeStatus id="675D88276817FFDEFB22FD01FB0DA956" box="[1166,1214,749,773]" pageId="3" pageNumber="174">type</typeStatus>
locality, in 70 % ethanol. 
<emphasis id="8A92EA976817FFDEFF55FCE3FEDAA97B" bold="true" box="[249,361,783,808]" pageId="3" pageNumber="174">P.103.1–4</emphasis>
, slide 
<emphasis id="8A92EA976817FFDEFE1AFCE3FD42A97B" bold="true" box="[438,753,783,808]" pageId="3" pageNumber="174">No. 1005, 1121, 1122, 1127</emphasis>
. Four adult whole mounts stained with borax-carmine from the 
<typeStatus id="675D88276817FFDEFF6BFCD9FF44A91E" box="[199,247,821,845]" pageId="3" pageNumber="174">type</typeStatus>
locality, 0 3.04.2014 and 
<date id="CC5810456817FFDEFD9CFCD8FD01A91E" box="[560,690,820,845]" pageId="3" pageNumber="174" value="2014-05-19">19.05.2014</date>
. 
<emphasis id="8A92EA976817FFDEFD68FCD8FCC9A91E" bold="true" box="[708,890,820,845]" pageId="3" pageNumber="174">P.103.5–103.10.</emphasis>
slide 
<emphasis id="8A92EA976817FFDEFC60FCD8FAD3A91E" bold="true" box="[972,1376,820,845]" pageId="3" pageNumber="174">
No. 1098, 1119, 1120, 
<date id="CC5810456817FFDEFB4AFCD8FAD3A91E" box="[1254,1376,820,845]" pageId="3" pageNumber="174" value="1128" valueMax="1130">1128–1130</date>
</emphasis>
. Six specimens from site 7, Dongjin-myeon, Buan-gun, Jeollabuk-do, 
<collectingCountry id="C0F176156817FFDEFC25FCB4FC63A93C" box="[905,976,856,879]" name="South Korea" pageId="3" pageNumber="174">Korea</collectingCountry>
, 
<geoCoordinate id="DDD250426817FFDEFC72FCBBFBC7A923" box="[990,1140,855,880]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="1" value="35.767944">35º46'04.6&quot;N</geoCoordinate>
<geoCoordinate id="DDD250426817FFDEFBD2FCBBFA91A923" box="[1150,1314,855,880]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="1" value="126.72078">126º43'14.8&quot;E</geoCoordinate>
, 
<quantity id="7F1E9B606817FFDEFA9DFCB4FADFA93C" box="[1329,1388,856,880]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="174" unit="m" value="23.0">23 m</quantity>
asl, soil of agronomical fields, leg. Y. Hong, 
<date id="CC5810456817FFDEFDF3FC90FD52A9C6" box="[607,737,892,917]" pageId="3" pageNumber="174" value="2014-05-19">19.05.2014</date>
. 
<emphasis id="8A92EA976817FFDEFD47FC90FCF8A9C6" bold="true" box="[747,843,892,917]" pageId="3" pageNumber="174">P.103.11</emphasis>
. Six specimens in 70 % ethanol from 
<typeStatus id="675D88276817FFDEFB5AFC91FA95A9C6" box="[1270,1318,893,917]" pageId="3" pageNumber="174">type</typeStatus>
locality.
</paragraph>
<paragraph id="B85936856817FFDEFF6BFC73FBDCA9EB" blockId="3.[151,1437,639,2033]" box="[199,1135,927,952]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF6BFC73FDBFA9EB" bold="true" box="[199,524,927,952]" pageId="3" pageNumber="174">Further material examined</emphasis>
. 12 and 
<specimenCount id="AEE0FD0C6817FFDEFDC0FC4CFCB7A9EB" box="[620,772,927,952]" pageId="3" pageNumber="174" type="generic">10 specimens</specimenCount>
from sites 7 and 1, respectively.
</paragraph>
</subSubSection>
<subSubSection id="F0FC650E6817FFDEFF6BFC28FBFDA98E" box="[199,1102,964,989]" pageId="3" pageNumber="174" type="etymology">
<paragraph id="B85936856817FFDEFF6BFC28FBFDA98E" blockId="3.[151,1437,639,2033]" box="[199,1102,964,989]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF6BFC28FEFDA98E" bold="true" box="[199,334,964,989]" pageId="3" pageNumber="174">Etymology.</emphasis>
Named after the large spermathecal ampulla (magna = large, Latin).
</paragraph>
</subSubSection>
<subSubSection id="F0FC650E6817FFDEFF6BFC0BFDA3AE8B" pageId="3" pageNumber="174" type="diagnosis">
<paragraph id="B85936856817FFDEFF6BFC0BFDA3AE8B" blockId="3.[151,1437,639,2033]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF6BFC0BFEF2AE53" bold="true" box="[199,321,999,1024]" pageId="3" pageNumber="174">Diagnosis.</emphasis>
The new species can be recognized by the following combination of characters: (1) medium-sized stout worms (
<quantity id="7F1E9B606817FFDEFE9CFBE0FE19AE77" box="[304,426,1036,1061]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="3" pageNumber="174" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11–16 mm</quantity>
long and 600–800 Μm wide at clitellum 
<emphasis id="8A92EA976817FFDEFCDFFBE1FC0CAE77" box="[883,959,1037,1060]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
, segments 47–58; (2) maximum 6 chaetae per bundle; (3) clitellum girdle-shaped: gland cells small in reticulate pattern dorsally and ventrally; (4) six preclitellar pairs of nephridia; (5) canals of the intestinal diverticula arranged longitudinally in VIII, dorsal vessel origin in IX; (6) coelomocytes large, rounded brownish, thickened at cell periphery; (7) seminal vesicle large; (8) sperm funnel large, cylindrical, collar wider as funnel body; (9) spermathecae with very large pear-shaped ampullae and without diverticula.
</paragraph>
</subSubSection>
<subSubSection id="F0FC650E6817FFD8FF6BFB08FD3DAD93" lastPageId="5" lastPageNumber="176" pageId="3" pageNumber="174" type="description">
<paragraph id="B85936856817FFDEFF6BFB08FB5AAC36" blockId="3.[151,1437,639,2033]" pageId="3" pageNumber="174">
<emphasis id="8A92EA976817FFDEFF6BFB08FEEAAEAE" bold="true" box="[199,345,1252,1277]" pageId="3" pageNumber="174">Description.</emphasis>
<typeStatus id="675D88276817FFDEFECDFB08FE7AAEAE" box="[353,457,1252,1277]" pageId="3" pageNumber="174" type="holotype">Holotype</typeStatus>
<quantity id="7F1E9B606817FFDEFE7EFB08FD84AEAF" box="[466,567,1252,1277]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="3" pageNumber="174" unit="mm" value="16.2">16.2 mm</quantity>
long, 590 µm wide at VIII and 620 µm at clitellum, 
<emphasis id="8A92EA976817FFDEFB34FB09FB56AEAF" box="[1176,1253,1253,1276]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
(
<quantity id="7F1E9B606817FFDEFB59FB08FAE8AEAF" box="[1269,1371,1252,1276]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="3" pageNumber="174" unit="mm" value="11.1">11.1 mm</quantity>
long, 490 µm wide at VIII and 660 µm at clitellum, fixed), 52 segments. Body length of 
<typeStatus id="675D88276817FFDEFBF1FAE5FB7AAF72" box="[1117,1225,1289,1313]" pageId="3" pageNumber="174" type="paratype">paratypes</typeStatus>
<quantity id="7F1E9B606817FFDEFB7EFAEBFAFCAF73" box="[1234,1359,1287,1312]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="3" pageNumber="174" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11–16 mm</quantity>
, width 520–700 µm at VIII and 530–800 µm at clitellum, 
<emphasis id="8A92EA976817FFDEFD60FAC1FCA5AF17" box="[716,790,1325,1348]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
. Length of fixed specimens 
<quantity id="7F1E9B606817FFDEFBFEFAC0FB5AAF17" box="[1106,1257,1324,1349]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.35" metricValueMin="0.8699999999999999" pageId="3" pageNumber="174" unit="mm" value="11.1" valueMax="13.5" valueMin="8.7">8.7–13.5 mm</quantity>
, width 400–670 µm at VIII and 400–730 µm at clitellum. Segments 47–58. Chaetal formula: 2,3,4,5 – 4,5,(3,2): 2,3,4,5,6 – 4,5,6,3. Chaetae straight, mostly unequal in size within a bundle; the largest 90–100 Μm long and 5 Μm wide, the smallest 63–70 Μm long and 4 Μm wide. Chaetae in XII absent. Head pore at 0/I, large transverse slit (
<figureCitation id="20DD2A006817FFDEFB16FA7BFB4EAFE3" box="[1210,1277,1431,1456]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
B). Epidermal gland cells arranged in 3–4 transverse rows per segment. Many glands also on the prostomium dorsally (
<figureCitation id="20DD2A006817FFDEFA9AFA50FACAAF87" box="[1334,1401,1468,1493]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
B). Clitellum girdle-shaped in 
<date id="CC5810456817FFDEFE67FA0CFDBFAFAB" box="[459,524,1504,1528]" bridgedPair="-" pageId="3" pageNumber="174">XII–1</date>
/2XIII, gland cells small in reticulate pattern (
<figureCitation id="20DD2A006817FFDEFBB2FA33FBD7AFAB" box="[1054,1124,1503,1528]" captionStart="FIGURE 2" captionStartId="5.[151,250,1385,1407]" captionTargetBox="[180,1398,193,1353]" captionTargetId="figure@5.[180,1408,193,1364]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Henlea magnaampullacea sp. n. (A – F) and Fridericia sphaerica sp. n. (G – I). H. magnaapullacea: A. Origin of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). B. Intestinal diverticula in VIII. C. Clitellar glands, dorsal view. D. Clitellar glands between the male copulatory organs, ventral view. E. Spermathecae of the holotype. F. Spermathecae of a fixed and stained specimens. Fridericia sphaerica: G. Clitellar glands dorsal view. H. Sperm funnels. I. Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs in vivo, except F. Scale bars 50 µm." httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="3" pageNumber="174">Fig. 2</figureCitation>
C), also between the bursal slits (
<figureCitation id="20DD2A006817FFDEFF79F9E8FEA8AC4F" box="[213,283,1540,1565]" captionStart="FIGURE 2" captionStartId="5.[151,250,1385,1407]" captionTargetBox="[180,1398,193,1353]" captionTargetId="figure@5.[180,1408,193,1364]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Henlea magnaampullacea sp. n. (A – F) and Fridericia sphaerica sp. n. (G – I). H. magnaapullacea: A. Origin of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). B. Intestinal diverticula in VIII. C. Clitellar glands, dorsal view. D. Clitellar glands between the male copulatory organs, ventral view. E. Spermathecae of the holotype. F. Spermathecae of a fixed and stained specimens. Fridericia sphaerica: G. Clitellar glands dorsal view. H. Sperm funnels. I. Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs in vivo, except F. Scale bars 50 µm." httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="3" pageNumber="174">Fig. 2</figureCitation>
D). Thickness of body wall about 40–60 µm, cuticle about 1 µm, 
<emphasis id="8A92EA976817FFDEFBBFF9E9FBD3AC4F" box="[1043,1120,1541,1564]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
. Brain concave posteriorly, about 150 Μm long, 
<emphasis id="8A92EA976817FFDEFED7F9C4FE74AC6C" box="[379,455,1576,1599]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
and slightly longer than wide (
<figureCitation id="20DD2A006817FFDEFC8BF9CBFCD9AC13" box="[807,874,1575,1600]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
A). Oesophagus in VI with a pattern of transverse streaks and one pair dorso-laterally and one pair ventro-laterally lobes of oesophageal appendages.
</paragraph>
<paragraph id="B85936856817FFDEFF6BF983FDA2ADA2" blockId="3.[151,1437,639,2033]" pageId="3" pageNumber="174">
Pharyngeal glands all separate dorsally, the second and third pairs with ventral lobes (
<figureCitation id="20DD2A006817FFDEFB24F983FB7FACDB" box="[1160,1228,1647,1672]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
E.). Chloragocytes from IV about 15–40 Μm long, 
<emphasis id="8A92EA976817FFDEFDB5F979FDD8ACFF" box="[537,619,1685,1708]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
(fixed 13–15 Μm). Dorsal vessel from IX, large heart-like pulsating expansion from IX to VII (
<figureCitation id="20DD2A006817FFDEFE68F95BFDB4AC83" box="[452,519,1719,1744]" captionStart="FIGURE 2" captionStartId="5.[151,250,1385,1407]" captionTargetBox="[180,1398,193,1353]" captionTargetId="figure@5.[180,1408,193,1364]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Henlea magnaampullacea sp. n. (A – F) and Fridericia sphaerica sp. n. (G – I). H. magnaapullacea: A. Origin of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). B. Intestinal diverticula in VIII. C. Clitellar glands, dorsal view. D. Clitellar glands between the male copulatory organs, ventral view. E. Spermathecae of the holotype. F. Spermathecae of a fixed and stained specimens. Fridericia sphaerica: G. Clitellar glands dorsal view. H. Sperm funnels. I. Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs in vivo, except F. Scale bars 50 µm." httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="3" pageNumber="174">Fig. 2</figureCitation>
A), blood colourless. Intestinal diverticula forming a ring around intestine in VIII, consisting of 4 spherical diverticula, which unite proximally (
<figureCitation id="20DD2A006817FFDEFCFAF930FC15ACA7" box="[854,934,1756,1781]" captionStart="FIGURE 2" captionStartId="5.[151,250,1385,1407]" captionTargetBox="[180,1398,193,1353]" captionTargetId="figure@5.[180,1408,193,1364]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Henlea magnaampullacea sp. n. (A – F) and Fridericia sphaerica sp. n. (G – I). H. magnaapullacea: A. Origin of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). B. Intestinal diverticula in VIII. C. Clitellar glands, dorsal view. D. Clitellar glands between the male copulatory organs, ventral view. E. Spermathecae of the holotype. F. Spermathecae of a fixed and stained specimens. Fridericia sphaerica: G. Clitellar glands dorsal view. H. Sperm funnels. I. Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs in vivo, except F. Scale bars 50 µm." httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="3" pageNumber="174">Figs. 2</figureCitation>
A,B, 3E). Canals of the diverticula arranged longitudinally and not including large hollows; especially well visible in subadult or juvenile specimens (
<figureCitation id="20DD2A006817FFDEFA95F913FACFAD4B" box="[1337,1404,1791,1816]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
F). Intestinal epithelium behind clitellum with tall, hyaline 
<typeStatus id="675D88276817FFDEFCB9F8C9FCF6AD6E" box="[789,837,1829,1853]" pageId="3" pageNumber="174">type</typeStatus>
of cells ventrally and laterally from XXIV–XXV to XXXI–XXXII (
<typeStatus id="675D88276817FFDEFEE4F8A4FEC9AD33" box="[328,378,1864,1888]" pageId="3" pageNumber="174">type</typeStatus>
2 after 
<bibRefCitation id="DC774B746817FFDEFE7CF8A5FD35AD33" author="Rota" box="[464,646,1864,1888]" pageId="3" pageNumber="174" refString="Rota, E., Healy, B. &amp; Erseus, C. (1998) Biogeography and taxonomy of terrestrial Enchytraeidae (Oligochaeta) in Northern Sweden, with comparative remarks on the genus Henlea. Zoologischer Anzeiger, 237, 155 - 169." type="journal article" year="1998">
Rota 
<emphasis id="8A92EA976817FFDEFDA7F8A5FDF0AD33" box="[523,579,1864,1888]" italics="true" pageId="3" pageNumber="174">et al.</emphasis>
1998
</bibRefCitation>
). Six pairs of preclitellar nephridia from 
<date id="CC5810456817FFDEFBE1F8ABFB60AD33" box="[1101,1235,1863,1888]" pageId="3" pageNumber="174" value="2011-06-05" valueMax="2011-10-05" valueMin="2011-06-05">5/6 to 10/11</date>
(in one case from 
<date id="CC5810456817FFDEFF3BF880FEB9ADD6" box="[151,266,1900,1925]" pageId="3" pageNumber="174" value="2011-05-04" valueMax="2011-10-04" valueMin="2011-05-04">4/5–10/11</date>
, absent at 5/6), anteseptale small, efferent duct origin posteroventrally (
<figureCitation id="20DD2A006817FFDEFBEFF880FB36ADD7" box="[1091,1157,1900,1925]" captionStart="FIGURE 1" captionStartId="4.[151,250,1886,1908]" captionTargetBox="[186,1403,208,1851]" captionTargetId="figure@4.[175,1411,195,1865]" captionTargetPageId="4" captionText="FIGURE 1. Henlea magnaampullacea sp. n. (A – D) and Fridericia sphaerica sp. n. (E – G). H. magnaampullacea: A. Spermathecae; B. Sperm funnel; C. Nephridium; D. Intestinal diverticulum (ventral view). F. sphaerica: E. Oesophageal appendage. F. Sperm funnel; G. Spermatheca." httpUri="https://zenodo.org/record/237088/files/figure.png" pageId="3" pageNumber="174">Fig. 1</figureCitation>
C). Coelomocytes large, rounded or ellipsoid (
<figureCitation id="20DD2A006817FFDEFE27F863FE7DADFB" box="[395,462,1935,1960]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
C), with fine brown granula in the cytoplasma (coelomocyte aggregations dark brown in transmitted light), and a peripherally thickened cell border, distinct at high magnification (
<figureCitation id="20DD2A006817FFDEFB1EF858FB47AD9F" box="[1202,1268,1972,1997]" captionStart="FIGURE 3" captionStartId="6.[151,250,1424,1446]" captionTargetBox="[178,1400,301,1393]" captionTargetId="figure@6.[178,1409,301,1403]" captionTargetPageId="6" captionText="FIGURE 3. Micrographs of Henlea magnaampullacea sp. n. A. Brain. B. Head pore (marked with arrow). C. Coelomocytes. D. Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). E. IV – IX, dorsal view of a juvenile specimen. F. Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. G. A lateral postclitellar chaetal bundle (fixed and stained). H – I. Sperm funnels. J. Spermathecae. All micrographs in vivo, except G. Scale bars 50 µm." httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="3" pageNumber="174">Fig. 3</figureCitation>
D) (length 35– 60 Μm, 
<emphasis id="8A92EA976817FFDEFF40F835FE8BADA3" box="[236,312,2009,2032]" italics="true" pageId="3" pageNumber="174">in vivo</emphasis>
, 20–30 µm, fixed).
</paragraph>
<caption id="EC99660D6810FFD9FF3BF8B2FDD4ADE1" httpUri="https://zenodo.org/record/237088/files/figure.png" pageId="4" pageNumber="175" targetBox="[186,1403,208,1851]" targetPageId="4">
<paragraph id="B85936856810FFD9FF3BF8B2FDD4ADE1" blockId="4.[151,1436,1886,1970]" pageId="4" pageNumber="175">
<emphasis id="8A92EA976810FFD9FF3BF8B2FEAAAD27" bold="true" box="[151,281,1886,1908]" pageId="4" pageNumber="175">FIGURE 1.</emphasis>
<taxonomicName id="7FE64D066810FFD9FE89F8B2FD85AD27" box="[293,566,1886,1908]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976810FFD9FE89F8B2FD85AD27" box="[293,566,1886,1908]" italics="true" pageId="4" pageNumber="175">Henlea magnaampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6810FFD9FDEEF8B3FD32AD27" box="[578,641,1887,1908]" pageId="4" pageNumber="175" rank="species">
<emphasis id="8A92EA976810FFD9FDEEF8B3FD32AD27" bold="true" box="[578,641,1887,1908]" pageId="4" pageNumber="175">sp. n.</emphasis>
</taxonomicNameLabel>
(A–D) and 
<taxonomicName id="7FE64D066810FFD9FCA1F8B2FC55AD27" box="[781,998,1886,1908]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="sphaerica" status="sp. nov.">
<emphasis id="8A92EA976810FFD9FCA1F8B2FC55AD27" box="[781,998,1886,1908]" italics="true" pageId="4" pageNumber="175">Fridericia sphaerica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6810FFD9FC5EF8B3FB82AD27" box="[1010,1073,1887,1908]" pageId="4" pageNumber="175" rank="species">
<emphasis id="8A92EA976810FFD9FC5EF8B3FB82AD27" bold="true" box="[1010,1073,1887,1908]" pageId="4" pageNumber="175">sp. n.</emphasis>
</taxonomicNameLabel>
(E–G). 
<taxonomicName id="7FE64D066810FFD9FB3CF88CFE9BADC0" authority="A. Spermathecae" authorityName="A. Spermathecae" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="magnaampullacea">
<emphasis id="8A92EA976810FFD9FB3CF88CFAC3AD27" box="[1168,1392,1886,1909]" italics="true" pageId="4" pageNumber="175">H. magnaampullacea</emphasis>
: 
<emphasis id="8A92EA976810FFD9FA2FF8B2FA2FAD20" bold="true" box="[1411,1436,1886,1907]" pageId="4" pageNumber="175">A.</emphasis>
Spermathecae
</taxonomicName>
; 
<emphasis id="8A92EA976810FFD9FE9AF892FEFEADC0" bold="true" box="[310,333,1918,1939]" pageId="4" pageNumber="175">B.</emphasis>
Sperm funnel; 
<emphasis id="8A92EA976810FFD9FE54F891FDA3ADC0" bold="true" box="[504,528,1917,1939]" pageId="4" pageNumber="175">C.</emphasis>
Nephridium; 
<emphasis id="8A92EA976810FFD9FD0BF892FD73ADC0" bold="true" box="[679,704,1918,1939]" pageId="4" pageNumber="175">D.</emphasis>
Intestinal diverticulum (ventral view). 
<taxonomicName id="7FE64D066810FFD9FBC8F892FA2FADC0" authority="E. Oesophageal" authorityName="E. Oesophageal" box="[1124,1436,1917,1939]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" kingdom="Animalia" order="Enchytraeida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="sphaerica">
<emphasis id="8A92EA976810FFD9FBC8F892FB55ADC0" box="[1124,1254,1917,1939]" italics="true" pageId="4" pageNumber="175">F. sphaerica</emphasis>
: 
<emphasis id="8A92EA976810FFD9FB5BF892FABDADC0" bold="true" box="[1271,1294,1918,1939]" pageId="4" pageNumber="175">E.</emphasis>
Oesophageal
</taxonomicName>
appendage. 
<emphasis id="8A92EA976810FFD9FEBCF870FE92ADE2" bold="true" box="[272,289,1948,1969]" pageId="4" pageNumber="175">F.</emphasis>
Sperm funnel; 
<emphasis id="8A92EA976810FFD9FE6CF870FE67ADE1" bold="true" box="[448,468,1948,1970]" pageId="4" pageNumber="175">G.</emphasis>
Spermatheca.
</paragraph>
</caption>
<caption id="EC99660D6811FFD8FF3BFA85FE57AC4B" httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="5" pageNumber="176" targetBox="[180,1398,193,1353]" targetPageId="5">
<paragraph id="B85936856811FFD8FF3BFA85FE57AC4B" blockId="5.[151,1436,1385,1560]" pageId="5" pageNumber="176">
<emphasis id="8A92EA976811FFD8FF3BFA85FEBFAF2D" bold="true" box="[151,268,1385,1407]" pageId="5" pageNumber="176">FIGURE 2</emphasis>
. Micrographs of 
<taxonomicName id="7FE64D066811FFD8FE17FA85FD76AF2C" box="[443,709,1385,1407]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="176" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976811FFD8FE17FA85FD76AF2C" box="[443,709,1385,1407]" italics="true" pageId="5" pageNumber="176">Henlea magnaampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6811FFD8FD67FA85FCB7AF2D" box="[715,772,1385,1406]" pageId="5" pageNumber="176" rank="species">
<emphasis id="8A92EA976811FFD8FD67FA85FCB7AF2D" bold="true" box="[715,772,1385,1406]" pageId="5" pageNumber="176">sp. n.</emphasis>
</taxonomicNameLabel>
(A–F) and 
<taxonomicName id="7FE64D066811FFD8FCD4FA85FBF9AF2C" box="[888,1098,1385,1407]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="176" phylum="Annelida" rank="species" species="sphaerica" status="sp. nov.">
<emphasis id="8A92EA976811FFD8FCD4FA85FBF9AF2C" box="[888,1098,1385,1407]" italics="true" pageId="5" pageNumber="176">Fridericia sphaerica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6811FFD8FBFCFA85FB31AF2D" box="[1104,1154,1385,1406]" pageId="5" pageNumber="176" rank="species">
<emphasis id="8A92EA976811FFD8FBFCFA85FB31AF2D" bold="true" box="[1104,1154,1385,1406]" pageId="5" pageNumber="176">sp.n.</emphasis>
</taxonomicNameLabel>
(G–I). 
<taxonomicName id="7FE64D066811FFD8FB61FA86FF4FAFCD" authority="A. Origin" authorityName="A. Origin" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="176" phylum="Annelida" rank="species" species="magnaapullacea">
<emphasis id="8A92EA976811FFD8FB61FA86FA26AF2C" box="[1229,1429,1385,1407]" italics="true" pageId="5" pageNumber="176">H. magnaapullacea</emphasis>
: 
<emphasis id="8A92EA976811FFD8FF3BFA64FF1CAFCE" bold="true" box="[151,175,1416,1437]" pageId="5" pageNumber="176">A.</emphasis>
Origin
</taxonomicName>
of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). 
<emphasis id="8A92EA976811FFD8FA28FA64FA28AFCE" bold="true" box="[1412,1435,1416,1437]" pageId="5" pageNumber="176">B.</emphasis>
Intestinal diverticula in VIII. 
<emphasis id="8A92EA976811FFD8FE69FA4AFE6EAFEF" bold="true" box="[453,477,1446,1468]" pageId="5" pageNumber="176">C.</emphasis>
Clitellar glands, dorsal view. 
<emphasis id="8A92EA976811FFD8FCBEFA4BFC97AFEF" bold="true" box="[786,804,1447,1468]" pageId="5" pageNumber="176">D</emphasis>
. Clitellar glands between the male copulatory organs, ventral view. 
<emphasis id="8A92EA976811FFD8FF79FA29FF5FAF89" bold="true" box="[213,236,1477,1498]" pageId="5" pageNumber="176">E.</emphasis>
Spermathecae of the holotype. 
<emphasis id="8A92EA976811FFD8FD99FA29FDFBAF89" bold="true" box="[565,584,1477,1498]" pageId="5" pageNumber="176">F.</emphasis>
Spermathecae of a fixed and stained specimens. 
<taxonomicName id="7FE64D066811FFD8FBEBFA29FA2FAF88" authority="G. Clitellar" authorityName="G. Clitellar" box="[1095,1436,1477,1499]" class="Clitellata" family="Enchytraeidae" genus="Fridericia" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="176" phylum="Annelida" rank="species" species="sphaerica">
<emphasis id="8A92EA976811FFD8FBEBFA29FAA8AF88" box="[1095,1307,1477,1499]" italics="true" pageId="5" pageNumber="176">Fridericia sphaerica</emphasis>
: 
<emphasis id="8A92EA976811FFD8FA86FA29FA8DAF88" bold="true" box="[1322,1342,1477,1499]" pageId="5" pageNumber="176">G.</emphasis>
Clitellar
</taxonomicName>
glands dorsal view. 
<emphasis id="8A92EA976811FFD8FECEFA08FECEAFAA" bold="true" box="[354,381,1508,1529]" pageId="5" pageNumber="176">H.</emphasis>
Sperm funnels. 
<emphasis id="8A92EA976811FFD8FD88FA08FD87AFAA" bold="true" box="[548,564,1508,1529]" pageId="5" pageNumber="176">I.</emphasis>
Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs 
<emphasis id="8A92EA976811FFD8FA24FA09FF72AC4B" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
, except F. Scale bars 50 µm.
</paragraph>
</caption>
<paragraph id="B85936856811FFD8FF6BF9D3FD3DAD93" blockId="5.[151,1437,1599,2021]" pageId="5" pageNumber="176">
Seminal vesicle large in XI or XI–XII. Sperm funnels cylindrical, large (
<figureCitation id="20DD2A006811FFD8FBAEF9D3FBFDAC0B" box="[1026,1102,1599,1624]" captionStart="FIGURE 1" captionStartId="4.[151,250,1886,1908]" captionTargetBox="[186,1403,208,1851]" captionTargetId="figure@4.[175,1411,195,1865]" captionTargetPageId="4" captionText="FIGURE 1. Henlea magnaampullacea sp. n. (A – D) and Fridericia sphaerica sp. n. (E – G). H. magnaampullacea: A. Spermathecae; B. Sperm funnel; C. Nephridium; D. Intestinal diverticulum (ventral view). F. sphaerica: E. Oesophageal appendage. F. Sperm funnel; G. Spermatheca." httpUri="https://zenodo.org/record/237088/files/figure.png" pageId="5" pageNumber="176">Figs. 1</figureCitation>
B, 3H–I), about 350–600 µm long and 4–6 times as long as wide, 
<emphasis id="8A92EA976811FFD8FD95F989FD35AC2F" box="[569,646,1637,1660]" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
, about ¾ as long as body diameter. Funnel length in fixed specimens 230–550 µm, 2.5–4 times longer than wide. Collar wider as funnel body. Spermatozoa about 140–170 µm long, heads 40–70 µm, 
<emphasis id="8A92EA976811FFD8FECFF941FE02AC97" box="[355,433,1709,1732]" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
. Diameter of sperm ducts 12–13 µm, 
<emphasis id="8A92EA976811FFD8FCC9F941FC0AAC97" box="[869,953,1709,1732]" italics="true" pageId="5" pageNumber="176">in vivo,</emphasis>
10–13 µm, fixed. Male copulatory organs (
<figureCitation id="20DD2A006811FFD8FF33F923FF50ACBB" box="[159,227,1743,1768]" captionStart="FIGURE 2" captionStartId="5.[151,250,1385,1407]" captionTargetBox="[180,1398,193,1353]" captionTargetId="figure@5.[180,1408,193,1364]" captionTargetPageId="5" captionText="FIGURE 2. Micrographs of Henlea magnaampullacea sp. n. (A – F) and Fridericia sphaerica sp. n. (G – I). H. magnaapullacea: A. Origin of dorsal vessel in IX (marked with black arrow), intestinal diverticula in VIII (marked with white arrow). B. Intestinal diverticula in VIII. C. Clitellar glands, dorsal view. D. Clitellar glands between the male copulatory organs, ventral view. E. Spermathecae of the holotype. F. Spermathecae of a fixed and stained specimens. Fridericia sphaerica: G. Clitellar glands dorsal view. H. Sperm funnels. I. Spermathecae; spiral loop of ectal duct canal marked with arrow. All micrographs in vivo, except F. Scale bars 50 µm." httpUri="https://zenodo.org/record/237089/files/figure.png" pageId="5" pageNumber="176">Fig. 2</figureCitation>
D) 120–190 µm long, 100–145 µm wide and 80–110 Μm high, fixed, bursal slits H-shaped, modiolus well developed. Subneural glands absent. Spermathecae (
<figureCitation id="20DD2A006811FFD8FCA9F918FCE6AD5F" box="[773,853,1780,1805]" captionStart="FIGURE 1" captionStartId="4.[151,250,1886,1908]" captionTargetBox="[186,1403,208,1851]" captionTargetId="figure@4.[175,1411,195,1865]" captionTargetPageId="4" captionText="FIGURE 1. Henlea magnaampullacea sp. n. (A – D) and Fridericia sphaerica sp. n. (E – G). H. magnaampullacea: A. Spermathecae; B. Sperm funnel; C. Nephridium; D. Intestinal diverticulum (ventral view). F. sphaerica: E. Oesophageal appendage. F. Sperm funnel; G. Spermatheca." httpUri="https://zenodo.org/record/237088/files/figure.png" pageId="5" pageNumber="176">Figs. 1</figureCitation>
A, 2E–F, 3J) very prominent, large; pear-shaped ampullae without diverticula, 90–150 µm wide, up to 200 µm when full with sperm. Ectal duct about 160–300 µm long and 40–60 (80) µm wide, slightly tapering distally, canal wide (13–15 µm, 
<emphasis id="8A92EA976811FFD8FBB9F8D1FBD2AD07" box="[1045,1121,1853,1876]" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
), (170–260 µm long, 45–60 µm wide, diameter of canal 10–12 µm in fixed specimens). Ampullae about 250–280 µm long 
<emphasis id="8A92EA976811FFD8FB4DF88CFA9CAD24" box="[1249,1327,1888,1911]" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
, merging entally and with joint opening into oesophagus in VI. At the orifice of spermathecal ducts 3–4 large glands (40–80 µm long, 
<emphasis id="8A92EA976811FFD8FEA8F844FEFDADEC" box="[260,334,1960,1983]" italics="true" pageId="5" pageNumber="176">in vivo</emphasis>
). 2–3 mature eggs at a time.
</paragraph>
</subSubSection>
<subSubSection id="F0FC650E6811FFDAFF6BF820FD81AB4E" lastPageId="7" lastPageNumber="178" pageId="5" pageNumber="176" type="distribution">
<paragraph id="B85936856811FFDBFF6BF820FD3EAAA4" blockId="5.[151,1437,1599,2021]" lastBlockId="6.[151,1436,151,248]" lastPageId="6" lastPageNumber="177" pageId="5" pageNumber="176">
<emphasis id="8A92EA976811FFD8FF6BF820FE4CADB6" bold="true" box="[199,511,1996,2021]" pageId="5" pageNumber="176">Distribution and habitat.</emphasis>
In 
<collectingCountry id="C0F176156811FFD8FD9FF821FDC4ADB7" box="[563,631,1997,2020]" name="South Korea" pageId="5" pageNumber="176">Korea</collectingCountry>
site 1: College of Agriculture &amp; Life Science, Chonbuk National University, Jeonju-si, Jeollabuk-do 
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<geoCoordinate id="DDD250426812FFDBFD7FFF7BFCC4AAE3" box="[723,887,151,176]" direction="east" orientation="longitude" pageId="6" pageNumber="177" precision="1" value="127.13233">127º07'56.4&quot;E</geoCoordinate>
, 
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asl), woodland, sites 6 and 7, Dongjinmyeon, Buan-gun, Jeollabuk-do 
<geoCoordinate id="DDD250426812FFDBFDA6FF50FD12AA86" box="[522,673,188,213]" direction="north" orientation="latitude" pageId="6" pageNumber="177" precision="1" value="35.767944">35º46'04.6&quot;N</geoCoordinate>
<geoCoordinate id="DDD250426812FFDBFD05FF50FCFDAA86" box="[681,846,188,213]" direction="east" orientation="longitude" pageId="6" pageNumber="177" precision="1" value="126.72078">126º43'14.8&quot;E</geoCoordinate>
, 
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asl and 35º46'04.2&quot; 
<geoCoordinate id="DDD250426812FFDBFB2EFF50FA95AA86" box="[1154,1318,188,213]" direction="east" orientation="longitude" pageId="6" pageNumber="177" precision="1" value="126.721">126º43'15.6&quot;E</geoCoordinate>
, 
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a 
<emphasis id="8A92EA976812FFDBFA2EFF50FA3EAA86" bold="true" box="[1410,1421,188,213]" pageId="6" pageNumber="177">s</emphasis>
l, agronomical fields, and planting garden tree.
</paragraph>
<caption id="EC99660D6812FFDBFF3BFA7CFD1EAC72" httpUri="https://zenodo.org/record/237090/files/figure.png" pageId="6" pageNumber="177" targetBox="[178,1400,301,1393]" targetPageId="6">
<paragraph id="B85936856812FFDBFF3BFA7CFD1EAC72" blockId="6.[151,1437,1424,1569]" pageId="6" pageNumber="177">
<emphasis id="8A92EA976812FFDBFF3BFA7CFEA0AFF6" bold="true" box="[151,275,1424,1446]" pageId="6" pageNumber="177">FIGURE 3.</emphasis>
Micrographs of 
<taxonomicName id="7FE64D066812FFDBFE12FA7CFD7AAFF5" box="[446,713,1424,1446]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976812FFDBFE12FA7CFD7AAFF5" box="[446,713,1424,1446]" italics="true" pageId="6" pageNumber="177">Henlea magnaampullacea</emphasis>
</taxonomicName>
<emphasis id="8A92EA976812FFDBFD7DFA7DFC99AFF6" bold="true" box="[721,810,1424,1446]" pageId="6" pageNumber="177">
<taxonomicNameLabel id="91A157EC6812FFDBFD7DFA7DFCB9AFF5" box="[721,778,1425,1446]" pageId="6" pageNumber="177" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Brain. 
<emphasis id="8A92EA976812FFDBFCDAFA7DFC3DAFF5" bold="true" box="[886,910,1425,1446]" pageId="6" pageNumber="177">B.</emphasis>
Head pore (marked with arrow). 
<emphasis id="8A92EA976812FFDBFB49FA7CFB4DAFF5" bold="true" box="[1253,1278,1424,1446]" pageId="6" pageNumber="177">C.</emphasis>
Coelomocytes. 
<emphasis id="8A92EA976812FFDBFF3BFA43FF1CAF97" bold="true" box="[151,175,1455,1476]" pageId="6" pageNumber="177">D.</emphasis>
Coelomocytes in 100 x magnification. (The characteristic thickened cell periphery is clearly visible). 
<emphasis id="8A92EA976812FFDBFB16FA43FB62AF97" bold="true" box="[1210,1233,1455,1476]" pageId="6" pageNumber="177">E.</emphasis>
IV–IX, dorsal view of a juvenile specimen. 
<emphasis id="8A92EA976812FFDBFE32FA22FE03AFB0" bold="true" box="[414,432,1486,1507]" pageId="6" pageNumber="177">F.</emphasis>
Intestinal diverticula of a subadult specimen, dorsal view, showing longitudinal and parellel arrangement of small canals. 
<emphasis id="8A92EA976812FFDBFE13FA01FE60AC50" bold="true" box="[447,467,1517,1539]" pageId="6" pageNumber="177">G.</emphasis>
A lateral postclitellar chaetal bundle (fixed and stained). 
<emphasis id="8A92EA976812FFDBFBB5FA01FBFAAC51" bold="true" box="[1049,1097,1517,1538]" pageId="6" pageNumber="177">H–I.</emphasis>
Sperm funnels. 
<emphasis id="8A92EA976812FFDBFB42FA01FAB2AC51" bold="true" box="[1262,1281,1517,1538]" pageId="6" pageNumber="177">J.</emphasis>
Spermathecae. All micrographs 
<emphasis id="8A92EA976812FFDBFEEEF9E0FE35AC72" box="[322,390,1548,1569]" italics="true" pageId="6" pageNumber="177">in vivo</emphasis>
, except G. Scale bars 50 µm.
</paragraph>
</caption>
<paragraph id="B85936856812FFDAFF6BF9A1FD81AB4E" blockId="6.[151,1437,1613,2035]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177">
<emphasis id="8A92EA976812FFDBFF6BF9A1FE61AC35" bold="true" box="[199,466,1613,1638]" pageId="6" pageNumber="177">Differential diagnosis.</emphasis>
<taxonomicName id="7FE64D066812FFDBFE72F9A2FCBAAC35" box="[478,777,1614,1638]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976812FFDBFE72F9A2FCBAAC35" box="[478,777,1614,1638]" italics="true" pageId="6" pageNumber="177">Henlea magnaampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6812FFDBFCB8F9A1FCEAAC35" box="[788,857,1613,1638]" pageId="6" pageNumber="177" rank="species">
<emphasis id="8A92EA976812FFDBFCB8F9A1FCEAAC35" bold="true" box="[788,857,1613,1638]" pageId="6" pageNumber="177">sp. n.</emphasis>
</taxonomicNameLabel>
and 6 other 
<taxonomicName id="7FE64D066812FFDBFC54F9A2FBFAAC35" box="[1016,1097,1614,1638]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="genus">
<emphasis id="8A92EA976812FFDBFC54F9A2FBFAAC35" box="[1016,1097,1614,1638]" italics="true" pageId="6" pageNumber="177">Henlea</emphasis>
</taxonomicName>
species are characterized by intestinal diverticula with multitubular substructure in VIII, and the dorsal vessel origin in IX. Among these species, 
<taxonomicName id="7FE64D066812FFDBFF5BF97BFE17ACFD" box="[247,420,1686,1710]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="ventriculosa">
<emphasis id="8A92EA976812FFDBFF5BF97BFE17ACFD" box="[247,420,1686,1710]" italics="true" pageId="6" pageNumber="177">H. ventriculosa</emphasis>
</taxonomicName>
(d' Udekem, 1854), 
<taxonomicName id="7FE64D066812FFDBFD3CF97BFBC4ACFD" authority="Nielsen &amp; Christensen, 1959" authorityName="Nielsen &amp; Christensen" authorityYear="1959" box="[656,1143,1685,1710]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="jutlandica">
<emphasis id="8A92EA976812FFDBFD3CF97BFC96ACFD" box="[656,805,1686,1710]" italics="true" pageId="6" pageNumber="177">H. jutlandica</emphasis>
<bibRefCitation id="DC774B746812FFDBFC81F979FBC4ACFD" author="Nielsen" box="[813,1143,1685,1710]" pageId="6" pageNumber="177" refString="Nielsen, C. O. &amp; Christensen, B. (1959) The Enchytraeidae. Critical revision and taxonomy of European species (studies on Enchytraeidae VII). Natura Jutlandica, 8 - 9, 1 - 160." type="book chapter" year="1959">Nielsen &amp; Christensen, 1959</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="7FE64D066812FFDBFB2FF97BFE90AC80" authority="Rodriguez &amp; Giani, 1986" authorityName="Rodriguez &amp; Giani" authorityYear="1986" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="andreae">
<emphasis id="8A92EA976812FFDBFB2FF97BFAB2ACFD" box="[1155,1281,1686,1710]" italics="true" pageId="6" pageNumber="177">H. andreae</emphasis>
Rodriguez &amp; Giani, 1986
</taxonomicName>
, 
<taxonomicName id="7FE64D066812FFDBFE9EF957FD78AC81" authority="Cernosvitov, 1929" authorityName="Cernosvitov" authorityYear="1929" box="[306,715,1722,1747]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="groenlandica">
<emphasis id="8A92EA976812FFDBFE9EF957FE5EAC81" box="[306,493,1722,1746]" italics="true" pageId="6" pageNumber="177">H. groenlandica</emphasis>
Černosvitov, 1929
</taxonomicName>
augm. 
<bibRefCitation id="DC774B746812FFDBFC8BF956FB0DAC80" author="Christensen" box="[807,1214,1722,1747]" pageId="6" pageNumber="177" refString="Christensen, B. &amp; Dozsa-Farkas, K. (2006) Invasion of terrestrial enchytraeids into two glacial tundras: North-eastern Greenland and the Actic Achipelago of Canada (Enchytraeidae, Oligochaeta). Polar Biology, 29, 454 - 466. http: // dx. doi. org / 10.1007 / s 00300 - 005 - 0076 - 3" type="journal article" year="2006">Christensen &amp; Dózsa-Farkas, 2006</bibRefCitation>
and 
<taxonomicName id="7FE64D066812FFDBFB51F957FD9EACA5" authority="Christensen &amp; Dozsa-Farkas, 1999" authorityName="Christensen &amp; Dozsa-Farkas" authorityYear="1999" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="conchifera">
<emphasis id="8A92EA976812FFDBFB51F957FA2FAC81" box="[1277,1436,1722,1746]" italics="true" pageId="6" pageNumber="177">H. conchifera</emphasis>
<bibRefCitation id="DC774B746812FFDBFF3BF931FD9EACA5" author="Christensen" box="[151,557,1757,1782]" pageId="6" pageNumber="177" refString="Christensen, B. &amp; Dozsa-Farkas, K. (1999) The enchytraeid fauna of the Palearctic tundra (Oligochaeta, Enchytraeidae). Biologiske Skrifter, The Royal Danish Academy of Sciences and Letters, 52, 1 - 37." type="journal article" year="1999">Christensen &amp; Dózsa-Farkas, 1999</bibRefCitation>
</taxonomicName>
differ from the new species in the spermathecal ampullae which are only slightly wider than the ectal ducts (
<bibRefCitation id="DC774B746812FFDBFDEEF8EEFCC7AD49" author="Schmelz" box="[578,884,1794,1819]" pageId="6" pageNumber="177" refString="Schmelz, R. M. &amp; Collado, R. (2010) A guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta). Soil Organisms, 82 (1), 1 - 176." type="journal article" year="2010">Schmelz &amp; Collado 2010</bibRefCitation>
; 
<bibRefCitation id="DC774B746812FFDBFC29F8EEFAA8AD49" author="Christensen" box="[901,1307,1794,1819]" pageId="6" pageNumber="177" refString="Christensen, B. &amp; Dozsa-Farkas, K. (1999) The enchytraeid fauna of the Palearctic tundra (Oligochaeta, Enchytraeidae). Biologiske Skrifter, The Royal Danish Academy of Sciences and Letters, 52, 1 - 37." type="journal article" year="1999">Christensen &amp; Dózsa-Farkas 1999</bibRefCitation>
, 
<bibRefCitation id="DC774B746812FFDBFA80F8EEFADAAD48" author="Dozsa-Farkas" box="[1324,1385,1794,1819]" pageId="6" pageNumber="177" refString="Christensen, B. &amp; Dozsa-Farkas, K. (2006) Invasion of terrestrial enchytraeids into two glacial tundras: North-eastern Greenland and the Actic Achipelago of Canada (Enchytraeidae, Oligochaeta). Polar Biology, 29, 454 - 466. http: // dx. doi. org / 10.1007 / s 00300 - 005 - 0076 - 3" type="journal article" year="2006">2006</bibRefCitation>
). 
<taxonomicName id="7FE64D066812FFDBFA2DF8EFFE27AD6D" authority="Burov, 1929" authorityName="Burov" authorityYear="1929" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="irkutensis">
<emphasis id="8A92EA976812FFDBFA2DF8EFFEB7AD6D" italics="true" pageId="6" pageNumber="177">H. irkutensis</emphasis>
<bibRefCitation id="DC774B746812FFDBFEA6F8CBFE27AD6D" author="Burov" box="[266,404,1830,1854]" pageId="6" pageNumber="177" refString="Burov, V. S. (1929) A new Henlea species from the vicinity of Irkutsk, Siberia (Russ. Germ. Summ.). Bulletin de l'Institut des Sciences, Section de Biologie et Geographie. Universite d'Irkutsk, 4, 99 - 132." type="journal article" year="1929">Burov, 1929</bibRefCitation>
</taxonomicName>
is much larger (60–90 segments, 
<quantity id="7F1E9B606812FFDBFCA7F8C9FC29AD6D" box="[779,922,1829,1854]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="5.5" metricValueMin="2.36" pageId="6" pageNumber="177" unit="mm" value="39.3" valueMax="55.0" valueMin="23.6">23.6–55 mm</quantity>
) (
<bibRefCitation id="DC774B746812FFDBFC1CF8CAFB89AD6D" author="Burov" box="[944,1082,1830,1854]" pageId="6" pageNumber="177" refString="Burov, V. S. (1929) A new Henlea species from the vicinity of Irkutsk, Siberia (Russ. Germ. Summ.). Bulletin de l'Institut des Sciences, Section de Biologie et Geographie. Universite d'Irkutsk, 4, 99 - 132." type="journal article" year="1929">Burov 1929</bibRefCitation>
). 
<taxonomicName id="7FE64D066812FFDBFBE1F8CBFA8EAD6D" box="[1101,1341,1830,1854]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="magnaampullacea" status="sp. nov.">
<emphasis id="8A92EA976812FFDBFBE1F8CBFA8EAD6D" box="[1101,1341,1830,1854]" italics="true" pageId="6" pageNumber="177">H. magnaampullacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157EC6812FFDBFAE8F8C9FA30AD6D" box="[1348,1411,1829,1854]" pageId="6" pageNumber="177" rank="species">
<emphasis id="8A92EA976812FFDBFAE8F8C9FA30AD6D" bold="true" box="[1348,1411,1829,1854]" pageId="6" pageNumber="177">sp. n.</emphasis>
</taxonomicNameLabel>
is most similar to 
<taxonomicName id="7FE64D066812FFDBFEE8F8A6FD12AD31" authority="Eisen, 1978" authorityName="Eisen" authorityYear="1978" box="[324,673,1866,1891]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="ochracea">
<emphasis id="8A92EA976812FFDBFEE8F8A6FDB7AD31" box="[324,516,1866,1890]" italics="true" pageId="6" pageNumber="177">Henlea ochracea</emphasis>
(Eisen, 1978)
</taxonomicName>
augm. Nurminen, 1973, because both are similarly large, have up to 6 chaetae per bundle, large sperm funnels and more robust spermathecae, but the collar of sperm funnels is not so high and bent backwards in the new species, and the spermathecal ampullae are abruptly widened, about 2–3 times wider than the diameter of ectal ducts, whereas in 
<taxonomicName id="7FE64D066812FFDBFD6AF85BFCFAAD9D" box="[710,841,1974,1998]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="177" phylum="Annelida" rank="species" species="ochracea">
<emphasis id="8A92EA976812FFDBFD6AF85BFCFAAD9D" box="[710,841,1974,1998]" italics="true" pageId="6" pageNumber="177">H ochracea</emphasis>
</taxonomicName>
the ampullae are about 1.5 times wider than the ectal ducts and the duct widens gradually into the ampullae (Dózsa-Farkas, personal observation, see 
<bibRefCitation id="DC774B746812FFDAFB54F836FEC7AAE3" author="Christensen" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177" refString="Christensen, B. &amp; Dozsa-Farkas, K. (1999) The enchytraeid fauna of the Palearctic tundra (Oligochaeta, Enchytraeidae). Biologiske Skrifter, The Royal Danish Academy of Sciences and Letters, 52, 1 - 37." type="journal article" year="1999">Christensen &amp; Dózsa-Farkas 1999</bibRefCitation>
, Fig 16). Based on the phylogenetic analysis (
<figureCitation id="20DD2A006813FFDAFC2CFF7BFC60AAE3" box="[896,979,151,176]" captionStart="FIGURE 14" captionStartId="22.[151,250,1863,1885]" captionTargetBox="[151,1434,553,1823]" captionTargetId="figure@22.[151,1435,553,1823]" captionTargetPageId="22" captionText="FIGURE 14. Henlea species, maximum likelihood (ML) trees of the ITS region (A), CO 1 (B) and H 3 genes (C). Bootstrap values greater than 50 are shown at the nodes. Sequences from new species described here appear in bold. A. ML tree of the ITS region based on 941 nucleotide positions using the T 92 + G substitution model. B. ML tree of the CO 1 gene based on 597 nucleotide positions using the GTR + I substitution model. C. ML tree of the H 3 gene based on 238 nucleotide positions using the T 92 + I substitution model. Scale bars, 0.05 substitutions per nucleotide position." httpUri="https://zenodo.org/record/237101/files/figure.png" pageId="7" pageNumber="178">Fig. 14</figureCitation>
), individuals from the new species were separated from other 
<taxonomicName id="7FE64D066813FFDAFE2AFF50FE64AA87" box="[390,471,188,212]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="178" phylum="Annelida" rank="genus">
<emphasis id="8A92EA976813FFDAFE2AFF50FE64AA87" box="[390,471,188,212]" italics="true" pageId="7" pageNumber="178">Henlea</emphasis>
</taxonomicName>
species collected in 
<collectingCountry id="C0F176156813FFDAFD6CFF51FCB4AA87" box="[704,775,189,212]" name="South Korea" pageId="7" pageNumber="178">Korea</collectingCountry>
, and were also distantly related to 
<taxonomicName id="7FE64D066813FFDAFB21FF51FAABAA87" box="[1165,1304,188,212]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="178" phylum="Annelida" rank="species" species="ochracea">
<emphasis id="8A92EA976813FFDAFB21FF51FAABAA87" box="[1165,1304,188,212]" italics="true" pageId="7" pageNumber="178">H. ochracea</emphasis>
</taxonomicName>
individuals from Alaska (identified morphologically by Dózsa-Farkas). Therefore molecular taxonomical results supported the description of a new 
<taxonomicName id="7FE64D066813FFDAFE2EFEE8FE60AB4F" box="[386,467,260,284]" class="Clitellata" family="Enchytraeidae" genus="Henlea" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="178" phylum="Annelida" rank="genus">
<emphasis id="8A92EA976813FFDAFE2EFEE8FE60AB4F" box="[386,467,260,284]" italics="true" pageId="7" pageNumber="178">Henlea</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>