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<document id="9247FCA1F0A94E40383790762A7BF62B" ID-CLB-Dataset="20325" ID-DOI="10.11646/zootaxa.5104.3.4" ID-GBIF-Dataset="17de5cf1-e498-4883-b0e7-8bcd5e976d47" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332094" ID-ZooBank="9A9BA43F-3B32-4595-A95A-853B8F72FD90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594675974" checkinUser="plazi" docAuthor="Kaneko, Naoki &amp; Wada, Kaoru" docDate="2022" docId="03EBE556FFFAFFA03795928DAEE6FA2E" docLanguage="en" docName="zootaxa.5104.3.4.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaia Heller 1892" docType="key" docVersion="7" lastPageNumber="431" masterDocId="FFD29D2EFFFFFFA53702931EAB7AFFB6" masterDocTitle="A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia" masterLastPageNumber="432" masterPageNumber="426" pageNumber="431" updateTime="1699316905168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3206307D743715EE3D7B61939EF98D92">A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia</mods:title>
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<mods:namePart id="144EFB3D2EB036E68475FE2EFF8C7B8F">Kaneko, Naoki</mods:namePart>
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<treatment id="03EBE556FFFAFFA03795928DAEE6FA2E" ID-DOI="http://doi.org/10.5281/zenodo.6343755" ID-GBIF-Taxon="193892594" ID-Zenodo-Dep="6343755" LSID="urn:lsid:plazi:treatment:03EBE556FFFAFFA03795928DAEE6FA2E" httpUri="http://treatment.plazi.org/id/03EBE556FFFAFFA03795928DAEE6FA2E" lastPageNumber="431" pageId="5" pageNumber="431">
<subSubSection id="C35807CBFFFAFFA03795928DAF46FE67" pageId="5" pageNumber="431" type="nomenclature">
<paragraph id="8BFD5440FFFAFFA03795928DAF65FE1A" blockId="5.[151,1084,402,465]" box="[151,1055,402,429]" pageId="5" pageNumber="431">
<heading id="D0B5E32CFFFAFFA03795928DAF65FE1A" bold="true" box="[151,1055,402,429]" fontSize="11" level="1" pageId="5" pageNumber="431" reason="1">
<emphasis id="B9368852FFFAFFA03795928DAF65FE1A" bold="true" box="[151,1055,402,429]" pageId="5" pageNumber="431">
Key for identification of Sulawesian species of
<taxonomicName id="4C422FC3FFFAFFA035DE928CA848FE1B" authorityName="Heller" authorityYear="1892" box="[732,818,402,429]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA035DE928CA848FE1B" bold="true" box="[732,818,402,429]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
(subgenus
<taxonomicName id="4C422FC3FFFAFFA034BC928CAF6EFE1B" authorityName="Heller" authorityYear="1892" box="[958,1044,402,429]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA034BC928CAF6EFE1B" bold="true" box="[958,1044,402,429]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379592A9AF46FE67" blockId="5.[151,1084,402,465]" box="[151,1084,438,465]" pageId="5" pageNumber="431">
Species distribution is adopted from
<bibRefCitation id="EFD329B1FFFAFFA0353192A9A862FE67" author="Machatschke, J. W." box="[563,792,438,465]" pageId="5" pageNumber="431" pagination="1 - 219" refId="ref4965" refString="Machatschke, J. W. (1957) Coleoptera Lamellicornia Fam. Scarabaeidae. Subfam. Rutelinae. Tribus Anomalini. In: Wytsman, P. A. G. (Ed.), Genera Insectorum. Fasc. 199 (B). Desmet-Verteneuil, Bruxelles, pp. 1 - 219, vi pls." type="book chapter" year="1957">Machatschke (1957)</bibRefCitation>
and
<bibRefCitation id="EFD329B1FFFAFFA0344C92A9A895FE67" author="Miyake, Y." box="[846,1007,439,465]" pageId="5" pageNumber="431" pagination="33 - 54" refId="ref5081" refString="Miyake, Y. (1996) New or little known scarabaeid beetles from Southeast Asia III (Coleoptera: Geotrupidae &amp; Scarabaeidae). Lamellicornia, Tokyo, 11, 33 - 54." type="journal article" year="1996">Miyake (1996</bibRefCitation>
,
<bibRefCitation id="EFD329B1FFFAFFA034F892A9AF4EFE67" author="Miyake, Y." box="[1018,1076,439,465]" pageId="5" pageNumber="431" pagination="105 - 120" refId="ref5116" refString="Miyake, Y. (2000) New or little known scarabaeid beetles from Southeast Asia IV (Insecta: Coleoptera: Scarabaeidae). Science Report of the Research Institute of Evolutionary Biology, Tokyo, 9, 105 - 120." type="journal article" year="2000">2000</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35807CBFFFAFFA0379592EBAEE6FA2E" pageId="5" pageNumber="431" type="key">
<keyStep id="30B64CE5FFFAFFA0379592EBAEE6FD91" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA0379592EBAEE6FDBC" blockId="5.[151,1436,501,1432]" box="[151,1436,501,523]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379592EBAEE6FDBC" box="[151,1436,501,523]" pageId="5" pageNumber="431">1. Elytra with 2–3 fine longitudinal ridges or clearly punctate rows............................................... 2</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA03795910DAEE6FD91" blockId="5.[151,1436,501,1432]" box="[151,1436,530,552]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795910DAEE6FD91" box="[151,1436,530,552]" pageId="5" pageNumber="431">- Elytra without longitudinal ridges and punctate rows........................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA03795912EAEE6FDD4" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA03795912EAEE6FDF3" blockId="5.[151,1436,501,1432]" box="[151,1436,560,582]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795912EAEE6FDF3" box="[151,1436,560,582]" pageId="5" pageNumber="431">2. Pronotum without hairs or with sparse setae only on lateral sides............................................... 3</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA037959153AEE6FDD4" blockId="5.[151,1436,501,1432]" box="[151,1436,589,611]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959153AEE6FDD4" box="[151,1436,589,611]" pageId="5" pageNumber="431">- Pronotum covered with hairs.......................................................................... …9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA037959174AEE6FD2B" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA037959174AEE6FDC9" blockId="5.[151,1436,501,1432]" box="[151,1436,618,640]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959174AEE6FDC9" box="[151,1436,618,640]" pageId="5" pageNumber="431">3. Pronotum densely transversely or longitudinally rugoso-punctate................................................4</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA037959196AEE6FD2B" blockId="5.[151,1436,501,1432]" box="[151,1436,647,670]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959196AEE6FD2B" box="[151,1436,647,670]" pageId="5" pageNumber="431">- Pronotum covered with coarsely spatulate punctures, except rugoso-punctate lateral sides............................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA0379591BBAEE6FD61" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA0379591BBAEE6FD0D" blockId="5.[151,1436,501,1432]" box="[151,1436,677,699]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379591BBAEE6FD0D" box="[151,1436,677,699]" pageId="5" pageNumber="431">4. Pronotum transversely rugoso-punctate................................................................... 6</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379591DDAEE6FD61" blockId="5.[151,1436,501,1432]" box="[151,1436,706,728]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379591DDAEE6FD61" box="[151,1436,706,728]" pageId="5" pageNumber="431">
- Pronotum longitudinally rugoso-punctate........................................
<taxonomicName id="4C422FC3FFFAFFA0335391DCAFECFD61" authorityName="Heller" authorityYear="1892" box="[1105,1174,706,727]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0335391DCAFECFD61" box="[1105,1174,706,727]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0339F91DCAEE6FD61" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1181,1436,706,728]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="ritsemae" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033A791DDAFC3FD6E" box="[1189,1209,707,728]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033CF91DDAE5AFD6E" box="[1229,1312,707,728]" italics="true" pageId="5" pageNumber="431">ritsemae</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA0322791DCAEE6FD61" author="Heller, K. M." box="[1317,1436,706,728]" pageId="5" pageNumber="431" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA0379591C1AEE6FCFB" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA0379591C1AEE6FCA5" blockId="5.[151,1436,501,1432]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379591C1AEE6FCA5" pageId="5" pageNumber="431">
5. Propygidium with dense fringe of hairs along posterior margin, pygidium with two large triangular hairy patches..............................................................................
<taxonomicName id="4C422FC3FFFAFFA0331591E3AF26FCA4" authorityName="Heller" authorityYear="1892" box="[1047,1116,765,786]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0331591E3AF26FCA4" box="[1047,1116,765,786]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0336091E3AEE6FCA5" authority="Miyake, 1996" authorityName="Miyake" authorityYear="1996" box="[1122,1436,765,787]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="tondanoensis" subGenus="M">
(
<emphasis id="B9368852FFFAFFA0336891E0AF04FCA5" box="[1130,1150,766,787]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA0339091E3AE6BFCA4" box="[1170,1297,765,786]" italics="true" pageId="5" pageNumber="431">tondanoensis</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA0321591E3AEE6FCA5" author="Miyake, Y." box="[1303,1436,765,787]" pageId="5" pageNumber="431" pagination="33 - 54" refId="ref5081" refString="Miyake, Y. (1996) New or little known scarabaeid beetles from Southeast Asia III (Coleoptera: Geotrupidae &amp; Scarabaeidae). Lamellicornia, Tokyo, 11, 33 - 54." type="journal article" year="1996">Miyake, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA037959005AEE6FCFB" blockId="5.[151,1436,501,1432]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959005AEE6FCFB" pageId="5" pageNumber="431">
- Propygidium without hairs along posterior margin, pygidium with two small and thin tufts at base..........................................................................................
<taxonomicName id="4C422FC3FFFAFFA0331A9026AF27FCFB" authorityName="Heller" authorityYear="1892" box="[1048,1117,824,845]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0331A9026AF27FCFB" box="[1048,1117,824,845]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA033669026AEE6FCFB" authority="Miyake, 2000" authorityName="Miyake" authorityYear="2000" box="[1124,1436,824,846]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="castanoptera" subGenus="M">
(
<emphasis id="B9368852FFFAFFA0336E9026AFFAFCFB" box="[1132,1152,824,845]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033969026AE6BFCFB" box="[1172,1297,824,845]" italics="true" pageId="5" pageNumber="431">castanoptera</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA032159026AEE6FCFB" author="Miyake, Y." box="[1303,1436,824,846]" pageId="5" pageNumber="431" pagination="105 - 120" refId="ref5116" refString="Miyake, Y. (2000) New or little known scarabaeid beetles from Southeast Asia IV (Insecta: Coleoptera: Scarabaeidae). Science Report of the Research Institute of Evolutionary Biology, Tokyo, 9, 105 - 120." type="journal article" year="2000">Miyake, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA03795904BAEE6FC31" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA03795904BAEE6FCDC" blockId="5.[151,1436,501,1432]" box="[151,1436,853,875]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795904BAEE6FCDC" box="[151,1436,853,875]" pageId="5" pageNumber="431">6. Propygidium with dense fringe of hairs along posterior margin................................................. 7</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA03795906DAEE6FC31" blockId="5.[151,1436,501,1432]" box="[151,1436,882,904]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795906DAEE6FC31" box="[151,1436,882,904]" pageId="5" pageNumber="431">- Propygidium without hairs along posterior margin............................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA03795908EAEE6FC75" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA03795908EAEE6FC13" blockId="5.[151,1436,501,1432]" box="[151,1436,911,934]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795908EAEE6FC13" box="[151,1436,911,934]" pageId="5" pageNumber="431">
7. Pygidium with two hairy tufts at base.......................................
<taxonomicName id="4C422FC3FFFAFFA0332E908EAF0BFC13" authorityName="Heller" authorityYear="1892" box="[1068,1137,912,933]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0332E908EAF0BFC13" box="[1068,1137,912,933]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0337A908EAEE6FC13" authority="(Boisduval, 1835)" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[1144,1436,911,934]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="nigrita" subGenus="M">
(
<emphasis id="B9368852FFFAFFA03382908EAFEEFC13" box="[1152,1172,912,933]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033AA908EAF93FC13" box="[1192,1257,912,933]" italics="true" pageId="5" pageNumber="431">nigrita</emphasis>
(Boisduval, 1835)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379590B3AEE6FC75" blockId="5.[151,1436,501,1432]" box="[151,1436,941,963]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379590B3AEE6FC75" box="[151,1436,941,963]" pageId="5" pageNumber="431">
- Pygidium bears scattered hairs, without hairy tufts …....................... ..
<taxonomicName id="4C422FC3FFFAFFA0330190B3AF32FC74" authorityName="Heller" authorityYear="1892" box="[1027,1096,941,962]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0330190B3AF32FC74" box="[1027,1096,941,962]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0334C90B3AEE6FC75" authority="(Schaufuss, 1885)" baseAuthorityName="Schaufuss" baseAuthorityYear="1885" box="[1102,1436,941,963]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="gnomonica" subGenus="M">
(
<emphasis id="B9368852FFFAFFA0335490B0AF10FC75" box="[1110,1130,942,963]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA0337C90B0AF93FC75" box="[1150,1257,942,963]" italics="true" pageId="5" pageNumber="431">gnomonica</emphasis>
(Schaufuss, 1885)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA0379590D4AEE6FC48" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA0379590D4AEE6FC56" blockId="5.[151,1436,501,1432]" box="[151,1436,970,992]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379590D4AEE6FC56" box="[151,1436,970,992]" pageId="5" pageNumber="431">
8. Pygidium with two small hairy tufts at base......................................
<taxonomicName id="4C422FC3FFFAFFA0335090D4AFEDFC69" authorityName="Heller" authorityYear="1892" box="[1106,1175,970,991]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0335090D4AFEDFC69" box="[1106,1175,970,991]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0339F90D4AEE6FC56" authority="Miyake, 1996" authorityName="Miyake" authorityYear="1996" box="[1181,1436,970,992]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="cyanea" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033A790D5AFC3FC56" box="[1189,1209,971,992]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033CF90D5AE6BFC56" box="[1229,1297,971,992]" italics="true" pageId="5" pageNumber="431">cyanea</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA0321590D4AEE6FC56" author="Miyake, Y." box="[1303,1436,970,992]" pageId="5" pageNumber="431" pagination="33 - 54" refId="ref5081" refString="Miyake, Y. (1996) New or little known scarabaeid beetles from Southeast Asia III (Coleoptera: Geotrupidae &amp; Scarabaeidae). Lamellicornia, Tokyo, 11, 33 - 54." type="journal article" year="1996">Miyake, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379590F6AEE6FC48" blockId="5.[151,1436,501,1432]" box="[151,1436,1000,1022]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379590F6AEE6FC48" box="[151,1436,1000,1022]" pageId="5" pageNumber="431">
- Pygidium with two large triangular hairy patches................................... ..
<taxonomicName id="4C422FC3FFFAFFA0337290F6AFCFFC4B" authorityName="Heller" authorityYear="1892" box="[1136,1205,1000,1021]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0337290F6AFCFFC4B" box="[1136,1205,1000,1021]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA033BE90F6AEE6FC48" authority="Miyake, 1996" authorityName="Miyake" authorityYear="1996" box="[1212,1436,1000,1022]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="taoi" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033C690F6AFA2FC4B" box="[1220,1240,1000,1021]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033EE90F6AE6BFC4B" box="[1260,1297,1000,1021]" italics="true" pageId="5" pageNumber="431">taoi</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA0321590F6AEE6FC48" author="Miyake, Y." box="[1303,1436,1000,1022]" pageId="5" pageNumber="431" pagination="33 - 54" refId="ref5081" refString="Miyake, Y. (1996) New or little known scarabaeid beetles from Southeast Asia III (Coleoptera: Geotrupidae &amp; Scarabaeidae). Lamellicornia, Tokyo, 11, 33 - 54." type="journal article" year="1996">Miyake, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA03795971BAEE6FBE3" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA03795971BAEE6FBAC" blockId="5.[151,1436,501,1432]" box="[151,1436,1029,1051]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795971BAEE6FBAC" box="[151,1436,1029,1051]" pageId="5" pageNumber="431">9. Pronotum densely covered with hairs.................................................................... 10</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA03795973DAEE6FBE3" blockId="5.[151,1436,501,1432]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795973DAEE6FBE3" pageId="5" pageNumber="431">
- Pronotum covered with scattered hairs, with coarse horseshoe-shaped punctures under the hairs …...........................................................................................
<taxonomicName id="4C422FC3FFFAFFA0332C975EAF09FBE3" authorityName="Heller" authorityYear="1892" box="[1070,1139,1088,1109]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0332C975EAF09FBE3" box="[1070,1139,1088,1109]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0337B975EAEE6FBE3" authority="Miyake, 2000" authorityName="Miyake" authorityYear="2000" box="[1145,1436,1088,1110]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="macassara" subGenus="M">
(
<emphasis id="B9368852FFFAFFA03383975EAFEFFBE3" box="[1153,1173,1088,1109]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033AB975EAE6BFBE3" box="[1193,1297,1088,1109]" italics="true" pageId="5" pageNumber="431">macassara</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA03215975EAEE6FBE3" author="Miyake, Y." box="[1303,1436,1088,1110]" pageId="5" pageNumber="431" pagination="105 - 120" refId="ref5116" refString="Miyake, Y. (2000) New or little known scarabaeid beetles from Southeast Asia IV (Insecta: Coleoptera: Scarabaeidae). Science Report of the Research Institute of Evolutionary Biology, Tokyo, 9, 105 - 120." type="journal article" year="2000">Miyake, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA037959743AEE6FB39" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA037959743AEE6FBC4" blockId="5.[151,1436,501,1432]" box="[151,1436,1117,1139]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959743AEE6FBC4" box="[151,1436,1117,1139]" pageId="5" pageNumber="431">10. Pronotum covered with longitudinal aciculate cracks, with fine punctures between these........................... 11</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA037959765AEE6FB39" blockId="5.[151,1436,501,1432]" box="[151,1436,1146,1168]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959765AEE6FB39" box="[151,1436,1146,1168]" pageId="5" pageNumber="431">- Pronotum covered with dense grained or annulate punctures...................................................12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA037959786AEE6FB7C" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA037959786AEE6FB1B" blockId="5.[151,1436,501,1432]" box="[151,1436,1175,1198]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959786AEE6FB1B" box="[151,1436,1175,1198]" pageId="5" pageNumber="431">
11. Pronotal hairs arranged in relatively low density...............................
<taxonomicName id="4C422FC3FFFAFFA0332C9786AF09FB1B" authorityName="Heller" authorityYear="1892" box="[1070,1139,1176,1197]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0332C9786AF09FB1B" box="[1070,1139,1176,1197]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0337B9786AEE6FB1B" authority="(Schaufuss, 1885)" baseAuthorityName="Schaufuss" baseAuthorityYear="1885" box="[1145,1436,1175,1198]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="ornata" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033839786AFEFFB1B" box="[1153,1173,1176,1197]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033AB9786AF93FB1B" box="[1193,1257,1176,1197]" italics="true" pageId="5" pageNumber="431">ornata</emphasis>
(Schaufuss, 1885)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379597ABAEE6FB7C" blockId="5.[151,1436,501,1432]" box="[151,1436,1205,1227]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379597ABAEE6FB7C" box="[151,1436,1205,1227]" pageId="5" pageNumber="431">
- Pronotal hairs arranged in extremely high density................................ ..
<taxonomicName id="4C422FC3FFFAFFA0335B97ABAFE4FB7C" authorityName="Heller" authorityYear="1892" box="[1113,1182,1205,1226]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0335B97ABAFE4FB7C" box="[1113,1182,1205,1226]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA033A797ABAEE6FB7C" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1189,1436,1205,1227]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="trajecta" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033AF97A8AFBBFB7D" box="[1197,1217,1206,1227]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033D797A8AE5AFB7D" box="[1237,1312,1206,1227]" italics="true" pageId="5" pageNumber="431">trajecta</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA0322797ABAEE6FB7C" author="Heller, K. M." box="[1317,1436,1205,1227]" pageId="5" pageNumber="431" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA0379597CCAEE6FAB0" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA0379597CCAEE6FB5E" blockId="5.[151,1436,501,1432]" box="[151,1436,1234,1256]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379597CCAEE6FB5E" box="[151,1436,1234,1256]" pageId="5" pageNumber="431">
12. Pronotum covered with dense grained punctures.................................
<taxonomicName id="4C422FC3FFFAFFA0333C97CCAFF9FB51" authorityName="Heller" authorityYear="1892" box="[1086,1155,1234,1255]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0333C97CCAFF9FB51" box="[1086,1155,1234,1255]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0338B97CCAEE6FB5E" authority="(Candeze, 1869)" baseAuthorityName="Candeze" baseAuthorityYear="1869" box="[1161,1436,1234,1256]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="vestita" subGenus="M">
(
<emphasis id="B9368852FFFAFFA0339397CDAFDFFB5E" box="[1169,1189,1235,1256]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033BB97CDAF82FB5E" box="[1209,1272,1235,1256]" italics="true" pageId="5" pageNumber="431">vestita</emphasis>
(Candèze, 1869)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA0379597EEAEE6FAB0" blockId="5.[151,1436,501,1432]" box="[151,1436,1264,1286]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA0379597EEAEE6FAB0" box="[151,1436,1264,1286]" pageId="5" pageNumber="431">
- Pronotum covered with dense annulate punctures................................
<taxonomicName id="4C422FC3FFFAFFA0334097EEAFFDFAB3" authorityName="Heller" authorityYear="1892" box="[1090,1159,1264,1285]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0334097EEAFFDFAB3" box="[1090,1159,1264,1285]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0338F97EEAEE6FAB0" authority="(Ohaus, 1936)" baseAuthorityName="Ohaus" baseAuthorityYear="1936" box="[1165,1436,1264,1286]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="celebica" subGenus="M">
(
<emphasis id="B9368852FFFAFFA0339797EEAFD3FAB3" box="[1173,1193,1264,1285]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033BF97EEAE77FAB3" box="[1213,1293,1264,1285]" italics="true" pageId="5" pageNumber="431">celebica</emphasis>
(Ohaus, 1936)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA037959613AEE6FA89" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA037959613AEE6FA94" blockId="5.[151,1436,501,1432]" box="[151,1436,1293,1315]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959613AEE6FA94" box="[151,1436,1293,1315]" pageId="5" pageNumber="431">13. Pronotum covered with dense granulose texture.............................................................14</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA037959635AEE6FA89" blockId="5.[151,1436,501,1432]" box="[151,1436,1322,1344]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959635AEE6FA89" box="[151,1436,1322,1344]" pageId="5" pageNumber="431">
- Pronotum fairly densely punctured, with lateral sides showing aciculate punctures.....
<taxonomicName id="4C422FC3FFFAFFA0333D9634AFFEFA89" authorityName="Heller" authorityYear="1892" box="[1087,1156,1322,1343]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA0333D9634AFFEFA89" box="[1087,1156,1322,1343]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA033889634AEE6FA89" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1162,1436,1322,1344]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="vandepolli" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033909635AFDCFAF6" box="[1170,1190,1323,1344]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033B89634AE65FA89" box="[1210,1311,1322,1343]" italics="true" pageId="5" pageNumber="431">vandepolli</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA032279634AEE6FA89" author="Heller, K. M." box="[1317,1436,1322,1344]" pageId="5" pageNumber="431" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B64CE5FFFAFFA037959656AEE6FA2E" pageId="5" pageNumber="431">
<paragraph id="8BFD5440FFFAFFA037959656AEE6FAEB" blockId="5.[151,1436,501,1432]" box="[151,1436,1352,1374]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA037959656AEE6FAEB" box="[151,1436,1352,1374]" pageId="5" pageNumber="431">
14. Pronotum with weak oblique impression on each side............................
<taxonomicName id="4C422FC3FFFAFFA033419656AFF2FAEB" authorityName="Heller" authorityYear="1892" box="[1091,1160,1352,1373]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA033419656AFF2FAEB" box="[1091,1160,1352,1373]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0338C9656AEE6FAEB" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1166,1436,1352,1374]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M">
(
<emphasis id="B9368852FFFAFFA033949656AFD0FAEB" box="[1174,1194,1352,1373]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA033BC9656AE65FAEB" box="[1214,1311,1352,1373]" italics="true" pageId="5" pageNumber="431">simulatrix</emphasis>
<bibRefCitation id="EFD329B1FFFAFFA032279656AEE6FAEB" author="Heller, K. M." box="[1317,1436,1352,1374]" pageId="5" pageNumber="431" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFD5440FFFAFFA03795967BAEE6FA2E" blockId="5.[151,1436,501,1432]" pageId="5" pageNumber="431">
<keyLead id="30B3F775FFFAFFA03795967BAEE6FA2E" pageId="5" pageNumber="431">
- Pronotum with strong oblique impression on each side and bearing sparkly gloss................................................................................................
<taxonomicName id="4C422FC3FFFAFFA034BC969CAF79FA21" authorityName="Heller" authorityYear="1892" box="[958,1027,1410,1431]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B9368852FFFAFFA034BC969CAF79FA21" box="[958,1027,1410,1431]" italics="true" pageId="5" pageNumber="431">Malaia</emphasis>
</taxonomicName>
<taxonomicName id="4C422FC3FFFAFFA0330B969CAE5AFA2E" authority="Kaneko &amp; Wada" authorityName="Kaneko &amp; Wada" authorityYear="2022" box="[1033,1312,1410,1432]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="medea" status="sp. nov." subGenus="M">
(
<emphasis id="B9368852FFFAFFA03313969DAF5FFA2E" box="[1041,1061,1411,1432]" italics="true" pageId="5" pageNumber="431">M</emphasis>
.)
<emphasis id="B9368852FFFAFFA0333B969CAF02FA21" box="[1081,1144,1410,1431]" italics="true" pageId="5" pageNumber="431">medea</emphasis>
Kaneko &amp; Wada
</taxonomicName>
,
<taxonomicNameLabel id="A2053529FFFAFFA0322B969DAEE6FA2E" box="[1321,1436,1410,1432]" pageId="5" pageNumber="431" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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