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<document id="92BDC0ED50A335CBD1763ACEBF58B718" ID-DOI="10.5281/zenodo.5353945" ID-ISSN="2345-7600" ID-Zenodo-Dep="5353945" ID-ZooBank="4CF42744-861A-4635-9703-E6639CEBFAA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212991585" checkinUser="tatiana" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2014" docId="03EB87DA3B18E370FC70FBB4FD7398B3" docLanguage="en" docName="62rbz396-483.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Mariaplax cyrtophallus Rahayu &amp; Ng 2014, new species" docType="treatment" docUuid="84AD3B94-00E0-4552-9153-2AFEC186AE16" docUuidSource="ZooBank" docVersion="4" lastPageNumber="434" masterDocId="FFD2FFA23B3BE356FFC6FFC9FF839D46" masterDocTitle="New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic" masterLastPageNumber="483" masterPageNumber="396" pageNumber="431" updateTime="1699188253459" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F0EF940BFFCAFB740B019BE46985036B">New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic</mods:title>
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<mods:name id="3F321DA2E8CF409D00851879A3EF9D70" type="personal">
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<mods:namePart id="2AB6093A75923302423F81C0433A2879">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="47385DE83A43658850E6D045E88E95C3">Ng, Peter K. L.</mods:namePart>
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<mods:title id="D9FD98E682ABBCC4FB9A3EEE1263536F">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="AB31CD59B4F5040E02569270D2DF6121">2014</mods:date>
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<mods:number id="380E466B3ADBA7AB736C92D7BA56C06C">2014-07-04</mods:number>
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<treatment id="03EB87DA3B18E370FC70FBB4FD7398B3" ID-DOI="http://doi.org/10.5281/zenodo.5451331" ID-Zenodo-Dep="5451331" LSID="urn:lsid:zoobank.org:act:84AD3B94-00E0-4552-9153-2AFEC186AE16" httpUri="http://treatment.plazi.org/id/03EB87DA3B18E370FC70FBB4FD7398B3" lastPageId="38" lastPageNumber="434" pageId="35" pageNumber="431">
<subSubSection id="C35865473B18E375FC70FBB4FAD399D3" box="[950,1360,1149,1173]" pageId="35" pageNumber="431" type="nomenclature">
<paragraph id="8BFD36CC3B18E375FC70FBB4FAD399D3" blockId="35.[950,1360,1149,1205]" box="[950,1360,1149,1173]" pageId="35" pageNumber="431">
<heading id="D0B581A03B18E375FC70FBB4FAD399D3" bold="true" box="[950,1360,1149,1173]" centered="true" fontSize="10" level="2" pageId="35" pageNumber="431" reason="2">
<emphasis id="B936EADE3B18E375FC70FBB4FAD399D3" bold="true" box="[950,1360,1149,1173]" pageId="35" pageNumber="431">
<taxonomicName id="4C424D4F3B18E375FC70FBB4FB3999D3" ID-CoL="3Y68V" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[950,1210,1149,1173]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="431" phylum="Arthropoda" rank="species" species="cyrtophallus" status="sp. nov.">
<emphasis id="B936EADE3B18E375FC70FBB4FB3999D3" bold="true" box="[950,1210,1149,1173]" italics="true" pageId="35" pageNumber="431">Mariaplax cyrtophallus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20557A53B18E375FB0CFBB4FAD399D3" box="[1226,1360,1149,1173]" pageId="35" pageNumber="431" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C35865473B18E375FBFFFB54FB4E99F3" box="[1081,1229,1181,1205]" pageId="35" pageNumber="431" type="description">
<paragraph id="8BFD36CC3B18E375FBFFFB54FB4E99F3" blockId="35.[950,1360,1149,1205]" box="[1081,1229,1181,1205]" pageId="35" pageNumber="431">
(
<figureCitation id="13792A493B18E375FB87FB54FB1F99F3" box="[1089,1180,1181,1205]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="35" pageNumber="431">Figs. 28</figureCitation>
,
<figureCitation id="13792A493B18E375FB6FFB54FB4699F3" box="[1193,1221,1181,1205]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35865473B18E375FC85FB15FB0E986D" pageId="35" pageNumber="431" type="materials_examined">
<paragraph id="8BFD36CC3B18E375FC85FB15FB0E986D" blockId="35.[835,1475,1244,1324]" pageId="35" pageNumber="431">
<emphasis id="B936EADE3B18E375FC85FB15FB8E99B4" bold="true" box="[835,1037,1244,1266]" pageId="35" pageNumber="431">Material examined.</emphasis>
<materialsCitation id="3B2A3C913B18E375FBD0FB15FB0E986D" collectingDate="2000-11-07" collectionCode="MZB" collectorName="A. Haris" country="Indonesia" latitude="-5.0255556" location="Otakwa" longLatPrecision="15" longitude="137.18242" municipality="Timika" pageId="35" pageNumber="431" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="54F9886E3B18E375FBD0FB15FBF099B4" box="[1046,1139,1244,1266]" pageId="35" pageNumber="431" type="holotype">Holotype</typeStatus>
: male (6.5 ×
<quantity id="4CBA9B293B18E375FB3FFB15FAC599B7" box="[1273,1350,1244,1265]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="35" pageNumber="431" unit="mm" value="4.2">4.2 mm</quantity>
) (
<collectionCode id="ED53AE093B18E375FA9DFB15FA1099B4" box="[1371,1427,1244,1266]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="431" type="Museum">MZB</collectionCode>
), off
<location id="8E9D60173B18E375FC85FB30FC139849" LSID="urn:lsid:plazi:treatment:03EB87DA3B18E370FC70FBB4FD7398B3:8E9D60173B18E375FC85FB30FC139849" box="[835,912,1273,1295]" country="Indonesia" latitude="-5.0255556" longLatPrecision="15" longitude="137.18242" municipality="Timika" name="Otakwa" pageId="35" pageNumber="431" stateProvince="Papua Barat">Otakwa</location>
,
<collectingMunicipality id="6B99ACB63B18E375FC5EFB30FC5C9849" box="[920,991,1273,1295]" pageId="35" pageNumber="431">Timika</collectingMunicipality>
,
<collectingRegion id="4986F82E3B18E375FC21FB30FBA79848" box="[999,1060,1273,1294]" country="Indonesia" name="Papua Barat" pageId="35" pageNumber="431">Papua</collectingRegion>
,
<collectingCountry id="F355765C3B18E375FBEBFB30FB0E9849" box="[1069,1165,1273,1295]" name="Indonesia" pageId="35" pageNumber="431">Indonesia</collectingCountry>
, near
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<geoCoordinate id="EE76500B3B18E375FAF4FB30FA3D9849" box="[1330,1470,1273,1295]" degrees="137" direction="east" minutes="10" orientation="longitude" pageId="35" pageNumber="431" precision="1" seconds="56.7" value="137.18242">137º10'56.7&quot;E</geoCoordinate>
, coll.
<collectorName id="26B7531A3B18E375FCB2FADFFC4A986A" box="[884,969,1302,1324]" pageId="35" pageNumber="431">A. Haris</collectorName>
,
<date id="FFFC100C3B18E375FC13FADFFB0A986D" box="[981,1161,1302,1324]" pageId="35" pageNumber="431" value="2000-11-07">
<collectingDate id="EFB8E9E43B18E375FC13FADFFB0A986D" box="[981,1161,1302,1324]" pageId="35" pageNumber="431" value="2000-11-07">7 November 2000</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35865473B18E372FC85FA9AFA919CB0" lastPageId="36" lastPageNumber="432" pageId="35" pageNumber="431" type="diagnosis">
<paragraph id="8BFD36CC3B18E372FC85FA9AFB149CF0" blockId="35.[835,1475,1363,2092]" lastBlockId="36.[812,1452,190,439]" lastPageId="36" lastPageNumber="432" pageId="35" pageNumber="431">
<emphasis id="B936EADE3B18E375FC85FA9AFC3B982D" bold="true" box="[835,952,1363,1387]" pageId="35" pageNumber="431">Diagnosis.</emphasis>
Carapace subquadrate, about 1.5 times as broad as long, dorsal surface strongly granulated; regions indistinct, with H-shaped shallow depression medially (
<figureCitation id="13792A493B18E375FAD2FA5DFAFE98EA" box="[1300,1405,1428,1452]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="35" pageNumber="431">Figs. 28A</figureCitation>
,
<figureCitation id="13792A493B18E375FA40FA5DFA3498EA" box="[1414,1463,1428,1452]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">29A</figureCitation>
). Eye small, slightly movable, cornea black, wider than granular peduncle. Pterygostomial region (
<figureCitation id="13792A493B18E375FB65FA1DFA8898AA" box="[1187,1291,1492,1516]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="35" pageNumber="431">Figs. 28B</figureCitation>
,
<figureCitation id="13792A493B18E375FAD2FA1DFAC098AA" box="[1300,1347,1492,1516]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">29D</figureCitation>
) covered by granules, with row of 5 oblique striae. Third maxillipeds broad (
<figureCitation id="13792A493B18E375FC8DF9DDFC379B6A" box="[843,948,1556,1580]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="35" pageNumber="431">Figs. 28B</figureCitation>
,
<figureCitation id="13792A493B18E375FC7BF9DDFC689B6A" box="[957,1003,1556,1580]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">29E</figureCitation>
), completely covering buccal cavity; ischium slightly longer than merus, slightly expanded subproximally, with oblique mesial margin, combined length of dactylus, propodus and carpus almost as long as that of merus and ischium; exopod relatively narrow, about 0.4 width of ischium, flagellum well developed. Chelipeds asymmetrical, covered by closely spaced tubercles (
<figureCitation id="13792A493B18E375FB24F91DFAEE9BAA" box="[1250,1389,1748,1772]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29G, H</figureCitation>
); major chela (
<figureCitation id="13792A493B18E375FC48F93DFC719A4A" box="[910,1010,1780,1804]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29G</figureCitation>
) with slightly gaping finger when closed; dactylus and fixed finger with longitudinal ridge at midline distally; palm slightly shorter than broad; minor chela (
<figureCitation id="13792A493B18E375FA51F8FDFCF09A2A" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29H</figureCitation>
) with narrow gap when fingers closed; dactylus and fixed finger with median longitudinal ridge distally. P2P4 (
<figureCitation id="13792A493B18E375FC8DF85DFC239AEA" box="[843,928,1940,1964]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29I</figureCitation>
) relatively long, slender, covered by dense granules; P3 longest (
<figureCitation id="13792A493B18E375FC0DF87DFBAA9A8A" box="[971,1065,1972,1996]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29I</figureCitation>
); merus of P4 relatively long, about 3.4 times as long as broad; dactylus of P4 gently upcurved (
<figureCitation id="13792A493B18E375FC8DF83DFC26954A" box="[843,933,2036,2060]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29J</figureCitation>
); merus without discernible distinct median groove on lateral surface (
<figureCitation id="13792A493B18E375FBD5F7DDFBF3956A" box="[1043,1136,2068,2092]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="35" pageNumber="431">Fig. 29J</figureCitation>
). Male thoracic sternum broad (
<figureCitation id="13792A493B1FE372FFBCFF77FE859D90" box="[122,262,190,214]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">Fig. 29B, C</figureCitation>
), surface prominently granular; sternites 1 and 2 fused, separated from sternite 3 by distinct furrow; sternite 3 separated from sternite 4 laterally, medially appear fused, thoracic sternal groove deep, transverse, extending laterally from edge of sternoabdominal cavity, parallel to frontal margin, not reaching base of chelipeds (
<figureCitation id="13792A493B1FE372FDBBFE97FD689C30" box="[637,747,350,374]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="36" pageNumber="432">Figs. 28B</figureCitation>
,
<figureCitation id="13792A493B1FE372FFB7FEB7FF209CD0" box="[113,163,382,406]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">29C</figureCitation>
); sternite 8 exposed, quadrangular, as long as male abdominal somite 1; sternoabdominal cavity not reaching middle to sternite 4 (
<figureCitation id="13792A493B1FE372FE99FE77FE469C90" box="[351,453,446,470]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">Fig. 29C</figureCitation>
). Male abdomen relatively narrow (
<figureCitation id="13792A493B1FE372FF08FE17FEB39CB0" box="[206,304,478,502]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">Fig. 29F</figureCitation>
), surface granular; somites 1 and 2 free, somites 35 fused, lateral margin slightly arched; somite 6 shorter than length of somites 35, median part of lateral margin expanded into blunt triangular projection; telson subpentagonal (
<figureCitation id="13792A493B1FE372FC12FED7FBBC9C70" box="[980,1087,286,310]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="36" pageNumber="432">Figs. 28B</figureCitation>
,
<figureCitation id="13792A493B1FE372FB8CFED7FBF59C70" box="[1098,1142,286,310]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">29F</figureCitation>
), slightly shorter than somite 6, distal margin rounded, sparsely setose. G1 (
<figureCitation id="13792A493B1FE372FAE1FEF7FA279C10" box="[1319,1444,318,342]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="36" pageNumber="432">Fig. 29J, K</figureCitation>
) bent 90° at midlength, directed anterolaterally, distal part tapered, not concealed under abdomen, exposed on thoracic sternal groove. Female unknown.
</paragraph>
<paragraph id="8BFD36CC3B1FE372FCEAFE17FA919CB0" blockId="36.[812,1298,478,502]" box="[812,1298,478,502]" pageId="36" pageNumber="432">
<emphasis id="B936EADE3B1FE372FCEAFE17FC039CB0" bold="true" box="[812,896,478,502]" pageId="36" pageNumber="432">Colour.</emphasis>
Dirty white when freshly preserved.
</paragraph>
</subSubSection>
<caption id="DF3D66443B1FE372FFB7F7E5FA2E9504" ID-DOI="http://doi.org/10.5281/zenodo.5354005" ID-Zenodo-Dep="5354005" box="[113,1453,2092,2114]" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="36" pageNumber="432" startId="36.[113,148,2092,2114]" targetBox="[113,1451,572,2059]" targetPageId="36">
<paragraph id="8BFD36CC3B1FE372FFB7F7E5FA2E9504" blockId="36.[113,1453,2092,2114]" box="[113,1453,2092,2114]" pageId="36" pageNumber="432">
Fig. 28.
<taxonomicName id="4C424D4F3B1FE372FF78F7E5FE1E9507" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[190,413,2092,2113]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="432" phylum="Arthropoda" rank="species" species="cyrtophallus" status="sp. nov.">
<emphasis id="B936EADE3B1FE372FF78F7E5FE1E9507" box="[190,413,2092,2113]" italics="true" pageId="36" pageNumber="432">Mariaplax cyrtophallus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20557A53B1FE372FE61F7E5FD989504" box="[423,539,2092,2114]" pageId="36" pageNumber="432" rank="species">new species</taxonomicNameLabel>
, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax.
</paragraph>
</caption>
<caption id="DF3D66443B1EE373FF4EF81BFC119578" ID-DOI="http://doi.org/10.5281/zenodo.5354007" ID-Zenodo-Dep="5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="37" pageNumber="433" startId="37.[136,171,2002,2024]" targetBox="[158,1434,194,1961]" targetPageId="37">
<paragraph id="8BFD36CC3B1EE373FF4EF81BFC119578" blockId="37.[136,1474,2002,2110]" pageId="37" pageNumber="433">
Fig. 29.
<taxonomicName id="4C424D4F3B1EE373FF18F81BFE469AA1" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[222,453,2002,2023]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="433" phylum="Arthropoda" rank="species" species="cyrtophallus" status="sp. nov.">
<emphasis id="B936EADE3B1EE373FF18F81BFE469AA1" box="[222,453,2002,2023]" italics="true" pageId="37" pageNumber="433">Mariaplax cyrtophallus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20557A53B1EE373FE12F81BFDCD9AAE" box="[468,590,2002,2024]" pageId="37" pageNumber="433" rank="species">new species</taxonomicNameLabel>
, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C35865473B1DE370FFB7FF74FE489C53" pageId="38" pageNumber="434" type="etymology">
<paragraph id="8BFD36CC3B1DE370FFB7FF74FE489C53" blockId="38.[113,752,189,277]" pageId="38" pageNumber="434">
<emphasis id="B936EADE3B1DE370FFB7FF74FF709D93" bold="true" box="[113,243,189,213]" pageId="38" pageNumber="434">Etymology.</emphasis>
From the Latin
<emphasis id="B936EADE3B1DE370FE76FF77FE649D93" box="[432,487,190,213]" italics="true" pageId="38" pageNumber="434">cyrto</emphasis>
, bend, and
<emphasis id="B936EADE3B1DE370FDA9FF74FD3D9D93" box="[623,702,189,213]" italics="true" pageId="38" pageNumber="434">phallus</emphasis>
, for penis; alluding to the bent structure of the G1. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C35865473B1DE370FFB7FEF4FE5098F3" pageId="38" pageNumber="434" type="discussion">
<paragraph id="8BFD36CC3B1DE370FFB7FEF4FE739950" blockId="38.[113,752,317,1046]" pageId="38" pageNumber="434">
<emphasis id="B936EADE3B1DE370FFB7FEF4FF5C9C13" bold="true" box="[113,223,317,341]" pageId="38" pageNumber="434">Remarks.</emphasis>
Several features of the present new species suggest its affinity to
<taxonomicName id="4C424D4F3B1DE370FEC4FE94FE2B9C33" authorityName=", Alcock" authorityYear="1902" box="[258,424,349,373]" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B1DE370FEC4FE94FE2B9C33" box="[258,424,349,373]" italics="true" pageId="38" pageNumber="434">Lambdophallus</emphasis>
</taxonomicName>
, notably the transverse sternal groove extends a good distance laterally and is parallel to frontal margin, the G1 bent at a 90° angle, and the shape of the third maxilliped that is slightly expanded subproximally and has straight mesial margin. However,
<taxonomicName id="4C424D4F3B1DE370FD8FFE14FD739CB3" authorityName=", Alcock" authorityYear="1902" box="[585,752,477,501]" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B1DE370FD8FFE14FD739CB3" box="[585,752,477,501]" italics="true" pageId="38" pageNumber="434">Lambdophallus</emphasis>
</taxonomicName>
has immovable and very short eyes peduncle, the third maxillipeds was rather narrow and leave wide gap between them, the male thoracic sternal groove starts above the articulation of the sixth abdominal somite and the telson was longer than the sixth somite (Alcock, 1902: pl. 62, fig. 1). These characters indicate that we cannot place this new species in
<taxonomicName id="4C424D4F3B1DE370FF18FD74FE079F93" authorityName=", Alcock" authorityYear="1902" box="[222,388,701,725]" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B1DE370FF18FD74FE079F93" box="[222,388,701,725]" italics="true" pageId="38" pageNumber="434">Lambdophallus</emphasis>
</taxonomicName>
. Although the longitudinal groove on the outer surface of the merus of P2P4 is indiscernible, obscured by tubercles, other characters such as the movable eyes peduncle, the male thoracic sternal groove that starts almost at subdistalmargin of the telson, the telson that was shorter than the sixth abdominal somite, and the oblique mesial margin of ischium of the third maxilliped clearly place this species in
<taxonomicName id="4C424D4F3B1DE370FE9EFC54FE4A9EF3" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[344,457,925,949]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B1DE370FE9EFC54FE4A9EF3" box="[344,457,925,949]" italics="true" pageId="38" pageNumber="434">Mariaplax</emphasis>
</taxonomicName>
. The structure of the male thoracic sternal groove (
<figureCitation id="13792A493B1DE370FEB5FC74FE729E93" box="[371,497,957,981]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="38" pageNumber="434">Fig. 29B, C</figureCitation>
) and shape of the G1 of
<taxonomicName id="4C424D4F3B1DE370FFB7FC17FE9F9EB3" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[113,284,989,1013]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="cyrtophallus" status="sp. nov.">
<emphasis id="B936EADE3B1DE370FFB7FC17FE9F9EB3" box="[113,284,989,1013]" italics="true" pageId="38" pageNumber="434">M. cyrtophallus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20557A53B1DE370FEEDFC17FE2D9EB3" box="[299,430,989,1013]" pageId="38" pageNumber="434" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="13792A493B1DE370FE78FC14FDB29EB3" box="[446,561,989,1013]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 15, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 15, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="38" pageNumber="434">Fig. 29I, J</figureCitation>
), however, easily distinguishes it from all congeners.
</paragraph>
<paragraph id="8BFD36CC3B1DE370FFB7FBF4FE5098F3" blockId="38.[113,752,1085,1461]" pageId="38" pageNumber="434">
Unusual among members of the genus is that
<taxonomicName id="4C424D4F3B1DE370FD8FFBF7FD6C9913" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[585,751,1085,1109]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B1DE370FD8FFBF7FD6C9913" box="[585,751,1085,1109]" italics="true" pageId="38" pageNumber="434">M. cyrtophallus</emphasis>
</taxonomicName>
does not have a discernible median groove on the lateral surface of the ambulatory merus. This is distinct in all other congeners, although it may be shallow at times. In
<taxonomicName id="4C424D4F3B1DE370FD15FB57FF759993" authorityName="Rahayu &amp; Ng" authorityYear="2014" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B1DE370FD15FB57FF759993" italics="true" pageId="38" pageNumber="434">M. cyrtophallus</emphasis>
</taxonomicName>
, the surface of the merus is covered with dense granules and no groove can be seen. In all other aspects, however,
<taxonomicName id="4C424D4F3B1DE370FF18FB37FE0E9853" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[222,397,1277,1301]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B1DE370FF18FB37FE0E9853" box="[222,397,1277,1301]" italics="true" pageId="38" pageNumber="434">M. cyrtophallus</emphasis>
</taxonomicName>
is a typical
<taxonomicName id="4C424D4F3B1DE370FDD8FB34FD739853" box="[542,752,1277,1301]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EADE3B1DE370FDD8FB34FD139853" box="[542,656,1277,1301]" italics="true" pageId="38" pageNumber="434">Mariaplax</emphasis>
species.
</taxonomicName>
It is apparently closest to
<taxonomicName id="4C424D4F3B1DE370FE49FAD7FE739873" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[399,496,1310,1333]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="secus" status="sp. nov.">
<emphasis id="B936EADE3B1DE370FE49FAD7FE739873" box="[399,496,1310,1333]" italics="true" pageId="38" pageNumber="434">M. secus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20557A53B1DE370FE39FAD7FD069873" box="[511,645,1309,1333]" pageId="38" pageNumber="434" rank="species">new species</taxonomicNameLabel>
, from the other side of the island of New
<collectingCountry id="F355765C3B1DE370FE1EFAF4FDAB9813" box="[472,552,1341,1365]" name="Guinea-Bissau" pageId="38" pageNumber="434">Guinea</collectingCountry>
, which also has a similar strongly bent G1; but in
<taxonomicName id="4C424D4F3B1DE370FE01FA97FDA69833" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[455,549,1374,1397]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="434" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B1DE370FE01FA97FDA69833" box="[455,549,1374,1397]" italics="true" pageId="38" pageNumber="434">M. secus</emphasis>
</taxonomicName>
, the lateral surface of the ambulatory merus has a distinct longitudinal groove (see discussion for that species).
</paragraph>
</subSubSection>
<subSubSection id="C35865473B1DE370FFB7FA14FD7398B3" box="[113,752,1501,1525]" pageId="38" pageNumber="434" type="distribution">
<paragraph id="8BFD36CC3B1DE370FFB7FA14FD7398B3" blockId="38.[113,752,1501,1525]" box="[113,752,1501,1525]" pageId="38" pageNumber="434">
<emphasis id="B936EADE3B1DE370FFB7FA14FE8098B3" bold="true" box="[113,259,1501,1525]" pageId="38" pageNumber="434">Distribution.</emphasis>
Timika,
<collectingRegion id="4986F82E3B1DE370FE99FA17FE2098B3" box="[351,419,1502,1525]" country="Indonesia" name="Papua Barat" pageId="38" pageNumber="434">Papua</collectingRegion>
,
<collectingCountry id="F355765C3B1DE370FE6AFA14FD9498B3" box="[428,535,1501,1525]" name="Indonesia" pageId="38" pageNumber="434">Indonesia</collectingCountry>
. Intertidal,
<quantity id="4CBA9B293B1DE370FD4AFA14FD6F98B3" box="[652,748,1501,1525]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="38" pageNumber="434" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">
15
<specimenCount id="9D44FD453B1DE370FD75FA14FD6F98B3" box="[691,748,1501,1525]" pageId="38" pageNumber="434" type="generic">30 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>