treatments-xml/data/03/EB/87/03EB87DA3B0EE36FFC1BFAB7FC459F11.xml

<document id="583B3A7CE13DE4756131921EAB1DD49D" ID-DOI="10.5281/zenodo.5353945" ID-ISSN="2345-7600" ID-Zenodo-Dep="5353945" ID-ZooBank="4CF42744-861A-4635-9703-E6639CEBFAA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212991585" checkinUser="tatiana" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2014" docId="03EB87DA3B0EE36FFC1BFAB7FC459F11" docLanguage="en" docName="62rbz396-483.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Mariaplax secus Rahayu &amp; Ng 2014, new species" docType="treatment" docUuid="A7D68C2F-6245-4C0B-A6B9-87224FACF8A5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="453" masterDocId="FFD2FFA23B3BE356FFC6FFC9FF839D46" masterDocTitle="New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic" masterLastPageNumber="483" masterPageNumber="396" pageNumber="449" updateTime="1699188253459" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4E1F9C876AC2573C5F39A2BB0253FF5B">New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic</mods:title>
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<mods:namePart id="B2ED5292AFBD8D562CD51B6ADCED36EF">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="A3C2A259D72D5C9DE42CB62D93B55B62">Ng, Peter K. L.</mods:namePart>
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<mods:date id="5D8706E513BFC73BB6A0EE6F784D52D1">2014</mods:date>
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<treatment id="03EB87DA3B0EE36FFC1BFAB7FC459F11" ID-DOI="http://doi.org/10.5281/zenodo.5451335" ID-Zenodo-Dep="5451335" LSID="urn:lsid:zoobank.org:act:A7D68C2F-6245-4C0B-A6B9-87224FACF8A5" httpUri="http://treatment.plazi.org/id/03EB87DA3B0EE36FFC1BFAB7FC459F11" lastPageId="57" lastPageNumber="453" pageId="53" pageNumber="449">
<subSubSection id="C35865473B0EE363FC1BFAB7FAAA98D0" box="[989,1321,1406,1430]" pageId="53" pageNumber="449" type="nomenclature">
<paragraph id="8BFD36CC3B0EE363FC1BFAB7FAAA98D0" blockId="53.[989,1321,1406,1462]" box="[989,1321,1406,1430]" pageId="53" pageNumber="449">
<heading id="D0B581A03B0EE363FC1BFAB7FAAA98D0" bold="true" box="[989,1321,1406,1430]" centered="true" fontSize="10" level="2" pageId="53" pageNumber="449" reason="2">
<emphasis id="B936EADE3B0EE363FC1BFAB7FAAA98D0" bold="true" box="[989,1321,1406,1430]" pageId="53" pageNumber="449">
<taxonomicName id="4C424D4F3B0EE363FC1BFAB7FB1798D3" ID-CoL="3Y697" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[989,1172,1406,1430]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="449" phylum="Arthropoda" rank="species" species="secus" status="sp. nov.">
<emphasis id="B936EADE3B0EE363FC1BFAB7FB1798D3" bold="true" box="[989,1172,1406,1430]" italics="true" pageId="53" pageNumber="449">Mariaplax secus</emphasis>
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, 
<taxonomicNameLabel id="A20557A53B0EE363FB62FAB7FAAA98D0" box="[1188,1321,1406,1430]" pageId="53" pageNumber="449" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C35865473B0EE363FBE0FA57FB6398F0" box="[1062,1248,1438,1462]" pageId="53" pageNumber="449" type="description">
<paragraph id="8BFD36CC3B0EE363FBE0FA57FB6398F0" blockId="53.[989,1321,1406,1462]" box="[1062,1248,1438,1462]" pageId="53" pageNumber="449">
(
<figureCitation id="13792A493B0EE363FBE8FA57FB0798F0" box="[1070,1156,1438,1462]" captionStart="Fig" captionStartId="2.[113,148,1860,1882]" captionTargetBox="[117,1446,189,1836]" captionTargetId="figure-11@2.[117,1448,189,1836]" captionTargetPageId="2" captionText="Fig. 1. Colours in life. A, Hexapinus simplex, new species, paratype male (8.7 × 5.0 mm) (MZB), Lombok, Indonesia; B, Hexapinus simplex, new species, male (9.3 × 5.8 mm) (ZRC), Lembeh Straits, Sulawesi Indonesia; C, Hexapinus simplex, new species, male (12.6 × 8.4 mm) (RUMF-ZC-2110), Tekebu, Amami-Ohshima Island, Japan; D, Hexapinus simplex, new species, male (12.6 × 8.4 mm) (ZRC 2013.1688), Funaura, Iriomote Island, Japan; E, Hexapinus simplex, new species, female (8.3 × 6.0 mm) (ZRC 2013.1687), Raffles Reserve near Raffles Lighthouse, Singapore; F, Hexapinus simplex, new species, male (11.2 × 7.0 mm) (ZRC), Pulau Senang, Singapore; G, Mariaplax galaxeae, new species, holotype female (4.2 × 3.2 mm) (ZRC 2013.1690), Singapore Strait; H, Mariaplax chenae, new species, East China Sea (courtsey W.-L. Liu, specimen not examined); I, Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916), Madang, Papua New Guinea; J, Mariaplax mica, new species, holotype female (6.7 × 4.5 mm) (NMCR 39069), Bohol Sea, Philippines." figureDoi="http://doi.org/10.5281/zenodo.5353947" httpUri="https://zenodo.org/record/5353947/files/figure.png" pageId="53" pageNumber="449">Figs. 1I</figureCitation>
, 
<figureCitation id="13792A493B0EE363FB54FA57FB2C98F0" box="[1170,1199,1438,1462]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="53" pageNumber="449">46</figureCitation>
, 
<figureCitation id="13792A493B0EE363FB7AFA57FB5B98F0" box="[1212,1240,1438,1462]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="53" pageNumber="449">47</figureCitation>
)
</paragraph>
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<subSubSection id="C35865473B0EE363FC82FA15FA199B0F" pageId="53" pageNumber="449" type="materials_examined">
<paragraph id="8BFD36CC3B0EE363FC82FA15FA199B0F" blockId="53.[836,1474,1500,1609]" pageId="53" pageNumber="449">
<emphasis id="B936EADE3B0EE363FC82FA15FB9598B4" bold="true" box="[836,1046,1500,1522]" pageId="53" pageNumber="449">Material examined.</emphasis>
<materialsCitation id="3B2A3C913B0EE363FBE6FA14FA199B0F" collectingDate="2012-11-07" collectionCode="MNHN" collectorName="Expedition" country="Papua New Guinea" elevation="135" latitude="-5.233333" location="Bismarck Sea" longLatPrecision="1305" longitude="145.81667" municipality="Cape Kussoron" pageId="53" pageNumber="449" specimenCode="IU-2013-916" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54F9886E3B0EE363FBE6FA14FB0398B5" box="[1056,1152,1501,1523]" pageId="53" pageNumber="449" type="holotype">Holotype</typeStatus>
: male (4.7 × 3.0 mm) (
<collectionCode id="ED53AE093B0EE363FAB0FA14FA4198B4" box="[1398,1474,1501,1522]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="53" pageNumber="449">MNHN</collectionCode>
<specimenCode id="DBE49EB73B0EE363FC82FA30FC469B49" box="[836,965,1529,1551]" collectionCode="IU-" pageId="53" pageNumber="449">IU-2013-916</specimenCode>
), station DW3953, 
<geoCoordinate id="EE76500B3B0EE363FB46FA30FB4F9B49" box="[1152,1228,1529,1551]" degrees="05" direction="south" minutes="14" orientation="latitude" pageId="53" pageNumber="449" precision="925" value="-5.233333">05°14'S</geoCoordinate>
<geoCoordinate id="EE76500B3B0EE363FB14FA30FAAF9B49" box="[1234,1324,1529,1551]" degrees="145" direction="east" minutes="49" orientation="longitude" pageId="53" pageNumber="449" precision="925" value="145.81667">145°49'E</geoCoordinate>
, 
<location id="8E9D60173B0EE363FAF1FA30FA3D9B49" LSID="urn:lsid:plazi:treatment:03EB87DA3B0EE36FFC1BFAB7FC459F11:8E9D60173B0EE363FAF1FA30FA3D9B49" box="[1335,1470,1529,1551]" country="Papua New Guinea" latitude="-5.233333" longLatPrecision="1305" longitude="145.81667" municipality="Cape Kussoron" name="Bismarck Sea" pageId="53" pageNumber="449" stateProvince="Madang">Bismarck Sea</location>
, 
<collectingRegion id="4986F82E3B0EE363FC82F9DFFC169B6A" box="[836,917,1558,1580]" country="Papua New Guinea" name="Madang" pageId="53" pageNumber="449">Madang</collectingRegion>
, south of 
<collectingMunicipality id="6B99ACB63B0EE363FC36F9DFFB079B6A" box="[1008,1156,1558,1580]" pageId="53" pageNumber="449">Cape Kussoron</collectingMunicipality>
, 
<quantity id="4CBA9B293B0EE363FB48F9DFFB7F9B6A" box="[1166,1276,1558,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="53" pageNumber="449" unit="m" value="135.0" valueMax="140.0" valueMin="130.0">
<elevation id="006FD1FF3B0EE363FB48F9DFFB7F9B6A" box="[1166,1276,1558,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="53" pageNumber="449" unit="m" value="135.0" valueMax="140.0" valueMin="130.0">130–140 m</elevation>
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, 
<collectingCountry id="F355765C3B0EE363FAC3F9DEFA3C9B6A" box="[1285,1471,1558,1580]" name="Papua New Guinea" pageId="53" pageNumber="449">Papua New Guinea</collectingCountry>
, coll. 
<collectorName id="26B7531A3B0EE363FCB0F9FAFC5C9B0F" box="[886,991,1587,1609]" pageId="53" pageNumber="449">Expedition</collectorName>
PAPUA NUIGINI, 2012, 
<date id="FFFC100C3B0EE363FB25F9FDFA179B0E" box="[1251,1428,1587,1609]" pageId="53" pageNumber="449" value="2012-11-07">
<collectingDate id="EFB8E9E43B0EE363FB25F9FDFA179B0E" box="[1251,1428,1587,1609]" pageId="53" pageNumber="449" value="2012-11-07">7 November 2012</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35865473B0EE361FC82F9B8FD969F13" lastPageId="55" lastPageNumber="451" pageId="53" pageNumber="449" type="diagnosis">
<paragraph id="8BFD36CC3B0EE361FC82F9B8FE709F53" blockId="53.[836,1475,1649,2090]" lastBlockId="55.[136,776,189,534]" lastPageId="55" lastPageNumber="451" pageId="53" pageNumber="449">
<emphasis id="B936EADE3B0EE363FC82F9B8FC3A9BCF" bold="true" box="[836,953,1649,1673]" pageId="53" pageNumber="449">Diagnosis.</emphasis>
Carapace subquadrate, about 1.6 times as broad as long, dorsal surface strongly granulated; regions indistinct, with inconspicuous H-shaped shallow depression medially (
<figureCitation id="13792A493B0EE363FC8AF918FC3A9BAF" box="[844,953,1745,1769]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="53" pageNumber="449">Figs. 46A</figureCitation>
, 
<figureCitation id="13792A493B0EE363FC03F918FC749BAF" box="[965,1015,1745,1769]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="53" pageNumber="449">47A</figureCitation>
). Eye small, very slightly movable, cornea black, wider than granular peduncle. Pterygostomial region covered with granules, with short row of oblique striae. Third maxillipeds (
<figureCitation id="13792A493B0EE363FC09F8F8FBB99A0F" box="[975,1082,1841,1865]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="53" pageNumber="449">Figs. 46B</figureCitation>
, 
<figureCitation id="13792A493B0EE363FB82F8F8FBF19A0F" box="[1092,1138,1841,1865]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="53" pageNumber="449">47C</figureCitation>
) broad, covering buccal cavity; ischium longer than merus, slightly expanded subproximally, with oblique mesial margin, combined length of dactylus, propodus and carpus almost as long as that of merus and ischium, dactylus as long as propodus; exopod relatively narrow, about 0.4 width of ischium, flagellum well developed. Chelipeds asymmetrical, covered by closely spaced tubercles and very short setae (
<figureCitation id="13792A493B0EE363FBFDF7D8FB26956F" box="[1083,1189,2065,2089]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="53" pageNumber="449">Fig. 46D</figureCitation>
); major chela with broad gaping finger when closed; dactylus and fixed finger with longitudinal ridge at midline distally; palm slightly shorter than broad; minor chela with broad gap when fingers closed; dactylus and fixed finger with median longitudinal ridge distally. P2–P4 (
<figureCitation id="13792A493B0DE360FEE5FEF7FE059C10" box="[291,390,318,342]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="54" pageNumber="450">Fig. 46A</figureCitation>
) relatively long, slender, covered by dense granules; P3 longest, lateral surface of merus with shallow longitudinal groove; dactylus of P4 upcurved, ventral margin with few granules proximally; merus of P4 long, about 4.3 times as long as broad. Male thoracic sternum broad (
<figureCitation id="13792A493B0DE360FAB1FF14FCD89C53" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="54" pageNumber="450">Figs. 46B</figureCitation>
, 
<figureCitation id="13792A493B0DE360FCAEFF34FC149C53" box="[872,919,253,277]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="54" pageNumber="450">47B</figureCitation>
), surface prominently granular; sternites 1 and 2 fused, separated from sternite 3 by distinct furrow; sternite 3 and 4 fused, thoracic sternal groove deep, relatively short, extending obliquely from edge of sternoabdominal cavity (
<figureCitation id="13792A493B0DE360FCF2FEB4FC239CD3" box="[820,928,381,405]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="54" pageNumber="450">Figs. 46B</figureCitation>
, 
<figureCitation id="13792A493B0DE360FC6AFEB4FC5F9CD3" box="[940,988,381,405]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="54" pageNumber="450">47B</figureCitation>
); sternite 8 exposed, quadrangular, slightly longer than male abdominal somite 1; sternoabdominal cavity reaching to middle of sternite 4 (
<figureCitation id="13792A493B0CE361FE3DFF14FDE29DB3" box="[507,609,221,245]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">Fig. 47B</figureCitation>
), distal margin sinuous. Male abdomen broad, short (
<figureCitation id="13792A493B0CE361FDE3FF34FD109C53" box="[549,659,253,277]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="55" pageNumber="451">Figs. 46B</figureCitation>
, 
<figureCitation id="13792A493B0CE361FD59FF34FD529C53" box="[671,721,253,277]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">47D</figureCitation>
), not covering sternoabdominal cavity, reaching only to fifth sternite; surface granular; somites 1 and 2 free, somites 3–6 fused, lateral margin slightly arched; telson subtriangular (
<figureCitation id="13792A493B0CE361FF56FEB4FE829CD3" box="[144,257,381,405]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">Figs. 47D</figureCitation>
), distal margin rounded. G1 (
<figureCitation id="13792A493B0CE361FD8EFEB4FD4B9CD3" box="[584,712,381,405]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">Fig. 47E, F</figureCitation>
) bent ca. 90° at midlength, directed anterolaterally, tuft of long setae on mesial surface near midlength, distal part tapered, not concealed under abdomen, exposed just before thoracic sternal groove. Female unknown.
</paragraph>
<caption id="DF3D66443B0DE360FFB7F7C6FC8A9504" ID-DOI="http://doi.org/10.5281/zenodo.5354039" ID-Zenodo-Dep="5354039" httpUri="https://zenodo.org/record/5354039/files/figure.png" pageId="54" pageNumber="450" startId="54.[113,148,2063,2085]" targetBox="[113,1451,455,2033]" targetPageId="54">
<paragraph id="8BFD36CC3B0DE360FFB7F7C6FC8A9504" blockId="54.[113,1452,2063,2114]" pageId="54" pageNumber="450">
Fig. 43. 
<taxonomicName id="4C424D4F3B0DE360FF02F7C6FE109563" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[196,403,2063,2085]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="54" pageNumber="450" phylum="Arthropoda" rank="species" species="propinqua" status="sp. nov.">
<emphasis id="B936EADE3B0DE360FF02F7C6FE109563" box="[196,403,2063,2085]" italics="true" pageId="54" pageNumber="450">Mariaplax propinqua</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20557A53B0DE360FE67F7C6FD9B9563" box="[417,536,2063,2085]" pageId="54" pageNumber="450" rank="species">new species</taxonomicNameLabel>
, holotype male (10.3 × 6.5 mm) (IOCAS). A, overall view of carapace, chelipeds and P2–P4; B, frontal view of cephalothorax; C, ventral view of cephalothorax.
</paragraph>
</caption>
<paragraph id="8BFD36CC3B0CE361FF4EFDF4FD969F13" blockId="55.[136,533,573,597]" box="[136,533,573,597]" pageId="55" pageNumber="451">
<emphasis id="B936EADE3B0CE361FF4EFDF4FF5E9F13" bold="true" box="[136,221,573,597]" pageId="55" pageNumber="451">Colour.</emphasis>
Dirty white in life (
<figureCitation id="13792A493B0CE361FE7DFDF4FD859F13" box="[443,518,573,597]" captionStart="Fig" captionStartId="2.[113,148,1860,1882]" captionTargetBox="[117,1446,189,1836]" captionTargetId="figure-11@2.[117,1448,189,1836]" captionTargetPageId="2" captionText="Fig. 1. Colours in life. A, Hexapinus simplex, new species, paratype male (8.7 × 5.0 mm) (MZB), Lombok, Indonesia; B, Hexapinus simplex, new species, male (9.3 × 5.8 mm) (ZRC), Lembeh Straits, Sulawesi Indonesia; C, Hexapinus simplex, new species, male (12.6 × 8.4 mm) (RUMF-ZC-2110), Tekebu, Amami-Ohshima Island, Japan; D, Hexapinus simplex, new species, male (12.6 × 8.4 mm) (ZRC 2013.1688), Funaura, Iriomote Island, Japan; E, Hexapinus simplex, new species, female (8.3 × 6.0 mm) (ZRC 2013.1687), Raffles Reserve near Raffles Lighthouse, Singapore; F, Hexapinus simplex, new species, male (11.2 × 7.0 mm) (ZRC), Pulau Senang, Singapore; G, Mariaplax galaxeae, new species, holotype female (4.2 × 3.2 mm) (ZRC 2013.1690), Singapore Strait; H, Mariaplax chenae, new species, East China Sea (courtsey W.-L. Liu, specimen not examined); I, Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916), Madang, Papua New Guinea; J, Mariaplax mica, new species, holotype female (6.7 × 4.5 mm) (NMCR 39069), Bohol Sea, Philippines." figureDoi="http://doi.org/10.5281/zenodo.5353947" httpUri="https://zenodo.org/record/5353947/files/figure.png" pageId="55" pageNumber="451">Fig. 1I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35865473B0CE361FF4EFDB4FE0D9F93" pageId="55" pageNumber="451" type="etymology">
<paragraph id="8BFD36CC3B0CE361FF4EFDB4FE0D9F93" blockId="55.[136,776,637,725]" pageId="55" pageNumber="451">
<emphasis id="B936EADE3B0CE361FF4EFDB4FE8A9FD3" bold="true" box="[136,265,637,661]" pageId="55" pageNumber="451">Etymology.</emphasis>
From the Latin 
<taxonomicName id="4C424D4F3B0CE361FE7CFDB7FE779FD3" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[442,500,638,661]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FE7CFDB7FE779FD3" box="[442,500,638,661]" italics="true" pageId="55" pageNumber="451">secus</emphasis>
</taxonomicName>
, for different; alluding to the unusual shape of the male abdomen. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C35865473B0CE36FFF4EFD34FAB89CD1" lastPageId="57" lastPageNumber="453" pageId="55" pageNumber="451" type="discussion">
<paragraph id="8BFD36CC3B0CE36FFF4EFD34FD129DB0" blockId="55.[136,776,765,821]" lastBlockId="57.[136,776,190,246]" lastPageId="57" lastPageNumber="453" pageId="55" pageNumber="451">
<emphasis id="B936EADE3B0CE361FF4EFD34FF759E53" bold="true" box="[136,246,765,789]" pageId="55" pageNumber="451">Remarks.</emphasis>
This is the second species in 
<taxonomicName id="4C424D4F3B0CE361FDE5FD34FD119E53" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[547,658,765,789]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B0CE361FDE5FD34FD119E53" box="[547,658,765,789]" italics="true" pageId="55" pageNumber="451">Mariaplax</emphasis>
</taxonomicName>
to have the G1 bent at a 90° angle, forming a L-shaped structure; the other species being 
<taxonomicName id="4C424D4F3B0CE361FC1FFF76FB019D90" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[985,1154,190,214]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus" status="sp. nov.">
<emphasis id="B936EADE3B0CE361FC1FFF76FB019D90" box="[985,1154,190,214]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20557A53B0CE361FB49FF76FA919D90" box="[1167,1298,190,214]" pageId="55" pageNumber="451" rank="species">new species</taxonomicNameLabel>
. The differences between these two species are as follow: the carapace is more rectangular, being 1.6 times as broad as long in 
<taxonomicName id="4C424D4F3B0CE361FAA7FF36FA419C50" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1377,1474,255,278]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FAA7FF36FA419C50" box="[1377,1474,255,278]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FC8AFED7FC339C70" box="[844,944,286,310]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="55" pageNumber="451">Fig. 46A</figureCitation>
) (1.5 times as broad as long in 
<taxonomicName id="4C424D4F3B0CE361FAD6FED6FA389C70" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1296,1467,286,310]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FAD6FED6FA389C70" box="[1296,1467,286,310]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B0CE361FC82FEF7FC2B9C10" box="[836,936,318,342]" captionStart="Fig" captionStartId="36.[113,148,2092,2114]" captionTargetBox="[113,1451,572,2059]" captionTargetId="figure-34@36.[113,1451,572,2059]" captionTargetPageId="36" captionText="Fig. 28. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB).A, dorsal view carapace; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5354005" httpUri="https://zenodo.org/record/5354005/files/figure.png" pageId="55" pageNumber="451">Fig. 28A</figureCitation>
); the ischium of the third maxilliped is as long as broad with the dactylus the same length as the propodus in 
<taxonomicName id="4C424D4F3B0CE361FCA7FEB6FC439CD0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[865,960,383,406]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FCA7FEB6FC439CD0" box="[865,960,383,406]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FC16FEB7FBB29CD0" box="[976,1073,382,406]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">Fig. 47C</figureCitation>
) (slightly longer than broad with the dactylus much longer than the propodus in 
<taxonomicName id="4C424D4F3B0CE361FAD4FE56FA389CF0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1298,1467,414,438]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FAD4FE56FA389CF0" box="[1298,1467,414,438]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B0CE361FC82FE77FC209C90" box="[836,931,446,470]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="55" pageNumber="451">Fig. 29E</figureCitation>
); the thoracic sternum is proportionately less broad in 
<taxonomicName id="4C424D4F3B0CE361FCA7FE16FC429CB0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[865,961,479,502]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FCA7FE16FC429CB0" box="[865,961,479,502]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
compared to that of 
<taxonomicName id="4C424D4F3B0CE361FB61FE16FAD29CB0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1191,1361,478,502]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FB61FE16FAD29CB0" box="[1191,1361,478,502]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FAA4FE17FA419CB0" box="[1378,1474,478,502]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">Fig. 47B</figureCitation>
versus 
<figureCitation id="13792A493B0CE361FC57FE37FC779F50" box="[913,1012,510,534]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="55" pageNumber="451">Fig. 29B</figureCitation>
); the distal part of sternoabdominal cavity is relatively long and narrow, with distal margin sinuous in 
<taxonomicName id="4C424D4F3B0CE361FC82FDF6FC219F10" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[836,930,575,598]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FC82FDF6FC219F10" box="[836,930,575,598]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FC77FDF7FB9F9F10" box="[945,1052,574,598]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="55" pageNumber="451">Figs. 46B</figureCitation>
, 
<figureCitation id="13792A493B0CE361FBE1FDF7FBD69F10" box="[1063,1109,574,598]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="55" pageNumber="451">47B</figureCitation>
) (short, broad with rounded distal margin in 
<taxonomicName id="4C424D4F3B0CE361FC7CFD96FBE99F30" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[954,1130,606,630]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FC7CFD96FBE99F30" box="[954,1130,606,630]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B0CE361FBBDFD97FB619F30" box="[1147,1250,606,630]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="55" pageNumber="451">Fig. 29B</figureCitation>
); the lateral surface of the P3 merus has a distinct shallow longitudinal groove in 
<taxonomicName id="4C424D4F3B0CE361FCA4FD56FC409FF0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[866,963,671,694]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FCA4FD56FC409FF0" box="[866,963,671,694]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FC13FD57FBBA9FF0" box="[981,1081,670,694]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="55" pageNumber="451">Fig. 46A</figureCitation>
) (an indistinct groove, obscured by tubercles, in 
<taxonomicName id="4C424D4F3B0CE361FC1EFD76FB0A9F90" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[984,1161,702,726]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FC1EFD76FB0A9F90" box="[984,1161,702,726]" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B0CE361FB5CFD77FB7A9F90" box="[1178,1273,702,726]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="55" pageNumber="451">Fig. 29I</figureCitation>
); the P4 merus is relatively much longer, being 4.3 times as long as broad in 
<taxonomicName id="4C424D4F3B0CE361FC82FD36FC229E50" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[836,929,767,790]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B0CE361FC82FD36FC229E50" box="[836,929,767,790]" italics="true" pageId="55" pageNumber="451">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B0CE361FC69FD37FB939E50" box="[943,1040,766,790]" captionStart="Fig" captionStartId="57.[136,171,2061,2083]" captionTargetBox="[135,1473,685,2028]" captionTargetId="figure-240@57.[135,1473,685,2030]" captionTargetPageId="57" captionText="Fig. 46. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, overall view of carapace, chelipeds and P2–P4; B, ventral view of cephalothorax; C, front view of cephalothorax; D, major chela and minor chela." figureDoi="http://doi.org/10.5281/zenodo.5354045" httpUri="https://zenodo.org/record/5354045/files/figure.png" pageId="55" pageNumber="451">Fig. 46A</figureCitation>
) (versus 3.4 times as long as broad in 
<taxonomicName id="4C424D4F3B0CE361FA63FD36FC4A9E70" authorityName="Rahayu &amp; Ng" authorityYear="2014" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B0CE361FA63FD36FC4A9E70" italics="true" pageId="55" pageNumber="451">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B0CE361FC11FCD7FBAD9E70" box="[983,1070,798,822]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="55" pageNumber="451">Fig. 29I</figureCitation>
); and the G1 has a tuft of long medial setae in 
<taxonomicName id="4C424D4F3B02E36FFF24FF76FEC29D90" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[226,321,191,214]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B02E36FFF24FF76FEC29D90" box="[226,321,191,214]" italics="true" pageId="57" pageNumber="453">M. secus</emphasis>
</taxonomicName>
(
<figureCitation id="13792A493B02E36FFE97FF77FE4D9D90" box="[337,462,190,214]" captionStart="Fig" captionStartId="58.[113,148,2032,2054]" captionTargetBox="[124,1447,215,1969]" captionTargetId="figure-11@58.[96,1489,180,2007]" captionTargetPageId="58" captionText="Fig. 47. Mariaplax secus, new species, holotype male (4.7 × 3.0 mm) (MNHN IU-2013-916). A, dorsolateral margin of carapace; B, thoracic sternites 1–6, showing sternoabdominal cavity, thoracic sternal groove and telson; C, left third maxilliped; D, male abdomen and telson; E, F, G1. Scale bars: A, B, D = 1.0 mm; C, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5354047" httpUri="https://zenodo.org/record/5354047/files/figure.png" pageId="57" pageNumber="453">Fig. 47E, F</figureCitation>
) (few setae along dorsal and ventral margin in 
<taxonomicName id="4C424D4F3B02E36FFE8AFF16FE759DB0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[332,502,222,246]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B02E36FFE8AFF16FE759DB0" box="[332,502,222,246]" italics="true" pageId="57" pageNumber="453">M. cyrtophallus</emphasis>
</taxonomicName>
, 
<figureCitation id="13792A493B02E36FFDC0FF17FD079DB0" box="[518,644,222,246]" captionStart="Fig" captionStartId="37.[136,171,2002,2024]" captionTargetBox="[158,1434,194,1961]" captionTargetId="figure-7@37.[140,1467,170,1980]" captionTargetPageId="37" captionText="Fig. 29. Mariaplax cyrtophallus, new species, holotype male (6.5 × 3.8 mm) (MZB). A, dorso-lateral margin of carapace; B, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; C, thoracic sternites 1–5, showing thoracic sternal groove and telson; D, anterior view of cephalothorax; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, left P4; J, K, left G1. Tubercles on thoracic sternum omitted. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5354007" httpUri="https://zenodo.org/record/5354007/files/figure.png" pageId="57" pageNumber="453">Fig. 29J, K</figureCitation>
).
</paragraph>
<caption id="DF3D66443B0CE361FF4EF7E3FA469506" ID-DOI="http://doi.org/10.5281/zenodo.5354041" ID-Zenodo-Dep="5354041" box="[136,1477,2090,2113]" httpUri="https://zenodo.org/record/5354041/files/figure.png" pageId="55" pageNumber="451" startId="55.[136,171,2090,2112]" targetBox="[138,1474,866,2065]" targetPageId="55">
<paragraph id="8BFD36CC3B0CE361FF4EF7E3FA469506" blockId="55.[136,1477,2090,2113]" box="[136,1477,2090,2113]" pageId="55" pageNumber="451">
Fig. 44. 
<taxonomicName id="4C424D4F3B0CE361FF10F7E3FE209506" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[214,419,2090,2112]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="451" phylum="Arthropoda" rank="species" species="propinqua" status="sp. nov.">
<emphasis id="B936EADE3B0CE361FF10F7E3FE209506" box="[214,419,2090,2112]" italics="true" pageId="55" pageNumber="451">Mariaplax propinqua</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20557A53B0CE361FE68F7E2FDA09506" box="[430,547,2090,2112]" pageId="55" pageNumber="451" rank="species">new species</taxonomicNameLabel>
, holotype male (10.3 × 6.5 mm) (IOCAS). A, abdomen and telson; B, major chela; C, minor chela.
</paragraph>
</caption>
<caption id="DF3D66443B03E36EFFB7F81CFE8A9507" ID-DOI="http://doi.org/10.5281/zenodo.5354043" ID-Zenodo-Dep="5354043" httpUri="https://zenodo.org/record/5354043/files/figure.png" pageId="56" pageNumber="452" startId="56.[113,148,2005,2027]" targetBox="[124,1438,217,1956]" targetPageId="56">
<paragraph id="8BFD36CC3B03E36EFFB7F81CFE8A9507" blockId="56.[113,1452,2005,2113]" pageId="56" pageNumber="452">
Fig. 45. 
<taxonomicName id="4C424D4F3B03E36EFF04F81CFE129AAC" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[194,401,2005,2026]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="452" phylum="Arthropoda" rank="species" species="propinqua" status="sp. nov.">
<emphasis id="B936EADE3B03E36EFF04F81CFE129AAC" box="[194,401,2005,2026]" italics="true" pageId="56" pageNumber="452">Mariaplax propinqua</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20557A53B03E36EFE5BF81CFD979AAD" box="[413,532,2005,2027]" pageId="56" pageNumber="452" rank="species">new species</taxonomicNameLabel>
, holotype male (10.3 × 6.5 mm) (IOCAS). A, dorsolateral margin of carapace; B, anterior view of cephalothorax; C, thoracic sternites 3–5, showing thoracic sternal groove and telson; D, thoracic sternites 1–5, showing sternoabdominal cavity and thoracic sternal groove; E, left third maxilliped; F, male abdomen and telson, G, major chela; H, minor chela; I, J, G1. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BFD36CC3B02E36FFF4EFED7FB949C5E" blockId="57.[136,776,286,599]" lastBlockId="57.[836,1475,192,280]" pageId="57" pageNumber="453">
The male abdomen in 
<taxonomicName id="4C424D4F3B02E36FFEBCFED6FE5B9C70" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[378,472,287,310]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B02E36FFEBCFED6FE5B9C70" box="[378,472,287,310]" italics="true" pageId="57" pageNumber="453">M. secus</emphasis>
</taxonomicName>
is unusual, being broad and very short, with somite 6 apparently fused to somites 3–5. It is completely different from all know hexapodids. However, this is likely to be associated with some parasitic infection (e.g., a rhizocephalan as it appears unusually swollen), or the result of past damage and anomalous regrowth. As such, two of the other observed differences between 
<taxonomicName id="4C424D4F3B02E36FFD93FE16FD379CB0" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[597,692,479,502]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B02E36FFD93FE16FD379CB0" box="[597,692,479,502]" italics="true" pageId="57" pageNumber="453">M. secus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C424D4F3B02E36FFD2DFE16FE8F9F50" authorityName="Rahayu &amp; Ng" authorityYear="2014" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B02E36FFD2DFE16FE8F9F50" italics="true" pageId="57" pageNumber="453">M. cyrtophallus</emphasis>
</taxonomicName>
: the differently structured male sternoabdominal cavities and G1s, should be used with caution. It is possible they have also been affected by the above factors. In any case, the differences in the carapace, third maxillipeds, thoracic sternum and P2–P4 argue for them to be recognised as separate species.
</paragraph>
<paragraph id="8BFD36CC3B02E36FFC82FE89FAB89CD1" blockId="57.[836,1475,320,408]" pageId="57" pageNumber="453">
Interestingly, both 
<taxonomicName id="4C424D4F3B02E36FFBD6FE89FB389C1E" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1040,1211,320,344]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="cyrtophallus">
<emphasis id="B936EADE3B02E36FFBD6FE89FB389C1E" box="[1040,1211,320,344]" italics="true" pageId="57" pageNumber="453">M. cyrtophallus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C424D4F3B02E36FFB35FE89FAD19C11" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[1267,1362,320,343]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="453" phylum="Arthropoda" rank="species" species="secus">
<emphasis id="B936EADE3B02E36FFB35FE89FAD19C11" box="[1267,1362,320,343]" italics="true" pageId="57" pageNumber="453">M. secus</emphasis>
</taxonomicName>
are found in the island of Indonesian Papua – 
<collectingCountry id="F355765C3B02E36FFB03FEA9FA149C3E" box="[1221,1431,352,376]" name="Papua New Guinea" pageId="57" pageNumber="453">Papua New Guinea</collectingCountry>
, the latter from the east, the former from the west.
</paragraph>
</subSubSection>
<subSubSection id="C35865473B02E36FFC82FE09FBD09CBE" pageId="57" pageNumber="453" type="materials_examined">
<paragraph id="8BFD36CC3B02E36FFC82FE09FBD09CBE" blockId="57.[836,1474,447,504]" pageId="57" pageNumber="453">
<materialsCitation id="3B2A3C913B02E36FFC82FE09FBD39CBE" country="Papua New Guinea" location="Cape Kussoron" municipality="Bismarck Sea" pageId="57" pageNumber="453" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="B936EADE3B02E36FFC82FE09FC5F9C91" bold="true" box="[836,988,447,472]" pageId="57" pageNumber="453">
<typeStatus id="54F9886E3B02E36FFC82FE09FCFF9C9E" box="[836,892,448,472]" pageId="57" pageNumber="453">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="4986F82E3B02E36FFC22FE09FBBD9C9E" box="[996,1086,448,472]" country="Papua New Guinea" name="Madang" pageId="57" pageNumber="453">Madang</collectingRegion>
, south of 
<location id="8E9D60173B02E36FFB6FFE09FACE9C91" LSID="urn:lsid:plazi:treatment:03EB87DA3B0EE36FFC1BFAB7FC459F11:8E9D60173B02E36FFB6FFE09FACE9C91" box="[1193,1357,448,472]" country="Papua New Guinea" municipality="Bismarck Sea" name="Cape Kussoron" pageId="57" pageNumber="453" stateProvince="Madang">Cape Kussoron</location>
, 
<collectingMunicipality id="6B99ACB63B02E36FFA9AFE09FCEE9CBE" pageId="57" pageNumber="453">Bismarck Sea</collectingMunicipality>
, 
<collectingCountry id="F355765C3B02E36FFCBCFE29FBD39CBE" box="[890,1104,480,504]" name="Papua New Guinea" pageId="57" pageNumber="453">Papua New Guinea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35865473B02E36FFC82FDD6FC459F11" pageId="57" pageNumber="453" type="distribution">
<paragraph id="8BFD36CC3B02E36FFC82FDD6FC459F11" blockId="57.[836,1473,543,600]" pageId="57" pageNumber="453">
<emphasis id="B936EADE3B02E36FFC82FDD6FC609F71" bold="true" box="[836,995,543,567]" pageId="57" pageNumber="453">Distribution.</emphasis>
<collectingRegion id="4986F82E3B02E36FFC36FDE9FBD39F7E" box="[1008,1104,544,568]" country="Papua New Guinea" name="Madang" pageId="57" pageNumber="453">Madang</collectingRegion>
, 
<collectingCountry id="F355765C3B02E36FFBA7FDE9FAC99F7E" box="[1121,1354,544,568]" name="Papua New Guinea" pageId="57" pageNumber="453">Papua New Guinea</collectingCountry>
. Subtidal, 
<quantity id="4CBA9B293B02E36FFC82FD89FC3C9F11" box="[836,959,576,600]" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="57" pageNumber="453" unit="m" value="135.0" valueMax="140.0" valueMin="130.0">
130– 
<specimenCount id="9D44FD453B02E36FFCBFFD89FC3C9F11" box="[889,959,576,600]" pageId="57" pageNumber="453" type="generic">140 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>