275 lines
34 KiB
XML
275 lines
34 KiB
XML
<document id="4B669C2F15B231C9BEB0607AE81FEA45" ID-DOI="10.11646/zootaxa.5319.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203222" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690883574739" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="03F74843FF8DFFFBB8F2F8925E08CB0E" docLanguage="en" docName="zootaxa.5319.4.3.pdf" docOrigin="Zootaxa 5319 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia prolongata Shavrin 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="528" masterDocId="FFCE303BFF8CFFFFB865FFFD5F1ACE03" masterDocTitle="Two new species and faunistic records of the genus Mannerheimia Mäklin, 1880 from India, Nepal and China, and additional notes on some eastern Palaearctic species of Omaliinae Macleay, 1825 (Coleoptera: Staphylinidae)" masterLastPageNumber="536" masterPageNumber="524" pageNumber="525" updateTime="1690883991847" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="505DD93E314872FF34B9AF904EBD4923">Two new species and faunistic records of the genus Mannerheimia Mäklin, 1880 from India, Nepal and China, and additional notes on some eastern Palaearctic species of Omaliinae Macleay, 1825 (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="D16631051A5018FB103A529F11E72ECA">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="8767A18731352EBCA041A9543A24CDDE">text</mods:typeOfResource>
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<mods:title id="13054E90B07B632AA919DB09E50F5FCD">Zootaxa</mods:title>
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<mods:part id="9D7E57DEC75264F0BF76218EAA7451EC">
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<mods:date id="9CFA7BBE246250C2A1B98E2E19880A77">2023</mods:date>
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<mods:number id="48EC532BCAAD9D224A1A98FAAC926485">2023-07-27</mods:number>
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<mods:number id="5CCC7D598E65F0D40544570A9E8F6E6B">5319</mods:number>
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<mods:start id="7CC14277B3AE47CF77B8AE6B9BA6379A">524</mods:start>
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<mods:url id="A02DED249E283309DDF91BD79E3B42B6">http://dx.doi.org/10.11646/zootaxa.5319.4.3</mods:url>
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<mods:identifier id="1F0C4D69569076BC8826F35DA568ECF3" type="DOI">10.11646/zootaxa.5319.4.3</mods:identifier>
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<mods:identifier id="194DF5F57CE8E07CE26467971B23393C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F74843FF8DFFFBB8F2F8925E08CB0E" LSID="urn:lsid:plazi:treatment:03F74843FF8DFFFBB8F2F8925E08CB0E" httpUri="http://treatment.plazi.org/id/03F74843FF8DFFFBB8F2F8925E08CB0E" lastPageId="4" lastPageNumber="528" pageId="1" pageNumber="525">
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<subSubSection id="C344AADEFF8DFFFEB8F2F8925D3BC989" box="[151,545,1903,1930]" pageId="1" pageNumber="525" type="nomenclature">
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<paragraph id="8BE1F955FF8DFFFEB8F2F8925D3BC989" blockId="1.[151,545,1903,1966]" box="[151,545,1903,1930]" pageId="1" pageNumber="525">
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<heading id="D0A94E39FF8DFFFEB8F2F8925D3BC989" bold="true" box="[151,545,1903,1930]" fontSize="11" level="1" pageId="1" pageNumber="525" reason="1">
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<emphasis id="B92A2547FF8DFFFEB8F2F8925D3BC989" bold="true" box="[151,545,1903,1930]" pageId="1" pageNumber="525">
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<taxonomicName id="4C5E82D6FF8DFFFEB8F2F8925ECDC989" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,471,1903,1930]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="prolongata" status="sp. n.">
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<emphasis id="B92A2547FF8DFFFEB8F2F8925ECDC989" bold="true" box="[151,471,1903,1930]" italics="true" pageId="1" pageNumber="525">Mannerheimia prolongata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A219983CFF8DFFFEB9BAF88D5D3BC989" box="[479,545,1904,1930]" pageId="1" pageNumber="525" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C344AADEFF8DFFFEB8F2F8695E50C9AD" box="[151,330,1940,1966]" pageId="1" pageNumber="525" type="description">
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<paragraph id="8BE1F955FF8DFFFEB8F2F8695E50C9AD" blockId="1.[151,545,1903,1966]" box="[151,330,1940,1966]" pageId="1" pageNumber="525">
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(
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<figureCitation id="1365E5D0FF8DFFFEB8FAF8695FF7C9AD" box="[159,237,1940,1966]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[163,1425,505,1683]" captionTargetId="figure-127@2.[151,1436,489,1693]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Mannerheimia: 1—Mannerheimia prolongata (paratype), 2—M. punctatissima (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8203224" httpUri="https://zenodo.org/record/8203224/files/figure.png" pageId="1" pageNumber="525">Figs. 1</figureCitation>
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,
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<figureCitation id="1365E5D0FF8DFFFEB89EF8695E3DC9AD" box="[251,295,1940,1966]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[153,1434,603,1551]" captionTargetId="figure-231@3.[151,1435,597,1553]" captionTargetPageId="3" captionText="FIGURES 3–5. Aedeagus of Mannerheimia prolongata (Figs. 3–4) and M. punctatissima (Fig. 5): 3, 5—parameral view, 4—lateral view. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.8203228" httpUri="https://zenodo.org/record/8203228/files/figure.png" pageId="1" pageNumber="525">3–4</figureCitation>
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,
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<figureCitation id="1365E5D0FF8DFFFEB956F8695E58C9AD" box="[307,322,1940,1966]" captionStart="FIGURE 6" captionStartId="5.[151,250,1108,1132]" captionTargetBox="[151,1436,273,1084]" captionTargetId="figure-64@5.[151,1436,273,1084]" captionTargetPageId="5" captionText="FIGURE 6. Distribution of Mannerheimia in the Himalayan Region: M. prolongata (circle), M. punctatissima (square)." figureDoi="http://doi.org/10.5281/zenodo.8203230" httpUri="https://zenodo.org/record/8203230/files/figure.png" pageId="1" pageNumber="525">6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C344AADEFF8DFFFDB8F2F8215EA9CF42" lastPageId="2" lastPageNumber="526" pageId="1" pageNumber="525" type="materials_examined">
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<paragraph id="8BE1F955FF8DFFFEB8F2F8215A86C9F5" blockId="1.[151,1436,2012,2040]" box="[151,1436,2012,2040]" pageId="1" pageNumber="525">
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<emphasis id="B92A2547FF8DFFFEB8F2F8215E5EC9F5" bold="true" box="[151,324,2012,2038]" pageId="1" pageNumber="525">Type material.</emphasis>
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<materialsCitation id="3B36F308FF8DFFFEB92FF8215A86C9F5" box="[330,1436,2012,2040]" collectingDate="1989-08-03" country="India" location="Valley of Flowers" pageId="1" pageNumber="525" specimenCount="1" stateProvince="Uttar Pradesh" typeStatus="holotype">
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<typeStatus id="54E547F7FF8DFFFEB92FF8215EA8C9F5" box="[330,434,2012,2038]" pageId="1" pageNumber="525">Holotype</typeStatus>
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♁: ‘
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<collectingCountry id="F349B9C5FF8DFFFEB985F8215D50C9F5" box="[480,586,2012,2038]" name="India" pageId="1" pageNumber="525">INDIEN</collectingCountry>
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:
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<collectingRegion id="499A37B7FF8DFFFEBA36F8205DF6C9F5" box="[595,748,2012,2039]" country="India" name="Uttar Pradesh" pageId="1" pageNumber="525">Uttar Pradesh</collectingRegion>
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<location id="8E81AF8EFF8DFFFEBA98F8215CD8C9F5" LSID="urn:lsid:plazi:treatment:03F74843FF8DFFFBB8F2F8925E08CB0E:8E81AF8EFF8DFFFEBA98F8215CD8C9F5" box="[765,962,2012,2038]" country="India" name="Valley of Flowers" pageId="1" pageNumber="525" stateProvince="Uttar Pradesh">Valley of Flowers</location>
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<date id="FFE0DF95FF8DFFFEBBB1F8215B2CC9F5" box="[980,1078,2012,2038]" pageId="1" pageNumber="525" value="1989-08-03">
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<collectingDate id="EFA4267DFF8DFFFEBBB1F8215B2CC9F5" box="[980,1078,2012,2038]" pageId="1" pageNumber="525" value="1989-08-03">3.8.1989</collectingDate>
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</date>
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3100- | leg. A. RIEDEL 3600m’
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE1F955FF8EFFFDB8F2FF6A5E7ECED6" blockId="2.[151,1437,151,465]" pageId="2" pageNumber="526">
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<yellow label, printed>, ‘
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<typeStatus id="54E547F7FF8EFFFDB9DFFF6A5D49CEB2" box="[442,595,151,177]" pageId="2" pageNumber="526">HOLOTYPE</typeStatus>
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<taxonomicName id="4C5E82D6FF8EFFFDBA0BFF6A5C08CEB2" authorityName="Maklin" authorityYear="1880" box="[622,786,151,177]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A2547FF8EFFFDBA0BFF6A5C08CEB2" box="[622,786,151,177]" italics="true" pageId="2" pageNumber="526">Mannerheimia</emphasis>
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</taxonomicName>
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<taxonomicName id="4C5E82D6FF8EFFFDBB48FF6A5CB2CEB2" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[813,936,151,177]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="prolongata" status="sp.n.">
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<emphasis id="B92A2547FF8EFFFDBB48FF6A5CB2CEB2" box="[813,936,151,177]" italics="true" pageId="2" pageNumber="526">prolongata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A219983CFF8EFFFDBBD6FF6A5CF2CEB2" box="[947,1000,151,177]" pageId="2" pageNumber="526" rank="species">sp.n.</taxonomicNameLabel>
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| Shavrin A.V. des. 2023’ <red label, printed> (
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<collectionCode id="ED4F6190FF8EFFFDB960FF465E4DCED6" box="[261,343,187,213]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="526">SMNS</collectionCode>
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).
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</paragraph>
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<paragraph id="8BE1F955FF8EFFFDB8A2FF225EA9CF42" blockId="2.[151,1437,151,465]" pageId="2" pageNumber="526">
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<materialsCitation id="3B36F308FF8EFFFDB8A2FF225ED5CF1E" collectingDate="1989-08-03" country="India" location="Valley of Flowers" pageId="2" pageNumber="526" specimenCount="4" specimenCount-female="2" stateProvince="Uttar Pradesh" typeStatus="paratype">
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<typeStatus id="54E547F7FF8EFFFDB8A2FF225E22CEFA" box="[199,312,223,249]" pageId="2" pageNumber="526">Paratypes</typeStatus>
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: 6 ♁♁ (
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<specimenCount id="9D5832DCFF8EFFFDB9F1FF225D27CEFA" box="[404,573,223,249]" pageId="2" pageNumber="526" type="generic">two specimens</specimenCount>
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dissected),
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<specimenCount id="9D5832DCFF8EFFFDBAA1FF225C1ECEFA" box="[708,772,223,249]" pageId="2" pageNumber="526" type="female">2 ♀♀</specimenCount>
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(
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<specimenCount id="9D5832DCFF8EFFFDBB71FF225CAACEFA" box="[788,944,223,249]" pageId="2" pageNumber="526" type="generic">one specimen</specimenCount>
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dissected): same data as for holotype, with additional red printed label
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</materialsCitation>
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: ‘
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<materialsCitation id="3B36F308FF8EFFFDB9BBFEFE5EB5CF42" collectionCode="SMNS" collectorName="Shavrin A. V." country="India" pageId="2" pageNumber="526" specimenCount="2" specimenCount-female="2" stateProvince="Uttar Pradesh" typeStatus="paratype">
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<typeStatus id="54E547F7FF8EFFFDB9BBFEFE5D74CF1E" box="[478,622,259,285]" pageId="2" pageNumber="526">PARATYPE</typeStatus>
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|
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<taxonomicName id="4C5E82D6FF8EFFFDBAE0FEFE5C33CF1E" authorityName="Maklin" authorityYear="1880" box="[645,809,259,285]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A2547FF8EFFFDBAE0FEFE5C33CF1E" box="[645,809,259,285]" italics="true" pageId="2" pageNumber="526">Mannerheimia</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4C5E82D6FF8EFFFDBB25FEFE5CA1CF1E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[832,955,259,285]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="prolongata" status="sp.n.">
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<emphasis id="B92A2547FF8EFFFDBB25FEFE5CA1CF1E" box="[832,955,259,285]" italics="true" pageId="2" pageNumber="526">prolongata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A219983CFF8EFFFDBBA1FEFE5CE3CF1E" box="[964,1017,259,285]" pageId="2" pageNumber="526" rank="species">sp.n.</taxonomicNameLabel>
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|
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<collectorName id="26AB9C83FF8EFFFDBC6AFEFE5BBACF1E" box="[1039,1184,259,285]" pageId="2" pageNumber="526">Shavrin A.V.</collectorName>
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des. 2023’ (1 ♁,
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<specimenCount id="9D5832DCFF8EFFFDBD0CFEFE5A8CCF1E" box="[1385,1430,259,285]" pageId="2" pageNumber="526" type="female">1 ♀</specimenCount>
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: cSh; 5 ♁♁,
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<specimenCount id="9D5832DCFF8EFFFDB978FEDA5E53CF42" box="[285,329,295,321]" pageId="2" pageNumber="526" type="female">1 ♀</specimenCount>
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||
:
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<collectionCode id="ED4F6190FF8EFFFDB933FEDA5EBDCF42" box="[342,423,295,321]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="526">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C344AADEFF8EFFFBB8A2FEB75D45CA36" lastPageId="4" lastPageNumber="528" pageId="2" pageNumber="526" type="description">
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||
<paragraph id="8BE1F955FF8EFFFDB8A2FEB75FC3CFD2" blockId="2.[151,1437,151,465]" pageId="2" pageNumber="526">
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||
<emphasis id="B92A2547FF8EFFFDB8A2FEB75E43CF67" bold="true" box="[199,345,330,356]" pageId="2" pageNumber="526">Description.</emphasis>
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Measurements (n=9): HW: 0.55–0.57; HL: 0.35–0.37; OL: 0.17–0.19; AL (
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<typeStatus id="54E547F7FF8EFFFDBCD3FEB65A05CF66" box="[1206,1311,331,357]" pageId="2" pageNumber="526">holotype</typeStatus>
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||
): 1.06; PL: 0.50–0.54; PWmax: 0.75–0.77; PWmin: 0.71–0.76; ESL: 0.84–0.91; EW: 1.08–1.14; MTbL (
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||
<typeStatus id="54E547F7FF8EFFFDBCC4FE925A10CF8A" box="[1185,1290,367,393]" pageId="2" pageNumber="526">holotype</typeStatus>
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||
): 0.62; MTrL (
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||
<typeStatus id="54E547F7FF8EFFFDB8FAFE6E5E12CFAE" box="[159,264,403,429]" pageId="2" pageNumber="526">holotype</typeStatus>
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||
): 0.40 (MTrL 1–4: 0.20; MTrL 5: 0.20); AW: 1.12–1.22; AedL: 0.82–0.89; BL: 2.85–3.34 (
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||
<typeStatus id="54E547F7FF8EFFFDBD54FE6E5A83CFAE" box="[1329,1433,403,429]" pageId="2" pageNumber="526">holotype</typeStatus>
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||
: 3.15).
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</paragraph>
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<caption id="DF21A9DDFF8EFFFDB8F2F9485E00C8F3" ID-DOI="http://doi.org/10.5281/zenodo.8203224" ID-Zenodo-Dep="8203224" httpUri="https://zenodo.org/record/8203224/files/figure.png" pageId="2" pageNumber="526" startId="2.[151,264,1717,1741]" targetBox="[163,1425,505,1683]" targetPageId="2" targetType="figure">
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||
<paragraph id="8BE1F955FF8EFFFDB8F2F9485E00C8F3" blockId="2.[151,1436,1717,1777]" pageId="2" pageNumber="526">
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||
<emphasis id="B92A2547FF8EFFFDB8F2F9485E27C8CE" bold="true" box="[151,317,1717,1741]" pageId="2" pageNumber="526">FIGURES 1–2.</emphasis>
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Habitus of
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<taxonomicName id="4C5E82D6FF8EFFFDB9DFF9485D55C8CE" authorityName="Maklin" authorityYear="1880" box="[442,591,1717,1741]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92A2547FF8EFFFDB9DFF9485D55C8CE" box="[442,591,1717,1741]" italics="true" pageId="2" pageNumber="526">Mannerheimia</emphasis>
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||
</taxonomicName>
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||
: 1—
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<taxonomicName id="4C5E82D6FF8EFFFDBAE1F9485C8BC8CE" authorityName="Shavrin" authorityYear="2023" box="[644,913,1717,1741]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="prolongata">
|
||
<emphasis id="B92A2547FF8EFFFDBAE1F9485C8BC8CE" box="[644,913,1717,1741]" italics="true" pageId="2" pageNumber="526">Mannerheimia prolongata</emphasis>
|
||
</taxonomicName>
|
||
(paratype), 2—
|
||
<taxonomicName id="4C5E82D6FF8EFFFDBC53F94B5BFCC8CE" authorityName="Shavrin" authorityYear="2023" box="[1078,1254,1718,1741]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="punctatissima">
|
||
<emphasis id="B92A2547FF8EFFFDBC53F94B5BFCC8CE" box="[1078,1254,1718,1741]" italics="true" pageId="2" pageNumber="526">M. punctatissima</emphasis>
|
||
</taxonomicName>
|
||
(paratype). Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1F955FF8EFFFCB8A2F8BE5A69CC3E" blockId="2.[151,1437,1859,2030]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="527" pageId="2" pageNumber="526">
|
||
Body moderately small, convex and elongate (
|
||
<figureCitation id="1365E5D0FF8EFFFDBA9DF8BE5C5AC95E" box="[760,832,1859,1885]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[163,1425,505,1683]" captionTargetId="figure-127@2.[151,1436,489,1693]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Mannerheimia: 1—Mannerheimia prolongata (paratype), 2—M. punctatissima (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8203224" httpUri="https://zenodo.org/record/8203224/files/figure.png" pageId="2" pageNumber="526">Fig. 1</figureCitation>
|
||
). Coloration yellow-brown to reddish-brown, with darker head and abdomen (darker specimens with paler lateral and basal portions of pronotum and paratergites); antennomeres 7–11 and legs brownish (yellow-brown specimens with yellowish antennae and legs); mouthparts, antennomeres 1–6 and tarsi yellow. Body shiny, lacking microsculpture except for abdomen, with dense and fine transverse microreticulation, indistinct in some specimens. Head with variable, irregular and sparse punctation, finer and sparser on clypeus, denser, larger and deeper between eyes (
|
||
<specimenCount id="9D5832DCFF8FFFFCBB1CFF6A5B3DCEB2" box="[889,1063,151,177]" pageId="3" pageNumber="527" type="generic">four specimens</specimenCount>
|
||
with moderately fine and sparse punctures in middle), narrow elongate regions of head in front of ocelli sometimes lacking punctures, infraorbital portions usually with very sparse punctation, neck region with very sparse and fine punctation; punctation of pronotum moderately dense, about as that as on middle portion of head or distinctly larger and deeper, sparser in median and basal portions, some specimens with finer punctation in medioapical portion, mediobasal third sometimes lacking punctures; scutellum without or with two-three very fine punctures in middle; punctation of elytra similar to that of pronotum, finer in parascutellar portion (
|
||
<typeStatus id="54E547F7FF8FFFFCBAEFFE925DF4CF8A" box="[650,750,367,393]" pageId="3" pageNumber="527">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D5832DCFF8FFFFCBB4DFE925CCFCF8A" box="[808,981,367,393]" pageId="3" pageNumber="527" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
|
||
) and along suture, each elytron without or with indistinct four to five very tangled longitudinal rows of punctures, these rows confused in basal and apical portions; abdominal tergites with fine and moderately dense punctation, sparser and indistinct on abdominal tergites VI–VIII. Apical portion of clypeus with sparse, moderately long and erect setae, each laterobasal portions with long erect setae (missing in some specimens); apical and lateral margins of pronotum usually with several long setae; lateral portions of elytra with several short erect setae; abdomen without or with very sparse and short setation.
|
||
</paragraph>
|
||
<caption id="DF21A9DDFF8FFFFCB8F2F9D45D11C867" ID-DOI="http://doi.org/10.5281/zenodo.8203228" ID-Zenodo-Dep="8203228" httpUri="https://zenodo.org/record/8203228/files/figure.png" pageId="3" pageNumber="527" startId="3.[151,264,1577,1601]" targetBox="[153,1434,603,1551]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BE1F955FF8FFFFCB8F2F9D45D11C867" blockId="3.[151,1437,1577,1637]" pageId="3" pageNumber="527">
|
||
<emphasis id="B92A2547FF8FFFFCB8F2F9D45E24C842" bold="true" box="[151,318,1577,1601]" pageId="3" pageNumber="527">FIGURES 3–5.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C5E82D6FF8FFFFCB9B1F9D45DF8C842" authorityName="Shavrin" authorityYear="2023" box="[468,738,1577,1601]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="prolongata">
|
||
<emphasis id="B92A2547FF8FFFFCB9B1F9D45DF8C842" box="[468,738,1577,1601]" italics="true" pageId="3" pageNumber="527">Mannerheimia prolongata</emphasis>
|
||
</taxonomicName>
|
||
(Figs. 3–4) and
|
||
<taxonomicName id="4C5E82D6FF8FFFFCBBF2F9D75B50C842" authorityName="Shavrin" authorityYear="2023" box="[919,1098,1577,1601]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="punctatissima">
|
||
<emphasis id="B92A2547FF8FFFFCBBF2F9D75CB6C842" box="[919,940,1578,1601]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.
|
||
<emphasis id="B92A2547FF8FFFFCBBD8F9D75B50C842" box="[957,1098,1578,1601]" italics="true" pageId="3" pageNumber="527">punctatissima</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 5): 3, 5—parameral view, 4—lateral view. Scale bar: 0.10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1F955FF8FFFFBB8A2F94A5C9CCED6" blockId="3.[151,1437,1719,2034]" lastBlockId="4.[151,1437,151,1293]" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527">Head 1.5 times as broad as long, with slightly elevated middle portion; anteocellar foveae very short and moderately deep, about as long as ocellus or slightly shorter; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes large and convex. Ocelli moderately large, slightly convex, situated at about level or slightly behind of posterior margin of eyes; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical palpomere of maxillary palpi distinctly more than twice as long as preceding palpomere, from basal portion gradually narrowed toward acute apex. Antennae reaching apical portion of elytra when turned backwards, with relatively elongate antennomeres 4–10; basal antennomere ovoid, distinctly more than twice as long as broad, antennomere 2 slightly narrower and shorter than basal antennomere, 3 about as long 2, 4 about twice shorter than 3, 5 slightly longer and broader than 4, 6 slightly broader than 5, 7 slightly broader than 6, 8–9 distinctly shorter than 7, 10 slightly shorter than 9, apical antennomere about twice as long as preceding antennomere, from middle gradually narrowing toward acute apex.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FF225A9DCF66" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">Pronotum 1.4–1.5 times as broad as long, 1.3 times as broad as head, widest slightly in front of middle, slightly more narrowed posteriad than anteriad; apical margin somewhat straight or distinctly rounded, distinctly narrower than rounded posterior margin; anterior angles widely rounded, slightly or distinctly protruding anteriad; latero-basal margins of pronotum slightly sinuate in front of rounded posterior angles; laterobasal portions widely impressed.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FE925E0ECFD2" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">Elytra 1.2 times as broad as long, slightly to strongly broadened apicad, 1.6 times as long as pronotum; apical margins rounded; surface of each elytron in mediobasal portion longitudinally elevated between punctures in some specimens.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FE265DA9CFF6" blockId="4.[151,1437,151,1293]" box="[199,691,474,501]" pageId="4" pageNumber="528">Metatarsus 1.5 times shorter than metatibia.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FE025B6CCC3E" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">Abdomen distinctly broader than elytra, with two small oval tomentose wing-folding spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FDBA5CE1CD16" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">
|
||
Male. Apical margins of abdominal tergite VIII and sternite VIII widely concave. Aedeagus with very wide basal portion, strongly narrowed apically toward subacute apex; parameres wide, significantly longer than apex of median lobe, distinctly widened apically and narrowed in apical portions toward subacute apices, with a row of very small 10–12 spines mixed with very fine and short setae along inner sides of each apical part of parameral lobe; internal sac very long, with two fields of small spines and with two long and narrow sclerotized structures in preapical part of median lobe (
|
||
<figureCitation id="1365E5D0FF88FFFBB98AFD065D29CD16" box="[495,563,763,789]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[153,1434,603,1551]" captionTargetId="figure-231@3.[151,1435,597,1553]" captionTargetPageId="3" captionText="FIGURES 3–5. Aedeagus of Mannerheimia prolongata (Figs. 3–4) and M. punctatissima (Fig. 5): 3, 5—parameral view, 4—lateral view. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.8203228" httpUri="https://zenodo.org/record/8203228/files/figure.png" pageId="4" pageNumber="528">Fig. 3</figureCitation>
|
||
). Lateral aspect of aedeagus as in
|
||
<figureCitation id="1365E5D0FF88FFFBBBD6FD065CEDCD16" box="[947,1015,763,789]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[153,1434,603,1551]" captionTargetId="figure-231@3.[151,1435,597,1553]" captionTargetPageId="3" captionText="FIGURES 3–5. Aedeagus of Mannerheimia prolongata (Figs. 3–4) and M. punctatissima (Fig. 5): 3, 5—parameral view, 4—lateral view. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.8203228" httpUri="https://zenodo.org/record/8203228/files/figure.png" pageId="4" pageNumber="528">Fig. 4</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FCE25FE1CD5E" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII widely rounded.</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FC9B5D45CA36" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">
|
||
<emphasis id="B92A2547FF88FFFBB8A2FC9B5EA8CD82" bold="true" box="[199,434,870,897]" pageId="4" pageNumber="528">Comparative notes.</emphasis>
|
||
Based on the general shape, coloration of the body, and the shape of relatively elongate antennomeres 4–10,
|
||
<taxonomicName id="4C5E82D6FF88FFFBB91AFC765D05CDA6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[383,543,907,933]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="prolongata" status="sp.n.">
|
||
<emphasis id="B92A2547FF88FFFBB91AFC765E8CCDA7" box="[383,406,907,932]" italics="true" pageId="4" pageNumber="528">M</emphasis>
|
||
.
|
||
<emphasis id="B92A2547FF88FFFBB9C1FC765D05CDA6" box="[420,543,907,933]" italics="true" pageId="4" pageNumber="528">prolongata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92A2547FF88FFFBBA42FC765D44CDA6" bold="true" box="[551,606,907,933]" pageId="4" pageNumber="528">
|
||
<taxonomicNameLabel id="A219983CFF88FFFBBA42FC765D44CDA6" box="[551,606,907,933]" pageId="4" pageNumber="528" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C5E82D6FF88FFFBBA95FC765B33CDA6" authority="(Erichson, 1840)" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[752,1065,907,933]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="arctica">
|
||
<emphasis id="B92A2547FF88FFFBBA95FC765C1DCDA7" box="[752,775,907,932]" italics="true" pageId="4" pageNumber="528">M</emphasis>
|
||
.
|
||
<emphasis id="B92A2547FF88FFFBBB70FC765C79CDA7" box="[789,867,907,932]" italics="true" pageId="4" pageNumber="528">arctica</emphasis>
|
||
(Erichson, 1840)
|
||
</taxonomicName>
|
||
, species widely distributed in the Palaearctic Region (
|
||
<bibRefCitation id="EFCF84A4FF88FFFBB91CFC525D3ECDCA" author="Shavrin" box="[377,548,943,969]" pageId="4" pageNumber="528" pagination="301 - 333" refId="ref6833" refString="Shavrin A. V. (2021 b) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021b</bibRefCitation>
|
||
). The new species can be distinguished from it by the less transverse pronotum (the pronotum of
|
||
<taxonomicName id="4C5E82D6FF88FFFBB905FC2E5EFACDEF" authorityName="Kashcheev" authorityYear="1999" box="[352,480,979,1005]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="asiatica">
|
||
<emphasis id="B92A2547FF88FFFBB905FC2E5E6DCDEF" box="[352,375,979,1004]" italics="true" pageId="4" pageNumber="528">M</emphasis>
|
||
.
|
||
<emphasis id="B92A2547FF88FFFBB9E2FC2E5EFACDEF" box="[391,480,979,1004]" italics="true" pageId="4" pageNumber="528">asiatica</emphasis>
|
||
</taxonomicName>
|
||
is distinctly broader, twice as broad as long), narrower apical portion of the median lobe and parameres, the presence of small spines on inner margin of the each apical portion of the parameres and by the different structure of the internal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C344AADEFF88FFFBB8A2FBC35E1DCAC6" pageId="4" pageNumber="528" type="distribution">
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FBC35D93CA7E" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">
|
||
<emphasis id="B92A2547FF88FFFBB8A2FBC35E7BCA5B" bold="true" box="[199,353,1086,1112]" pageId="4" pageNumber="528">Distribution.</emphasis>
|
||
<taxonomicName id="4C5E82D6FF88FFFBB90CFBC25D8BCA5A" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[361,657,1087,1113]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="prolongata" status="sp. n.">
|
||
<emphasis id="B92A2547FF88FFFBB90CFBC25D8BCA5A" box="[361,657,1087,1113]" italics="true" pageId="4" pageNumber="528">Mannerheimia prolongata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92A2547FF88FFFBBAFCFBC25DC0CA5A" bold="true" box="[665,730,1087,1113]" pageId="4" pageNumber="528">
|
||
<taxonomicNameLabel id="A219983CFF88FFFBBAFCFBC25DC0CA5A" box="[665,730,1087,1113]" pageId="4" pageNumber="528" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="54E547F7FF88FFFBBB90FBC25B3FCA5A" box="[1013,1061,1087,1113]" pageId="4" pageNumber="528">type</typeStatus>
|
||
locality in the Valley of Flowers National Park in
|
||
<collectingRegion id="499A37B7FF88FFFBB933FB9E5EF9CA7E" box="[342,483,1123,1149]" country="India" name="Uttarakhand" pageId="4" pageNumber="528">Uttarakhand</collectingRegion>
|
||
,
|
||
<collectingCountry id="F349B9C5FF88FFFBB98BFB9E5D32CA7E" box="[494,552,1123,1149]" name="India" pageId="4" pageNumber="528">India</collectingCountry>
|
||
(
|
||
<figureCitation id="1365E5D0FF88FFFBBA52FB9E5D61CA7E" box="[567,635,1123,1149]" captionStart="FIGURE 6" captionStartId="5.[151,250,1108,1132]" captionTargetBox="[151,1436,273,1084]" captionTargetId="figure-64@5.[151,1436,273,1084]" captionTargetPageId="5" captionText="FIGURE 6. Distribution of Mannerheimia in the Himalayan Region: M. prolongata (circle), M. punctatissima (square)." figureDoi="http://doi.org/10.5281/zenodo.8203230" httpUri="https://zenodo.org/record/8203230/files/figure.png" pageId="4" pageNumber="528">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FB7B5E1DCAC6" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">
|
||
<emphasis id="B92A2547FF88FFFBB8A2FB7B5E51CAA3" bold="true" box="[199,331,1158,1184]" pageId="4" pageNumber="528">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="4CA654B0FF88FFFBBB2FFB7A5CE3CAA2" box="[842,1017,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="4" pageNumber="528" unit="m" value="3350.0" valueMax="3600.0" valueMin="3100.0">3100 to 3600 m</quantity>
|
||
a.s.l. The detailed ecological data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C344AADEFF88FFFBB8A2FB325E08CB0E" pageId="4" pageNumber="528" type="etymology">
|
||
<paragraph id="8BE1F955FF88FFFBB8A2FB325E08CB0E" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="528">
|
||
<emphasis id="B92A2547FF88FFFBB8A2FB325E54CAEA" bold="true" box="[199,334,1231,1257]" pageId="4" pageNumber="528">Etymology.</emphasis>
|
||
The specific epithet is the Latin
|
||
<emphasis id="B92A2547FF88FFFBBAD5FB325C2CCAEA" box="[688,822,1231,1257]" italics="true" pageId="4" pageNumber="528">prolongatus</emphasis>
|
||
, -
|
||
<emphasis id="B92A2547FF88FFFBBB2EFB325C43CAEB" box="[843,857,1231,1256]" italics="true" pageId="4" pageNumber="528">a</emphasis>
|
||
, -
|
||
<emphasis id="B92A2547FF88FFFBBB0AFB325C8BCAEB" box="[879,913,1231,1256]" italics="true" pageId="4" pageNumber="528">um</emphasis>
|
||
(lengthened). It alludes to the shape of elongate parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |