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<document id="878336F84E25D5D505B86D92B1252604" ID-DOI="10.11646/zootaxa.3584.1.1" ID-ISSN="1175­5334" ID-ZooBank="072032C4-63FC-499A-A61D-58B428051302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629830673067" checkinUser="felipe" docAuthor="Michat, Mariano C., Alarie, Yves &amp; Watts, Chris H. S." docDate="2012" docId="256287ACFFF02760FBE8FE25E0418713" docLanguage="en" docName="zt03584p110.pdf" docOrigin="Zootaxa 3584 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3584.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Limbodessus macrohinkleri Watts &amp; Humphreys 2006" docType="treatment" docVersion="3" lastPageNumber="56" masterDocId="D95BFFD4FFC52757FB7FFF92E231831C" masterDocTitle="3584" masterLastPageNumber="110" masterPageNumber="1" pageNumber="54" updateTime="1699118123178" updateUser="plazi" zenodo-license-document="CLOSED">
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<treatment id="256287ACFFF02760FBE8FE25E0418713" ID-DOI="http://doi.org/10.5281/zenodo.5462380" ID-Zenodo-Dep="5462380" LSID="urn:lsid:plazi:treatment:256287ACFFF02760FBE8FE25E0418713" httpUri="http://treatment.plazi.org/id/256287ACFFF02760FBE8FE25E0418713" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<subSubSection id="E5D16531FFF02762FBE8FE25E37382EE" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="AD7436BAFFF02762FBE8FE25E10A82CD" blockId="53.[151,827,439,498]" box="[151,827,439,465]" pageId="53" pageNumber="54">
<heading id="F63C81D6FFF02762FBE8FE25E10A82CD" bold="true" box="[151,827,439,465]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
<taxonomicName id="6ACB4D39FFF02762FBE8FE25E10A82CD" ID-CoL="72BFC" authority="Watts &amp; Humphreys, 2006" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[151,827,439,465]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="macrohinkleri">
<emphasis id="9FBFEAA8FFF02762FBE8FE25E10A82CD" bold="true" box="[151,827,439,465]" pageId="53" pageNumber="54">
<emphasis id="9FBFEAA8FFF02762FBE8FE25E3D682CD" bold="true" box="[151,487,439,465]" italics="true" pageId="53" pageNumber="54">Limbodessus macrohinkleri</emphasis>
<bibRefCitation id="C95A4B4BFFF02762FA91FE25E10A82CD" author="Watts &amp; Humphreys" box="[494,827,439,465]" firstAuthor="Watts" pageId="53" pageNumber="54" pagination="123 - 185" refId="ref60425" refString="Watts, C. H. S. &amp; Humphreys, W. F. (2006) Twenty-six new Dytiscidae (Coleoptera) of the genera Limbodessus Guignot and Nirripirti Watts &amp; Humphreys, from underground waters in Australia. Transactions of the Royal Society of South Australia, 130, 123 - 185." type="journal article" year="2006">Watts &amp; Humphreys, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD7436BAFFF02762FBE8FE48E37382EE" blockId="53.[151,827,439,498]" box="[151,322,474,498]" pageId="53" pageNumber="54">(Figs 107–110)</paragraph>
</subSubSection>
<subSubSection id="E5D16531FFF02762FBE8FDB0E31081BA" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="AD7436BAFFF02762FBE8FDB0E31081BA" blockId="53.[151,1436,546,930]" pageId="53" pageNumber="54">
<emphasis id="9FBFEAA8FFF02762FBE8FDB0E34C8127" bold="true" box="[151,381,546,571]" inLineHeading="true" pageId="53" pageNumber="54" reason="1">Source of material.</emphasis>
<materialsCitation id="1DA33CE7FFF02762FAF9FDB0E31081BA" accessionNumber="BES 10493" collectingDate="2004-06-03" collectorName="W. F. Humphreys &amp; Humphreys, C. H. S &amp; C. Clay." country="Australia" county="Dawson's Well" latitude="-26.88651" location="One" longLatPrecision="1" longitude="120.16203" municipality="Lake Way Station" pageId="53" pageNumber="54" specimenCount="1">
<location id="A8146061FFF02762FAF9FDB0E3858126" LSID="urn:lsid:plazi:treatment:256287ACFFF02760FBE8FE25E0418713:A8146061FFF02762FAF9FDB0E3858126" box="[390,436,546,570]" country="Australia" county="Dawson's Well" latitude="-26.88651" longLatPrecision="1" longitude="120.16203" municipality="Lake Way Station" name="One" pageId="53" pageNumber="54">One</location>
specimen of instar II was used for the description (
<tableCitation id="E0490301FFF02762FF7EFDB0E6678126" box="[1025,1110,546,570]" captionStart="TABLE 1" captionStartId="5.[151,238,608,632]" captionText="TABLE 1. Species and instars described in the present study. The numbers under each instar indicate the specimens studied." pageId="53" pageNumber="54">Table 1</tableCitation>
).
<location id="A8146061FFF02762FF0FFDB0E6AA8126" LSID="urn:lsid:plazi:treatment:256287ACFFF02760FBE8FE25E0418713:A8146061FFF02762FF0FFDB0E6AA8126" box="[1136,1179,546,570]" country="Australia" county="Dawson's Well" latitude="-26.88651" longLatPrecision="1" longitude="120.16203" municipality="Lake Way Station" name="The" pageId="53" pageNumber="54">The</location>
larva was collected in association with adults at the following locality:
<emphasis id="9FBFEAA8FFF02762F9C3FDD4E62C8143" bold="true" box="[700,1053,582,607]" pageId="53" pageNumber="54">
<collectingCountry id="D5DC762AFFF02762F9C3FDD4E11F8143" box="[700,814,582,607]" name="Australia" pageId="53" pageNumber="54">Australia</collectingCountry>
,
<location id="A8146061FFF02762F844FDD4E1B58143" LSID="urn:lsid:plazi:treatment:256287ACFFF02760FBE8FE25E0418713:A8146061FFF02762F844FDD4E1B58143" box="[827,900,582,607]" country="Australia" county="Dawson's Well" latitude="-26.88651" longLatPrecision="1" longitude="120.16203" municipality="Lake Way Station" name="Carey" pageId="53" pageNumber="54">Carey</location>
palaeovalley
</emphasis>
,
<location id="A8146061FFF02762FF54FDD4E68D8142" LSID="urn:lsid:plazi:treatment:256287ACFFF02760FBE8FE25E0418713:A8146061FFF02762FF54FDD4E68D8142" box="[1067,1212,582,606]" country="Australia" county="Dawson's Well" latitude="-26.88651" longLatPrecision="1" longitude="120.16203" municipality="Lake Way Station" name="Hinkler Well" pageId="53" pageNumber="54">Hinkler Well</location>
calcrete,
<collectingMunicipality id="4D10ACC0FFF02762FE56FDD4E2D9819E" pageId="53" pageNumber="54">Lake Way Station</collectingMunicipality>
,
<collectingCounty id="44154E36FFF02762FB8BFDF8E393819E" box="[244,418,618,642]" pageId="53" pageNumber="54">Dawson’s Well</collectingCounty>
, BES 10493,
<geoCoordinate id="C8FF507DFFF02762F941FDF8E08B819E" box="[574,698,618,642]" degrees="26.88651" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-26.88651">26.88651S</geoCoordinate>
,
<geoCoordinate id="C8FF507DFFF02762F9B9FDF8E161819E" box="[710,848,618,642]" degrees="120.16203" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="120.16203">120.16203E</geoCoordinate>
,
<date id="D975107AFFF02762F822FDF8E1EE819E" box="[861,991,618,642]" pageId="53" pageNumber="54" value="2004-06-03">
<collectingDate id="C931E992FFF02762F822FDF8E1EE819E" box="[861,991,618,642]" pageId="53" pageNumber="54" value="2004-06-03">3–VI–2004</collectingDate>
</date>
, coll.
<collectorName id="003E536CFFF02762FF5BFDF9E6DF819E" box="[1060,1262,618,642]" pageId="53" pageNumber="54">W. F. Humphreys</collectorName>
, C. H. S. Watts and
<collectorName id="003E536CFFF02762FBB9FD1CE31081BA" box="[198,289,654,678]" pageId="53" pageNumber="54">C. Clay.</collectorName>
</materialsCitation>
</paragraph>
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<subSubSection id="E5D16531FFF02760FBB8FD20E3D28008" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="AD7436BAFFF02762FBB8FD20E3D380BE" blockId="53.[151,1436,546,930]" pageId="53" pageNumber="54">
<emphasis id="9FBFEAA8FFF02762FBB8FD20E3F081D7" bold="true" box="[199,449,690,715]" inLineHeading="true" pageId="53" pageNumber="54" reason="1">Diagnosis (instar II).</emphasis>
Larger species (HL more than
<quantity id="6A339B5FFFF02762F857FD20E1A381D6" box="[808,914,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="53" pageNumber="54" unit="mm" value="1.05">1.05 mm</quantity>
); head (Fig. 107) pyriform; nasale digitiform; half-circle of dense spinulae on lateroventral margins of nasale absent; hole-like structure on ventrodistal surface of nasale absent; lateral margins of nasale not inflated in dorsal view; lateral branches of nasale well developed; slender spinulae anterior to seta FR13 scarce (20 or less); occipital foramen strongly reduced (HW/OCW more than 2.70); occipital suture absent; lateral margins of parietal curved; secondary spiniform setae on lateral margins of parietal numerous; seta AN2 present; distal half of MN broad; setae LA3, LA4, LA5 and LA8 hair-like; secondary setae on U present (Fig. 110).
</paragraph>
<caption id="F9B46632FFF02762FBE8FC40E7A98731" pageId="53" pageNumber="54" startId="53.[151,239,978,1002]">
<paragraph id="AD7436BAFFF02762FBE8FC40E7A98731" blockId="53.[151,1436,978,1069]" pageId="53" pageNumber="54">
<emphasis id="9FBFEAA8FFF02762FBE8FC40E32B80F6" bold="true" box="[151,282,978,1002]" pageId="53" pageNumber="54">TABLE 14.</emphasis>
Number and position of secondary setae on the legs of second-instar larvae of species of
<taxonomicName id="6ACB4D39FFF02762FE73FC40E7A780F5" authorityName="Guignot" authorityYear="1939" box="[1292,1430,978,1001]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="9FBFEAA8FFF02762FE73FC40E7A780F5" box="[1292,1430,978,1001]" italics="true" pageId="53" pageNumber="54">Limbodessus</emphasis>
</taxonomicName>
. Numbers between slash marks refer to pro-, meso- and metathoracic leg, respectively. A = anterior, D = dorsal, P = posterior, Pr = proximal, V = ventral, Total = total number of secondary setae on the segment (excluding primary setae).
</paragraph>
</caption>
<paragraph id="AD7436BAFFF02762FBD0FBDFE7608492" pageId="53" pageNumber="54">
<table id="DFCBC41AFFF0D8A8FBD0FBDFE76584FF" box="[175,1364,1101,2019]" gridcols="6" gridrows="22" pageId="53" pageNumber="54">
<tr id="13FB34F8FFF0D8A8FBD0FBDFE765877F" box="[175,1364,1101,1123]" gridrow="0" pageId="53" pageNumber="54">
<th id="502A5D84FFF0D8A8FBD0FBDFE32F877F" box="[175,286,1101,1123]" gridcol="0" gridrow="0" pageId="53" pageNumber="54">Segment</th>
<th id="502A5D84FFF0D8A8FAEFFBDFE3D2877F" box="[400,483,1101,1123]" gridcol="1" gridrow="0" pageId="53" pageNumber="54">Position</th>
<th id="502A5D84FFF0D8A8F94BFBDFE0E0877F" box="[564,721,1101,1123]" gridcol="2" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="6ACB4D39FFF02762F94BFBDCE0E0877F" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[564,721,1101,1123]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="millbilliensis">
<emphasis id="9FBFEAA8FFF02762F94BFBDCE0E0877F" box="[564,721,1101,1123]" italics="true" pageId="53" pageNumber="54">L. millbilliensis</emphasis>
</taxonomicName>
</th>
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<taxonomicName id="6ACB4D39FFF02762F852FBDCE1AC877F" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[813,925,1102,1123]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FFF02762F852FBDCE1AC877F" box="[813,925,1102,1123]" italics="true" pageId="53" pageNumber="54">L. morgani</emphasis>
</taxonomicName>
</th>
<th id="502A5D84FFF0D8A8F884FBDFE6A4877F" box="[1019,1173,1101,1123]" gridcol="4" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="6ACB4D39FFF02762FF7AFBDCE6BA877F" authorityName="Watts &amp; Humphreys" authorityYear="2009" box="[1029,1163,1101,1123]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ordinarius">
<emphasis id="9FBFEAA8FFF02762FF7AFBDCE6BA877F" box="[1029,1163,1101,1123]" italics="true" pageId="53" pageNumber="54">L. ordinarius</emphasis>
</taxonomicName>
</th>
<th id="502A5D84FFF0D8A8FE7FFBDFE765877F" box="[1280,1364,1101,1123]" gridcol="5" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="6ACB4D39FFF02762FE7FFBDCE765877F" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[1280,1364,1101,1123]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pulpa">
<emphasis id="9FBFEAA8FFF02762FE7FFBDCE765877F" box="[1280,1364,1101,1123]" italics="true" pageId="53" pageNumber="54">L. pulpa</emphasis>
</taxonomicName>
</th>
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<th id="502A5D84FFF0D8A8FBD0FBEAE32F8792" box="[175,286,1144,1166]" gridcol="0" gridrow="1" pageId="53" pageNumber="54">Coxa</th>
<td id="502A5D84FFF0D8A8FAEFFBEAE3D28792" box="[400,483,1144,1166]" gridcol="1" gridrow="1" pageId="53" pageNumber="54">A</td>
<td id="502A5D84FFF0D8A8F94BFBEAE0E08792" box="[564,721,1144,1166]" gridcol="2" gridrow="1" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F852FBEAE1AC8792" box="[813,925,1144,1166]" gridcol="3" gridrow="1" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F884FBEAE6A48792" box="[1019,1173,1144,1166]" gridcol="4" gridrow="1" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8FE7FFBEAE7658792" box="[1280,1364,1144,1166]" gridcol="5" gridrow="1" pageId="53" pageNumber="54">0 / 0 / 0</td>
</tr>
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<td id="502A5D84FFF0D8A8FAEFFB30E3D287A4" box="[400,483,1186,1208]" gridcol="1" gridrow="2" pageId="53" pageNumber="54">PD</td>
<td id="502A5D84FFF0D8A8F94BFB30E0E087A4" box="[564,721,1186,1208]" gridcol="2" gridrow="2" pageId="53" pageNumber="54">2 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852FB30E1AC87A4" box="[813,925,1186,1208]" gridcol="3" gridrow="2" pageId="53" pageNumber="54">2 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884FB30E6A487A4" box="[1019,1173,1186,1208]" gridcol="4" gridrow="2" pageId="53" pageNumber="54">1–2 / 1–2 / 1–2</td>
<td id="502A5D84FFF0D8A8FE7FFB30E76587A4" box="[1280,1364,1186,1208]" gridcol="5" gridrow="2" pageId="53" pageNumber="54">1 / 1 / 1</td>
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<td id="502A5D84FFF0D8A8FAEFFB5FE3D287FF" box="[400,483,1229,1251]" gridcol="1" gridrow="3" pageId="53" pageNumber="54">V</td>
<td id="502A5D84FFF0D8A8F94BFB5FE0E087FF" box="[564,721,1229,1251]" gridcol="2" gridrow="3" pageId="53" pageNumber="54">1 / 0 / 1</td>
<td id="502A5D84FFF0D8A8F852FB5FE1AC87FF" box="[813,925,1229,1251]" gridcol="3" gridrow="3" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884FB5FE6A487FF" box="[1019,1173,1229,1251]" gridcol="4" gridrow="3" pageId="53" pageNumber="54">1 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FFB5FE76587FF" box="[1280,1364,1229,1251]" gridcol="5" gridrow="3" pageId="53" pageNumber="54">1 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FB6AE7658612" box="[175,1364,1272,1294]" gridrow="4" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFFB6AE3D28612" box="[400,483,1272,1294]" gridcol="1" gridrow="4" pageId="53" pageNumber="54">Total</td>
<td id="502A5D84FFF0D8A8F94BFB6AE0E08612" box="[564,721,1272,1294]" gridcol="2" gridrow="4" pageId="53" pageNumber="54">3 / 1 / 2</td>
<td id="502A5D84FFF0D8A8F852FB6AE1AC8612" box="[813,925,1272,1294]" gridcol="3" gridrow="4" pageId="53" pageNumber="54">2 / 2 / 2</td>
<td id="502A5D84FFF0D8A8F884FB6AE6A48612" box="[1019,1173,1272,1294]" gridcol="4" gridrow="4" pageId="53" pageNumber="54">2–3 / 2–3 / 2–3</td>
<td id="502A5D84FFF0D8A8FE7FFB6AE7658612" box="[1280,1364,1272,1294]" gridcol="5" gridrow="4" pageId="53" pageNumber="54">2 / 2 / 2</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FAB0E7658624" box="[175,1364,1314,1336]" gridrow="5" pageId="53" pageNumber="54">
<th id="502A5D84FFF0D8A8FBD0FAB0E32F8624" box="[175,286,1314,1336]" gridcol="0" gridrow="5" pageId="53" pageNumber="54">Trochanter</th>
<td id="502A5D84FFF0D8A8FAEFFAB0E3D28624" box="[400,483,1314,1336]" gridcol="1" gridrow="5" pageId="53" pageNumber="54">Pr</td>
<td id="502A5D84FFF0D8A8F94BFAB0E0E08624" box="[564,721,1314,1336]" gridcol="2" gridrow="5" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852FAB0E1AC8624" box="[813,925,1314,1336]" gridcol="3" gridrow="5" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884FAB0E6A48624" box="[1019,1173,1314,1336]" gridcol="4" gridrow="5" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FFAB0E7658624" box="[1280,1364,1314,1336]" gridcol="5" gridrow="5" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FADFE765867F" box="[175,1364,1357,1379]" gridrow="6" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFFADFE3D2867F" box="[400,483,1357,1379]" gridcol="1" gridrow="6" pageId="53" pageNumber="54">Total</td>
<td id="502A5D84FFF0D8A8F94BFADFE0E0867F" box="[564,721,1357,1379]" gridcol="2" gridrow="6" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852FADFE1AC867F" box="[813,925,1357,1379]" gridcol="3" gridrow="6" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884FADFE6A4867F" box="[1019,1173,1357,1379]" gridcol="4" gridrow="6" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FFADFE765867F" box="[1280,1364,1357,1379]" gridcol="5" gridrow="6" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FAEAE7658692" box="[175,1364,1400,1422]" gridrow="7" pageId="53" pageNumber="54">
<th id="502A5D84FFF0D8A8FBD0FAEAE32F8692" box="[175,286,1400,1422]" gridcol="0" gridrow="7" pageId="53" pageNumber="54">Femur</th>
<td id="502A5D84FFF0D8A8FAEFFAEAE3D28692" box="[400,483,1400,1422]" gridcol="1" gridrow="7" pageId="53" pageNumber="54">AD</td>
<td id="502A5D84FFF0D8A8F94BFAEAE0E08692" box="[564,721,1400,1422]" gridcol="2" gridrow="7" pageId="53" pageNumber="54">2 / 2 / 2</td>
<td id="502A5D84FFF0D8A8F852FAEAE1AC8692" box="[813,925,1400,1422]" gridcol="3" gridrow="7" pageId="53" pageNumber="54">1 / 2 / 2</td>
<td id="502A5D84FFF0D8A8F884FAEAE6A48692" box="[1019,1173,1400,1422]" gridcol="4" gridrow="7" pageId="53" pageNumber="54">2 / 2 / 2</td>
<td id="502A5D84FFF0D8A8FE7FFAEAE7658692" box="[1280,1364,1400,1422]" gridcol="5" gridrow="7" pageId="53" pageNumber="54">2 / 2 / 2</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FA30E76586A4" box="[175,1364,1442,1464]" gridrow="8" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFFA30E3D286A4" box="[400,483,1442,1464]" gridcol="1" gridrow="8" pageId="53" pageNumber="54">AV</td>
<td id="502A5D84FFF0D8A8F94BFA30E0E086A4" box="[564,721,1442,1464]" gridcol="2" gridrow="8" pageId="53" pageNumber="54">1 / 1 / 0</td>
<td id="502A5D84FFF0D8A8F852FA30E1AC86A4" box="[813,925,1442,1464]" gridcol="3" gridrow="8" pageId="53" pageNumber="54">1 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884FA30E6A486A4" box="[1019,1173,1442,1464]" gridcol="4" gridrow="8" pageId="53" pageNumber="54">1 / 1–2 / 1–2</td>
<td id="502A5D84FFF0D8A8FE7FFA30E76586A4" box="[1280,1364,1442,1464]" gridcol="5" gridrow="8" pageId="53" pageNumber="54">1 / 2 / 2</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FA5FE76586FF" box="[175,1364,1485,1507]" gridrow="9" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFFA5FE3D286FF" box="[400,483,1485,1507]" gridcol="1" gridrow="9" pageId="53" pageNumber="54">PD</td>
<td id="502A5D84FFF0D8A8F94BFA5FE0E086FF" box="[564,721,1485,1507]" gridcol="2" gridrow="9" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F852FA5FE1AC86FF" box="[813,925,1485,1507]" gridcol="3" gridrow="9" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F884FA5FE6A486FF" box="[1019,1173,1485,1507]" gridcol="4" gridrow="9" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8FE7FFA5FE76586FF" box="[1280,1364,1485,1507]" gridcol="5" gridrow="9" pageId="53" pageNumber="54">0 / 0 / 0</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0FA6AE7658512" box="[175,1364,1528,1550]" gridrow="10" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFFA6AE3D28512" box="[400,483,1528,1550]" gridcol="1" gridrow="10" pageId="53" pageNumber="54">PV</td>
<td id="502A5D84FFF0D8A8F94BFA6AE0E08512" box="[564,721,1528,1550]" gridcol="2" gridrow="10" pageId="53" pageNumber="54">0 / 0 / 1</td>
<td id="502A5D84FFF0D8A8F852FA6AE1AC8512" box="[813,925,1528,1550]" gridcol="3" gridrow="10" pageId="53" pageNumber="54">1 / 1 / 2</td>
<td id="502A5D84FFF0D8A8F884FA6AE6A48512" box="[1019,1173,1528,1550]" gridcol="4" gridrow="10" pageId="53" pageNumber="54">1 / 1 / 2</td>
<td id="502A5D84FFF0D8A8FE7FFA6AE7658512" box="[1280,1364,1528,1550]" gridcol="5" gridrow="10" pageId="53" pageNumber="54">1 / 2 / 2</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F9B0E7658524" box="[175,1364,1570,1592]" gridrow="11" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF9B0E3D28524" box="[400,483,1570,1592]" gridcol="1" gridrow="11" pageId="53" pageNumber="54">Total</td>
<td id="502A5D84FFF0D8A8F94BF9B0E0E08524" box="[564,721,1570,1592]" gridcol="2" gridrow="11" pageId="53" pageNumber="54">3 / 3 / 3</td>
<td id="502A5D84FFF0D8A8F852F9B0E1AC8524" box="[813,925,1570,1592]" gridcol="3" gridrow="11" pageId="53" pageNumber="54">3 / 4 / 5</td>
<td id="502A5D84FFF0D8A8F884F9B0E6A48524" box="[1019,1173,1570,1592]" gridcol="4" gridrow="11" pageId="53" pageNumber="54">4 / 4–5 / 5–6</td>
<td id="502A5D84FFF0D8A8FE7FF9B0E7658524" box="[1280,1364,1570,1592]" gridcol="5" gridrow="11" pageId="53" pageNumber="54">4 / 6 / 6</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F9DFE765857F" box="[175,1364,1613,1635]" gridrow="12" pageId="53" pageNumber="54">
<th id="502A5D84FFF0D8A8FBD0F9DFE32F857F" box="[175,286,1613,1635]" gridcol="0" gridrow="12" pageId="53" pageNumber="54">Tibia</th>
<td id="502A5D84FFF0D8A8FAEFF9DFE3D2857F" box="[400,483,1613,1635]" gridcol="1" gridrow="12" pageId="53" pageNumber="54">AD</td>
<td id="502A5D84FFF0D8A8F94BF9DFE0E0857F" box="[564,721,1613,1635]" gridcol="2" gridrow="12" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852F9DFE1AC857F" box="[813,925,1613,1635]" gridcol="3" gridrow="12" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884F9DFE6A4857F" box="[1019,1173,1613,1635]" gridcol="4" gridrow="12" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FF9DFE765857F" box="[1280,1364,1613,1635]" gridcol="5" gridrow="12" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F9EAE7658592" box="[175,1364,1656,1678]" gridrow="13" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF9EAE3D28592" box="[400,483,1656,1678]" gridcol="1" gridrow="13" pageId="53" pageNumber="54">AV</td>
<td id="502A5D84FFF0D8A8F94BF9EAE0E08592" box="[564,721,1656,1678]" gridcol="2" gridrow="13" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852F9EAE1AC8592" box="[813,925,1656,1678]" gridcol="3" gridrow="13" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884F9EAE6A48592" box="[1019,1173,1656,1678]" gridcol="4" gridrow="13" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FF9EAE7658592" box="[1280,1364,1656,1678]" gridcol="5" gridrow="13" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F930E76585A4" box="[175,1364,1698,1720]" gridrow="14" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF930E3D285A4" box="[400,483,1698,1720]" gridcol="1" gridrow="14" pageId="53" pageNumber="54">PD</td>
<td id="502A5D84FFF0D8A8F94BF930E0E085A4" box="[564,721,1698,1720]" gridcol="2" gridrow="14" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F852F930E1AC85A4" box="[813,925,1698,1720]" gridcol="3" gridrow="14" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F884F930E6A485A4" box="[1019,1173,1698,1720]" gridcol="4" gridrow="14" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8FE7FF930E76585A4" box="[1280,1364,1698,1720]" gridcol="5" gridrow="14" pageId="53" pageNumber="54">0 / 0 / 0</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F95FE76585FF" box="[175,1364,1741,1763]" gridrow="15" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF95FE3D285FF" box="[400,483,1741,1763]" gridcol="1" gridrow="15" pageId="53" pageNumber="54">PV</td>
<td id="502A5D84FFF0D8A8F94BF95FE0E085FF" box="[564,721,1741,1763]" gridcol="2" gridrow="15" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F852F95FE1AC85FF" box="[813,925,1741,1763]" gridcol="3" gridrow="15" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F884F95FE6A485FF" box="[1019,1173,1741,1763]" gridcol="4" gridrow="15" pageId="53" pageNumber="54">1 / 0 / 0</td>
<td id="502A5D84FFF0D8A8FE7FF95FE76585FF" box="[1280,1364,1741,1763]" gridcol="5" gridrow="15" pageId="53" pageNumber="54">0 / 0 / 0</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F96AE7658412" box="[175,1364,1784,1806]" gridrow="16" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF96AE3D28412" box="[400,483,1784,1806]" gridcol="1" gridrow="16" pageId="53" pageNumber="54">Total</td>
<td id="502A5D84FFF0D8A8F94BF96AE0E08412" box="[564,721,1784,1806]" gridcol="2" gridrow="16" pageId="53" pageNumber="54">0 / 2 / 2</td>
<td id="502A5D84FFF0D8A8F852F96AE1AC8412" box="[813,925,1784,1806]" gridcol="3" gridrow="16" pageId="53" pageNumber="54">0 / 2 / 2</td>
<td id="502A5D84FFF0D8A8F884F96AE6A48412" box="[1019,1173,1784,1806]" gridcol="4" gridrow="16" pageId="53" pageNumber="54">1 / 2 / 2–4</td>
<td id="502A5D84FFF0D8A8FE7FF96AE7658412" box="[1280,1364,1784,1806]" gridcol="5" gridrow="16" pageId="53" pageNumber="54">0 / 2 / 2</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F8B0E7658424" box="[175,1364,1826,1848]" gridrow="17" pageId="53" pageNumber="54">
<th id="502A5D84FFF0D8A8FBD0F8B0E32F8424" box="[175,286,1826,1848]" gridcol="0" gridrow="17" pageId="53" pageNumber="54">Tarsus</th>
<td id="502A5D84FFF0D8A8FAEFF8B0E3D28424" box="[400,483,1826,1848]" gridcol="1" gridrow="17" pageId="53" pageNumber="54">AD</td>
<td id="502A5D84FFF0D8A8F94BF8B0E0E08424" box="[564,721,1826,1848]" gridcol="2" gridrow="17" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F852F8B0E1AC8424" box="[813,925,1826,1848]" gridcol="3" gridrow="17" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8F884F8B0E6A48424" box="[1019,1173,1826,1848]" gridcol="4" gridrow="17" pageId="53" pageNumber="54">0 / 0 / 0</td>
<td id="502A5D84FFF0D8A8FE7FF8B0E7658424" box="[1280,1364,1826,1848]" gridcol="5" gridrow="17" pageId="53" pageNumber="54">0 / 0 / 0</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F8DFE765847F" box="[175,1364,1869,1891]" gridrow="18" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF8DFE3D2847F" box="[400,483,1869,1891]" gridcol="1" gridrow="18" pageId="53" pageNumber="54">AV</td>
<td id="502A5D84FFF0D8A8F94BF8DFE0E0847F" box="[564,721,1869,1891]" gridcol="2" gridrow="18" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852F8DFE1AC847F" box="[813,925,1869,1891]" gridcol="3" gridrow="18" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884F8DFE6A4847F" box="[1019,1173,1869,1891]" gridcol="4" gridrow="18" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FF8DFE765847F" box="[1280,1364,1869,1891]" gridcol="5" gridrow="18" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
<tr id="13FB34F8FFF0D8A8FBD0F8EAE7658492" box="[175,1364,1912,1934]" gridrow="19" pageId="53" pageNumber="54" rowspan-0="1">
<td id="502A5D84FFF0D8A8FAEFF8EAE3D28492" box="[400,483,1912,1934]" gridcol="1" gridrow="19" pageId="53" pageNumber="54">PD</td>
<td id="502A5D84FFF0D8A8F94BF8EAE0E08492" box="[564,721,1912,1934]" gridcol="2" gridrow="19" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F852F8EAE1AC8492" box="[813,925,1912,1934]" gridcol="3" gridrow="19" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8F884F8EAE6A48492" box="[1019,1173,1912,1934]" gridcol="4" gridrow="19" pageId="53" pageNumber="54">0 / 1 / 1</td>
<td id="502A5D84FFF0D8A8FE7FF8EAE7658492" box="[1280,1364,1912,1934]" gridcol="5" gridrow="19" pageId="53" pageNumber="54">0 / 1 / 1</td>
</tr>
</table>
</paragraph>
<paragraph id="AD7436BAFFF22760FBB8FF0AE3FC83B3" blockId="55.[151,1437,151,1039]" box="[199,461,151,176]" pageId="55" pageNumber="56">
<emphasis id="9FBFEAA8FFF22760FBB8FF0AE31883AC" bold="true" box="[199,297,152,176]" pageId="55" pageNumber="56">Instar I.</emphasis>
Not available.
</paragraph>
<paragraph id="AD7436BAFFF22760FBB8FF2EE32F81F3" blockId="55.[151,1437,151,1039]" pageId="55" pageNumber="56">
<emphasis id="9FBFEAA8FFF22760FBB8FF2EE31F83C8" bold="true" box="[199,302,188,212]" pageId="55" pageNumber="56">Instar II</emphasis>
(Figs 107–110). Head (Fig. 107). A3 more than 2.50 times longer than A1; A4 less than 0.70 times as long as A3; MN less than 4.70 times longer than broad; MP2 1.05–1.45 times longer than MP1; MP2 more than 3.20 times longer than MP3; LP2 less than 0.90 times as long as LP1. Legs (Figs 108–109). L3 2.25–2.95 times longer than HW; CL(L3) less than 0.40 times as long as TA. Abdomen (Fig. 110). U more than 4.15 times longer than LAS; U less than 2.15 times longer than HW; U1 more than 1.65 times longer than U2. Chaetotaxy. Anteroventral margin of nasale with 228 lamellae clypeales distributed in 3–4 rows; anterior secondary setae on proCO present; meso- and metaCO with less than 5 posterodorsal secondary setae; ventral secondary setae on pro- and mesoCO present; proFE with more than 3 posteroventral secondary setae; metaFE with more than 11 secondary setae; anterodorsal secondary setae on pro-, meso- and metaTI absent; anteroventral and posterodorsal secondary setae on proTI present; mesoTI with more than 3 posterodorsal secondary setae; metaTI with more than 2 posterodorsal secondary setae; posteroventral secondary setae on pro-, meso- and metaTI present; metaTI with more than 5 secondary setae; anterodorsal secondary setae on proTA absent; anteroventral and posteroventral secondary setae on proTA present; posterodorsal and posteroventral secondary setae on meso- and metaTA present; metaTA with more than 5 secondary setae; secondary setae on U present. Measurements and ratios that characterize the body shape are shown in
<tableCitation id="E0490301FFF22760F8AEFD26E61981D0" box="[977,1064,692,716]" captionStart="TABLE 5" captionStartId="10.[151,239,300,324]" captionText="TABLE 5. Measurements and ratios for the second-instar larvae of species of Limbodessus." pageId="55" pageNumber="56">Table 5</tableCitation>
. Secondary leg setation detailed in
<tableCitation id="E0490301FFF22760FBCBFD45E32B81F3" box="[180,282,727,751]" captionStart="TABLE 13" captionStartId="47.[151,239,916,940]" captionText="TABLE 13. Number and position of secondary setae on the legs of second-instar larvae of species of Limbodessus. Numbers between slash marks refer to pro-, meso- and metathoracic leg, respectively. A = anterior, D = dorsal, P = posterior, Pr = proximal, V = ventral, Total = total number of secondary setae on the segment (excluding primary setae)." pageId="55" pageNumber="56">Table 13</tableCitation>
.
</paragraph>
<paragraph id="AD7436BAFFF22760FBB8FD6EE3D28008" blockId="55.[151,1437,151,1039]" box="[199,483,764,788]" pageId="55" pageNumber="56">
<emphasis id="9FBFEAA8FFF22760FBB8FD6EE30F8008" bold="true" box="[199,318,764,788]" pageId="55" pageNumber="56">Instar III.</emphasis>
Not available.
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FFF22760FBB0FCB2E0418713" pageId="55" pageNumber="56" type="discussion">
<paragraph id="AD7436BAFFF22760FBB0FCB2E0418713" blockId="55.[151,1437,151,1039]" pageId="55" pageNumber="56">
<emphasis id="9FBFEAA8FFF22760FBB0FCB2E3728024" bold="true" box="[207,323,800,824]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Remarks.</emphasis>
The absence of instar III specimens of
<taxonomicName id="6ACB4D39FFF22760F86DFCB3E1E08024" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[786,977,800,824]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="macrohinkleri">
<emphasis id="9FBFEAA8FFF22760F86DFCB3E1E08024" box="[786,977,800,824]" italics="true" pageId="55" pageNumber="56">L. macrohinkleri</emphasis>
</taxonomicName>
hampers the comparison with the other species described here, particularly regarding some chaetotaxic characters.
<taxonomicName id="6ACB4D39FFF22760F896FCD6E72E8040" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[1001,1311,836,860]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="macrohinkleri">
<emphasis id="9FBFEAA8FFF22760F896FCD6E72E8040" box="[1001,1311,836,860]" italics="true" pageId="55" pageNumber="56">Limbodessus macrohinkleri</emphasis>
</taxonomicName>
belongs to the group of stygobitic species characterized by the presence of secondary setae on the urogomphus (
<taxonomicName id="6ACB4D39FFF22760FEFAFCFBE37480B8" authorityName="Watts &amp; Humphreys" authorityYear="2006" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="barwidgeeensis">
<emphasis id="9FBFEAA8FFF22760FEFAFCFBE37480B8" italics="true" pageId="55" pageNumber="56">L. barwidgeeensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760FA2DFC1FE3C480B8" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[338,501,908,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="bigbellensis">
<emphasis id="9FBFEAA8FFF22760FA2DFC1FE3C480B8" box="[338,501,908,932]" italics="true" pageId="55" pageNumber="56">L. bigbellensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760F97DFC1FE04580B8" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[514,628,909,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="9FBFEAA8FFF22760F97DFC1FE04580B8" box="[514,628,909,932]" italics="true" pageId="55" pageNumber="56">L. cooperi</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760F9FDFC1FE13D80B8" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[642,780,908,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eberhardi">
<emphasis id="9FBFEAA8FFF22760F9FDFC1FE13D80B8" box="[642,780,908,932]" italics="true" pageId="55" pageNumber="56">L. eberhardi</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760F866FC1FE18580B8" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[793,948,908,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="nambiensis">
<emphasis id="9FBFEAA8FFF22760F866FC1FE18580B8" box="[793,948,908,932]" italics="true" pageId="55" pageNumber="56">L. nambiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760F8BEFC1FE65880B8" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[961,1129,908,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="raesideensis">
<emphasis id="9FBFEAA8FFF22760F8BEFC1FE65880B8" box="[961,1129,908,932]" italics="true" pageId="55" pageNumber="56">L. raesideensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FFF22760FF09FC1FE72680B8" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[1142,1303,908,932]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="yandalensis">
<emphasis id="9FBFEAA8FFF22760FF09FC1FE72680B8" box="[1142,1303,908,932]" italics="true" pageId="55" pageNumber="56">L. yandalensis</emphasis>
</taxonomicName>
) (Fig. 110), and within this group it can be distinguished by the following combination of characters: head pyriform (Fig. 107), occipital suture absent (Fig. 107), occipital foramen strongly reduced (Fig. 107), and absence of anterodorsal secondary setae on the protibia (Fig. 108).
</paragraph>
</subSubSection>
</treatment>
</document>
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