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<document id="81232C39EFD3FB902A2591A23AC3DF45" ID-CLB-Dataset="28034" ID-DOI="10.11646/zootaxa.4508.4.1" ID-GBIF-Dataset="a7ff59ca-84b3-4132-8a5d-367b14c84cee" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607312" ID-ZooBank="D268E210-6688-4B41-9A76-AE54C2FC07BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586215440" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="03EB87F8F169FFAEFF31FC017EE44B8E" docLanguage="en" docName="zootaxa.4508.4.1.pdf" docOrigin="Zootaxa 4508 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium denticulatum Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="478" masterDocId="FFD2FF80F173FFB5FFA6FFEC7C18480E" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group" masterLastPageNumber="506" masterPageNumber="451" pageNumber="477" updateTime="1698713119729" updateUser="ExternalLinkService">
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<mods:title id="AB8A66E2F14685810FA102BD2BD0AC66">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group</mods:title>
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<mods:namePart id="CE24D177E18385CC1758C6EE72BB26CF">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="1C74C14C4A7CF3ACE90B2213F2991615">Smetana, Aleš</mods:namePart>
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<mods:typeOfResource id="94537F4C52CBA2305D1A69944EA93ABC">text</mods:typeOfResource>
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<mods:title id="6BF69F6C0FFE99D472FDB68A572F2FCB">Zootaxa</mods:title>
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<mods:date id="F78B7FAFAD7CCBDD5DD073323E82A89B">2018</mods:date>
<mods:detail id="724B42DB0D6E3D9AB75477575C897163" type="pubDate">
<mods:number id="34A9F10B5344E0D6AF601AB720D3000F">2018-11-02</mods:number>
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<mods:detail id="D5F94E573443451E81ABF831BC4E432D" type="volume">
<mods:number id="A5A6A0F7B3CE6C5EE66D436D1B6DDF6F">4508</mods:number>
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<mods:classification id="822BEAA8C5950A3A6BBDA2EE8F8D8D40">journal article</mods:classification>
<mods:identifier id="4E1D799484FC234C58EC6172F1E24373" type="CLB-Dataset">28034</mods:identifier>
<mods:identifier id="62C7F420B615B4C6670D4757FDA9CC50" type="DOI">10.11646/zootaxa.4508.4.1</mods:identifier>
<mods:identifier id="EFC89AC1797CF5D209C15640461D6E00" type="GBIF-Dataset">a7ff59ca-84b3-4132-8a5d-367b14c84cee</mods:identifier>
<mods:identifier id="0EFD08C7C34DB85C04EF53F4FBCC4A6B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03EB87F8F169FFAEFF31FC017EE44B8E" ID-DOI="http://doi.org/10.5281/zenodo.6495070" ID-GBIF-Taxon="154496007" ID-Zenodo-Dep="6495070" LSID="urn:lsid:plazi:treatment:03EB87F8F169FFAEFF31FC017EE44B8E" httpUri="http://treatment.plazi.org/id/03EB87F8F169FFAEFF31FC017EE44B8E" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<subSubSection id="C3586565F169FFAFFF31FC017F174C09" box="[151,783,1005,1031]" pageId="26" pageNumber="477" type="nomenclature">
<paragraph id="8BFD36EEF169FFAFFF31FC017F174C09" blockId="26.[151,783,1005,1064]" box="[151,783,1005,1031]" pageId="26" pageNumber="477">
<heading id="D0B58182F169FFAFFF31FC017F174C09" bold="true" box="[151,783,1005,1031]" fontSize="11" level="1" pageId="26" pageNumber="477" reason="1">
<emphasis id="B936EAFCF169FFAFFF31FC017F174C09" bold="true" box="[151,783,1005,1031]" pageId="26" pageNumber="477">
<taxonomicName id="4C424D6DF169FFAFFF31FC017EDF4C09" ID-CoL="67CCD" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,711,1005,1031]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B936EAFCF169FFAFFF31FC017DDF4C09" bold="true" box="[151,455,1005,1031]" italics="true" pageId="26" pageNumber="477">Anthobium denticulatum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="A2055787F169FFAFFD72FC017F174C09" box="[724,783,1005,1031]" pageId="26" pageNumber="477" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586565F169FFAFFF31FBE37D404C26" box="[151,344,1039,1064]" pageId="26" pageNumber="477" type="description">
<paragraph id="8BFD36EEF169FFAFFF31FBE37D404C26" blockId="26.[151,783,1005,1064]" box="[151,344,1039,1064]" pageId="26" pageNumber="477">
(
<figureCitation id="13792A6BF169FFAFFF39FBE37CE44C26" box="[159,252,1039,1064]" captionStart="FIGURES 45–49" captionStartId="12.[151,264,1735,1758]" captionTargetBox="[201,1372,376,1683]" captionTargetId="figure@12.[150,1436,373,1734]" captionTargetPageId="12" captionText="FIGURES 45–49. Habitus of Anthobium fusculum group: 45—A. confucii (paratype; CHINA: Hubei), 46—A. cuccodoroi (paratype), 47—A. denticulatum (paratype), 48—A. laozii (paratype; CHINA: Yunnan, Cangshan), 49—A. latissimum (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607326/files/figure.png" pageId="26" pageNumber="477">Figs. 47</figureCitation>
,
<figureCitation id="13792A6BF169FFAFFEAEFBE37D484C26" box="[264,336,1039,1064]" captionStart-0="FIGURE 54" captionStart-1="FIGURES 55–58" captionStartId-0="15.[151,250,982,1004]" captionStartId-1="17.[151,264,1972,1995]" captionTargetBox-0="[179,1408,193,961]" captionTargetBox-1="[374,1237,209,1940]" captionTargetId-0="figure@15.[179,1408,193,961]" captionTargetId-1="figure@17.[268,1328,193,1959]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionText-0="FIGURE 54. Distribution of Anthobium altivagans (black circles), A. besucheti (black squares), A. rugosum (black triangles) and A. denticulatum (black star) in the Himalayan Region." captionText-1="FIGURES 55–58. Aedeagus of Anthobium denticulatum (55–56: paratype) and A. capitale (57–58: holotype): 55, 57—parameral view, 56, 58—lateral view. Scale bar: 0.1 mm." httpUri-0="https://zenodo.org/record/2607330/files/figure.png" httpUri-1="https://zenodo.org/record/2607334/files/figure.png" pageId="26" pageNumber="477">54–56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586565F169FFAFFF31FBBB7D9F4D46" pageId="26" pageNumber="477" type="materials_examined">
<paragraph id="8BFD36EEF169FFAFFF31FBBB7E074CD2" blockId="26.[151,1437,1111,2037]" pageId="26" pageNumber="477">
<emphasis id="B936EAFCF169FFAFFF31FBBB7DD24C7E" bold="true" box="[151,458,1111,1136]" pageId="26" pageNumber="477">Type material examined:</emphasis>
<materialsCitation id="3B2A3CB3F169FFAFFE70FBBB7D2C4CB6" ID-GBIF-Occurrence="2028139715" collectingDate="2013-09-13" collectorName="Kaski, Wl. &amp; Hagge &amp; Schmidt'" country="Nepal" latitude="28.380001" location="Madi Khola valley" municipality="Sikles" pageId="26" pageNumber="477" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F9884CF169FFAFFE70FBBB7E264C7E" box="[470,574,1111,1136]" pageId="26" pageNumber="477" type="holotype">Holotype</typeStatus>
<specimenCount id="9D44FD67F169FFAFFDECFBB47E7A4C61" box="[586,610,1112,1135]" pageId="26" pageNumber="477" type="male">♂</specimenCount>
: ‘
<collectingCountry id="F355767EF169FFAFFDD0FBB47ECE4C7E" box="[630,726,1112,1136]" name="Nepal" pageId="26" pageNumber="477">NEPAL</collectingCountry>
, D:
<collectorName id="26B75338F169FFAFFCABFBBB7F934C7E" box="[781,907,1111,1136]" pageId="26" pageNumber="477">Kaski, Wl.</collectorName>
|
<location id="8E9D6035F169FFAFFC01FBBB789A4C7E" LSID="urn:lsid:plazi:treatment:03EB87F8F169FFAEFF31FC017EE44B8E:8E9D6035F169FFAFFC01FBBB789A4C7E" box="[935,1154,1111,1136]" country="Nepal" latitude="28.380001" municipality="Sikles" name="Madi Khola valley" pageId="26" pageNumber="477">Madi Khola valley</location>
| above
<collectingMunicipality id="6B99AC94F169FFAFFB48FBBB792D4C7E" box="[1262,1333,1111,1136]" pageId="26" pageNumber="477">Sikles</collectingMunicipality>
, 3200m’ &lt;rectangular label, printed&gt;, ‘
<geoCoordinate id="EE765029F169FFAFFE47FB907E724C9A" box="[481,618,1148,1172]" direction="north" orientation="latitude" pageId="26" pageNumber="477" precision="15" value="28.380001">28°22'48''N</geoCoordinate>
, 84°03' | 54''E,
<date id="FFFC102EF169FFAFFCB3FB907F804C9A" box="[789,920,1148,1172]" pageId="26" pageNumber="477" value="2013-09-13">
<collectingDate id="EFB8E9C6F169FFAFFCB3FB907F804C9A" box="[789,920,1148,1172]" pageId="26" pageNumber="477" value="2013-09-13">13.IX.2013</collectingDate>
</date>
, leg. | [J.]
<collectorName id="26B75338F169FFAFFBACFB91784B4C9B" box="[1034,1107,1149,1173]" pageId="26" pageNumber="477">Hagge</collectorName>
&amp; [J.]
<collectorName id="26B75338F169FFAFFB07FB9079104C9B" box="[1185,1288,1148,1173]" pageId="26" pageNumber="477">Schmidt’</collectorName>
&lt;rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="3B2A3CB3F169FFAFFEF1FB4C7E034CD2" ID-GBIF-Occurrence="2028139733" collectionCode="NME" collectorName="Shavrin A. &amp; Smetana A." country="Nepal" pageId="26" pageNumber="477" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F9884CF169FFAFFEF1FB4C7DE84CB6" box="[343,496,1184,1208]" pageId="26" pageNumber="477" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C424D6DF169FFAFFDA8FB737E964CB6" authorityName="Leach" authorityYear="1819" box="[526,654,1183,1208]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="505" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| denticulatum
<emphasis id="B936EAFCF169FFAFFCEDFB737F9A4CB6" bold="true" box="[843,898,1183,1208]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="A2055787F169FFAFFCEDFB737F9A4CB6" box="[843,898,1183,1208]" pageId="26" pageNumber="477" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="26B75338F169FFAFFC07FB7378384CB6" box="[929,1056,1183,1208]" pageId="26" pageNumber="477">Shavrin A.</collectorName>
&amp;
<collectorName id="26B75338F169FFAFFBE8FB4C78CE4CB6" box="[1102,1238,1184,1208]" pageId="26" pageNumber="477">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="ED53AE2BF169FFAFFE68FB287E0B4CD2" box="[462,531,1220,1244]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="26" pageNumber="477">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD36EEF169FFAFFF61FB047D9F4D46" blockId="26.[151,1437,1111,2037]" pageId="26" pageNumber="477">
<typeStatus id="54F9884CF169FFAFFF61FB047D2C4D0E" box="[199,308,1256,1280]" pageId="26" pageNumber="477" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D44FD67F169FFAFFEE2FB047DFA4D0E" box="[324,482,1255,1280]" pageId="26" pageNumber="477" type="generic">19 specimens</specimenCount>
):
<specimenCount id="9D44FD67F169FFAFFE52FB0B7E2B4CF1" box="[500,563,1255,1280]" pageId="26" pageNumber="477" type="male">6 ♂♂</specimenCount>
[three specimens dissected],
<materialsCitation id="3B2A3CB3F169FFAFFCDBFB0478E54D0E" ID-GBIF-Occurrence="2028139727" box="[893,1277,1255,1280]" collectionCode="NME" collectorName="Shavrin A. &amp; Smetana A." country="Nepal" pageId="26" pageNumber="477" specimenCount="13" specimenCount-female="13" typeStatus="paratype">
<specimenCount id="9D44FD67F169FFAFFCDBFB047FD54CF1" box="[893,973,1256,1280]" pageId="26" pageNumber="477" type="female">13 ♀♀</specimenCount>
: same data as the holotype
</materialsCitation>
, ‘
<materialsCitation id="3B2A3CB3F169FFAFFAAAFB047D9C4D46" ID-GBIF-Occurrence="2028139711" collectionCode="CS" country="Nepal" pageId="26" pageNumber="477" specimenCount="19" specimenCount-female="13" specimenCount-male="6" typeStatus="paratype">
<typeStatus id="54F9884CF169FFAFFAAAFB0479844D0E" box="[1292,1436,1256,1280]" pageId="26" pageNumber="477" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C424D6DF169FFAFFF03FAE07D3D4D2B" authorityName="Leach" authorityYear="1819" box="[165,293,1292,1317]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="505" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| denticulatum
<emphasis id="B936EAFCF169FFAFFE7EFAE07E084D2B" bold="true" box="[472,528,1292,1317]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="A2055787F169FFAFFE7EFAE07E084D2B" box="[472,528,1292,1317]" pageId="26" pageNumber="477" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<bibRefCitation id="EFD34B1FF169FFAFFD8EFAE07F8E4D2A" author="Shavrin, A. V. &amp; Smetana, A." box="[552,918,1292,1317]" pageId="26" pageNumber="477" pagination="1 - 39" refId="ref40158" refString="Shavrin, A. V. &amp; Smetana, A. (2017) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups. Zootaxa, 4365 (1), 1 - 39. https: // doi. org / 10.11646 / zootaxa. 4365.1.1" type="journal article" year="2017">Shavrin A. &amp; Smetana A. 2017</bibRefCitation>
’ &lt;red rectangular label, printed&gt; (
<specimenCount id="9D44FD67F169FFAFFAB9FAE079554D2A" box="[1311,1357,1292,1317]" pageId="26" pageNumber="477" type="male">5 ♂</specimenCount>
,
<specimenCount id="9D44FD67F169FFAFFAFDFAE0798E4D2A" box="[1371,1430,1292,1316]" pageId="26" pageNumber="477" type="female">11 ♀</specimenCount>
: NME;
<specimenCount id="9D44FD67F169FFAFFF42FADC7D084D49" box="[228,272,1328,1352]" pageId="26" pageNumber="477" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D44FD67F169FFAFFEBAFADC7D504D49" box="[284,328,1328,1352]" pageId="26" pageNumber="477" type="female">2 ♀</specimenCount>
:
<collectionCode id="ED53AE2BF169FFAFFEF3FADC7D634D46" box="[341,379,1328,1352]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="26" pageNumber="477">CS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586565F169FFAEFF61FAB87D614AA9" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477" type="description">
<paragraph id="8BFD36EEF169FFAFFF61FAB87F604DBB" blockId="26.[151,1437,1111,2037]" pageId="26" pageNumber="477">
<emphasis id="B936EAFCF169FFAFFF61FAB87D494D63" bold="true" box="[199,337,1364,1389]" pageId="26" pageNumber="477">Description</emphasis>
. Measurements (n=20): HW: 0.55–0.58; HL: 0.33–0.40; AL(
<typeStatus id="54F9884CF169FFAFFBA3FAB878744D63" box="[1029,1132,1364,1389]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
): 0.85; OL: 0.12–0.14; PL: 0.42–0.49; PW: 0.78–0.87; ESL: 0.85–0.93; EW: 0.96–1.12; AW: 0.86–0.97; MTbL: 0.45, MTrL: 0.20 (MTrL 1–4: 0.13; MTrL 5: 0.07); AedL: 1.10; TL: 2.30–3.30 (2.70–
<typeStatus id="54F9884CF169FFAFFCA1FA707F754DBB" box="[775,877,1436,1461]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BFD36EEF169FFAFFF61FA2C7D9F4F4E" blockId="26.[151,1437,1111,2037]" pageId="26" pageNumber="477">
In external appearance similar to
<taxonomicName id="4C424D6DF169FFAFFDE7FA2C7ED34DD6" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[577,715,1472,1496]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B936EAFCF169FFAFFDE7FA2C7ED34DD6" box="[577,715,1472,1496]" italics="true" pageId="26" pageNumber="477">A. besucheti</emphasis>
</taxonomicName>
. Body and antennomeres 9–11 yellow-brown to reddish brown, with darker head and abdomen in some specimens; mouthparts, antennomeres and legs yellow to yellow-brown. Head with distinct microsculpture, transverse in middle and between ocelli, isodiametric on anterior portions of infraorbital ridges, posterior portions of infraorbital ridges with slightly rugose vertical waves of sculpture; anterior part of middle elevation of pronotum with indistinct transverse meshes in some specimens; scutellum with fine isodiametric microsculpture. Head with irregular small punctation, denser and coarser on posteriolateral portions; pronotum with irregular, large and deep punctation, sparser on lateral portions, with small impunctate area between mediobasal impression in some specimens; apex of scutellum with several small punctures; punctation of elytra as that on pronotum, some specimens with distinctly smaller punctures on parascutellar area and along suture, each elytron with five to eight vague and tangled longitudinal, and with diagonal rows of punctures in some specimens. Habitus as in
<figureCitation id="13792A6BF169FFAFFE96F8CB7D9A4F4E" box="[304,386,1831,1856]" captionStart="FIGURES 45–49" captionStartId="12.[151,264,1735,1758]" captionTargetBox="[201,1372,376,1683]" captionTargetId="figure@12.[150,1436,373,1734]" captionTargetPageId="12" captionText="FIGURES 45–49. Habitus of Anthobium fusculum group: 45—A. confucii (paratype; CHINA: Hubei), 46—A. cuccodoroi (paratype), 47—A. denticulatum (paratype), 48—A. laozii (paratype; CHINA: Yunnan, Cangshan), 49—A. latissimum (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607326/files/figure.png" pageId="26" pageNumber="477">Fig. 47</figureCitation>
.
</paragraph>
<paragraph id="8BFD36EEF169FFAFFF61F8A07D064FFB" blockId="26.[151,1437,1111,2037]" pageId="26" pageNumber="477">
Head 1.4–1.6 times as wide as long; middle portion of head slightly and irregularly elevated, with indistinct to distinct transverse impression between eyes at posterior third, with narrow grooves in front of ocelli reaching posterior third or middle length of eyes; postocular ridges indistinct, very smooth. Length × width of antennomeres (
<typeStatus id="54F9884CF169FFAFFF39F8547D104FDF" box="[159,264,1976,2001]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
): 1: 0.12 × 0.06; 2: 0.07 × 0.04; 3: 0.08 × 0.03; 4–7: 0.07 × 0.04; 8: 0.06 × 0.04; 9–10: 0.06 × 0.05; 11: 0.12 × 0.05.
</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FF747F9048F6" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">Pronotum 1.7–1.8 times as wide as long, 1.4–1.5 times as wide as head; lateral edges of pronotum with regular, small and smooth crenulation; pronotum usually with deep longitudinal impression in apical part, and with very deep transverse impression in mediobasal third in some specimens.</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FEE87EC64931" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">Elytra 1.1–1.2 times as wide as long, twice as long as pronotum; middle part of each elytron usually with irregular longitudinal elevations between punctures.</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FEA07EE749DE" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">
Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="13792A6BF168FFAEFAA4FEA0794F496B" box="[1282,1367,332,357]" captionStart="FIGURES 55–58" captionStartId="17.[151,264,1972,1995]" captionTargetBox="[374,1237,209,1940]" captionTargetId="figure@17.[268,1328,193,1959]" captionTargetPageId="17" captionText="FIGURES 55–58. Aedeagus of Anthobium denticulatum (55–56: paratype) and A. capitale (57–58: holotype): 55, 57—parameral view, 56, 58—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607334/files/figure.png" pageId="27" pageNumber="478">Fig. 55</figureCitation>
) with moderately wide median lobe with subtruncate apex; parameres significantly exceeding apex of median lobe, with two apical and two preapical setae; internal sac narrow and long, with rows of sclerotized widened teeth in both middle and basal part. Aedeagus laterally as in
<figureCitation id="13792A6BF168FFAEFD0EFE5B7EE349DE" box="[680,763,439,464]" captionStart="FIGURES 55–58" captionStartId="17.[151,264,1972,1995]" captionTargetBox="[374,1237,209,1940]" captionTargetId="figure@17.[268,1328,193,1959]" captionTargetPageId="17" captionText="FIGURES 55–58. Aedeagus of Anthobium denticulatum (55–56: paratype) and A. capitale (57–58: holotype): 55, 57—parameral view, 56, 58—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607334/files/figure.png" pageId="27" pageNumber="478">Fig. 56</figureCitation>
.
</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FE307CE24A16" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">Female. Apical margin of abdominal tergite VIII slightly sinuate. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FDC87D614AA9" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">
<emphasis id="B936EAFCF168FFAEFF61FDC87DAB4A33" bold="true" box="[199,435,548,573]" pageId="27" pageNumber="478">Comparative notes.</emphasis>
Based on the body size, coloration, shape of the body with moderately short elytra,
<taxonomicName id="4C424D6DF168FFAEFA22FDC87E0D4A6E" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B936EAFCF168FFAEFA22FDC87D314A6E" italics="true" pageId="27" pageNumber="478">A. denticulatum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<emphasis id="B936EAFCF168FFAEFD84FDAB7E414A6E" bold="true" box="[546,601,583,608]" pageId="27" pageNumber="478">
<taxonomicNameLabel id="A2055787F168FFAEFD84FDAB7E414A6E" box="[546,601,583,608]" pageId="27" pageNumber="478" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C424D6DF168FFAEFC92FDA478B14A6E" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[820,1193,583,608]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="B936EAFCF168FFAEFC92FDA47FA64A6E" box="[820,958,584,608]" italics="true" pageId="27" pageNumber="478">A. besucheti</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<emphasis id="B936EAFCF168FFAEFB11FDAB78F74A6E" bold="true" box="[1207,1263,583,608]" pageId="27" pageNumber="478">
<taxonomicNameLabel id="A2055787F168FFAEFB11FDAB78F74A6E" box="[1207,1263,583,608]" pageId="27" pageNumber="478" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the slightly narrower pronotum, by wider and larger aedeagus, and by the presence of rows of sclerotized teeth in internal sac.
</paragraph>
</subSubSection>
<subSubSection id="C3586565F168FFAEFF61FD587D8D4B36" pageId="27" pageNumber="478" type="distribution">
<paragraph id="8BFD36EEF168FFAEFF61FD587D9D4AFE" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">
<emphasis id="B936EAFCF168FFAEFF61FD587D784AC3" bold="true" box="[199,352,692,717]" pageId="27" pageNumber="478">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="54F9884CF168FFAEFC64FD597FEA4AC3" box="[962,1010,693,717]" pageId="27" pageNumber="478">type</typeStatus>
locality (
<figureCitation id="13792A6BF168FFAEFBC4FD5878AE4AC3" box="[1122,1206,692,717]" captionStart="FIGURE 54" captionStartId="15.[151,250,982,1004]" captionTargetBox="[179,1408,193,961]" captionTargetId="figure@15.[179,1408,193,961]" captionTargetPageId="15" captionText="FIGURE 54. Distribution of Anthobium altivagans (black circles), A. besucheti (black squares), A. rugosum (black triangles) and A. denticulatum (black star) in the Himalayan Region." httpUri="https://zenodo.org/record/2607330/files/figure.png" pageId="27" pageNumber="478">Fig. 54</figureCitation>
) in the Madi Khola valley, central
<collectingCountry id="F355767EF168FFAEFE9DFD3B7D9A4AFE" box="[315,386,727,752]" name="Nepal" pageId="27" pageNumber="478">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8BFD36EEF168FFAEFF61FD107D8D4B36" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">
<emphasis id="B936EAFCF168FFAEFF61FD107D524B1B" bold="true" box="[199,330,764,789]" pageId="27" pageNumber="478">Bionomics.</emphasis>
Specimens were taken by sifting leaf litter in
<taxonomicName id="4C424D6DF168FFAEFCC6FD10781D4B1A" box="[864,1029,764,788]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="478" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFCF168FFAEFCC6FD10781D4B1A" box="[864,1029,764,788]" italics="true" pageId="27" pageNumber="478">Rhododendron</emphasis>
</taxonomicName>
forest at elevation
<quantity id="4CBA9B0BF168FFAEFB42FD1079234B1A" box="[1252,1339,764,788]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="478" unit="m" value="3200.0">3200 m</quantity>
a.s.l. (J. Schmidt, pers. comm.)
</paragraph>
</subSubSection>
<subSubSection id="C3586565F168FFAEFF61FCA87EE44B8E" pageId="27" pageNumber="478" type="etymology">
<paragraph id="8BFD36EEF168FFAEFF61FCA87EE44B8E" blockId="27.[151,1437,151,896]" pageId="27" pageNumber="478">
<emphasis id="B936EAFCF168FFAEFF61FCA87D564B53" bold="true" box="[199,334,836,861]" pageId="27" pageNumber="478">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="B936EAFCF168FFAEFC97FCA87FA24B52" box="[817,954,836,860]" italics="true" pageId="27" pageNumber="478">denticulatus</emphasis>
, -
<emphasis id="B936EAFCF168FFAEFC75FCA97FF94B52" box="[979,993,837,860]" italics="true" pageId="27" pageNumber="478">a</emphasis>
, -
<emphasis id="B936EAFCF168FFAEFC5FFCA978034B52" box="[1017,1051,837,860]" italics="true" pageId="27" pageNumber="478">um</emphasis>
(toothed). It refers to lateral rows of sclerotized teeth on the internal sac of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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