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<document id="221749DA7AE71D06D1799044D617D5AD" ID-CLB-Dataset="31260" ID-DOI="10.11646/zootaxa.4359.1.1" ID-GBIF-Dataset="a5cbd5ef-7f49-4a94-a2f4-61adcc8c8d01" ID-ISSN="1175-5326" ID-Zenodo-Dep="1069168" ID-ZooBank="8D06174D-B19F-4B5C-B9B0-DA74E6D43C75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512122415347" checkinUser="plazi" docAuthor="Tibiriçá, Yara, Pola, Marta &amp; Cervera, Juan Lucas" docDate="2017" docId="03EB87A2FFE9FFBD9790FCAEFAFCFA78" docLanguage="en" docName="zootaxa.4359.1.1.pdf.imf" docOrigin="Zootaxa 4359 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phyllidia haegeli" docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFD2FFDAFFD0FF849707FF83FFB7FFC0" masterDocTitle="Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique" masterLastPageNumber="133" masterPageNumber="1" pageNumber="58" updateTime="1698506029818" updateUser="ExternalLinkService">
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<mods:title id="D9CCC8ADF358BBD8128E682C35E52B01">Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique</mods:title>
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<mods:namePart id="DE744ABCFECB0EDBD9A261BEAF02EE29">Tibiriçá, Yara</mods:namePart>
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<mods:namePart id="26441494313A40B768591CC6D99954EC">Pola, Marta</mods:namePart>
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<mods:namePart id="F514F7C1EC55B32CEC80725D8D4FAB73">Cervera, Juan Lucas</mods:namePart>
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<mods:title id="CE74093C859B3328379B663A91CD498A">Zootaxa</mods:title>
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<subSubSection id="C358653FFFE9FFBD9790FCAEFEF8FCA8" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BFD36B4FFE9FFBD9790FCAEFD0EFC87" blockId="57.[151,697,813,872]" box="[151,697,813,839]" pageId="57" pageNumber="58">
<heading id="D0B581D8FFE9FFBD9790FCAEFD0EFC87" bold="true" box="[151,697,813,839]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<taxonomicName id="4C424D37FFE9FFBD9790FCAEFD0EFC87" ID-CoL="4H4XL" authority="(Fahrner &amp; Beck, 2000)" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" box="[151,697,813,839]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD9790FCAEFD0EFC87" bold="true" box="[151,697,813,839]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD9790FCAEFEB3FC87" bold="true" box="[151,260,813,839]" italics="true" pageId="57" pageNumber="58">Phyllidia</emphasis>
cf.
<emphasis id="B936EAA6FFE9FFBD9636FCAEFE3EFC87" bold="true" box="[305,393,813,839]" italics="true" pageId="57" pageNumber="58">haegeli</emphasis>
(
<bibRefCitation id="EFD34B45FFE9FFBD9690FCAEFD06FC87" author="Fahrner" box="[407,689,813,839]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Beck, L. A. (2000) Identification key to the Indo-Pacific species of the nudibranch family Phyllidiidae Rafinesque, 1814 including the description of two new species. Archiv fur Molluskenkunde, 128, 189 - 211." type="journal article" year="2000">Fahrner &amp; Beck, 2000</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFD36B4FFE9FFBD9790FCCCFEF8FCA8" blockId="57.[151,697,813,872]" box="[151,335,847,872]" pageId="57" pageNumber="58">
(
<figureCitation id="13792A31FFE9FFBD9799FCCCFEF1FCA8" box="[158,326,847,872]" captionStart="FIGURE 19" captionStartId="58.[151,250,1921,1943]" captionTargetBox="[215,1372,195,1896]" captionTargetId="figure@58.[215,1372,195,1896]" captionTargetPageId="58" captionText="FIGURE 19. A, Sclerodoris tuberculata (MB28-005086). BC, Phyllidia alyta (MB28-004883) dorsal and ventral view.DE, Phyllidia coelestis (MB28-004781 and MB28-004896, respectively). FG Phyllidia cf. haegeli (ZMBN117018) dorsal and ventral view. HI, Phyllidia marindica (MB28-004780) dorsal and ventral view. J, Phyllidia ocellata (MB28-005092). KL, Phyllidia varicosa (MB28-004450) dorsal and ventral view." httpUri="https://zenodo.org/record/1069209/files/figure.png" pageId="57" pageNumber="58">Figure 19 FG</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFE9FFBD9790FC14FD89FC13" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BFD36B4FFE9FFBD9790FC14FBC3FC70" blockId="57.[151,1436,919,1464]" box="[151,1140,919,944]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD9790FC14FE37FC70" bold="true" box="[151,384,919,944]" pageId="57" pageNumber="58">Material examined.</emphasis>
One specimen. ZMBN117018, ZGWS, 0
<date id="FFFC1074FFE9FFBD9455FC1BFC6DFC70" box="[850,986,919,944]" pageId="57" pageNumber="58" value="2012-02-02">
2 Feb.
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</date>
, 21m,
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.
</paragraph>
<paragraph id="8BFD36B4FFE9FFBD97C0FC39FD89FC13" blockId="57.[151,1436,919,1464]" box="[199,574,954,979]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD97C0FC39FE82FC13" bold="true" box="[199,309,954,979]" pageId="57" pageNumber="58">Habitats.</emphasis>
Subtropical rocky reef.
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFE9FFBD97C0FC5EFA24FBD8" pageId="57" pageNumber="58" type="description">
<paragraph id="8BFD36B4FFE9FFBD97C0FC5EFE75FC35" blockId="57.[151,1436,919,1464]" box="[199,450,989,1014]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD97C0FC5EFED3FC36" bold="true" box="[199,356,989,1014]" pageId="57" pageNumber="58">Occurrences.</emphasis>
Zavora.
</paragraph>
<paragraph id="8BFD36B4FFE9FFBD97C0FC7CFA24FBD8" blockId="57.[151,1436,919,1464]" box="[199,1427,1023,1048]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD97C0FC7CFE58FBD8" bold="true" box="[199,495,1023,1048]" pageId="57" pageNumber="58">Geographic distribution.</emphasis>
Indo-west Pacific.
<collectingCountry id="F3557624FFE9FFBD95CFFC7CFC82FBD8" box="[712,821,1023,1048]" name="Indonesia" pageId="57" pageNumber="58">Indonesia</collectingCountry>
(
<bibRefCitation id="EFD34B45FFE9FFBD9444FC7CFB8CFBD8" author="Fahrner" box="[835,1083,1023,1048]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Beck, L. A. (2000) Identification key to the Indo-Pacific species of the nudibranch family Phyllidiidae Rafinesque, 1814 including the description of two new species. Archiv fur Molluskenkunde, 128, 189 - 211." type="journal article" year="2000">Fahrner &amp; Beck 2000</bibRefCitation>
) and potentially
<collectingCountry id="F3557624FFE9FFBD93F0FC7CFA38FBD8" box="[1271,1423,1023,1048]" name="Mozambique" pageId="57" pageNumber="58">Mozambique</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFE9FFBD97C0FBA1FAFCFA78" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BFD36B4FFE9FFBD97C0FBA1FAFCFA78" blockId="57.[151,1436,919,1464]" pageId="57" pageNumber="58">
<emphasis id="B936EAA6FFE9FFBD97C0FBA1FE8CFBFB" bold="true" box="[199,315,1058,1083]" pageId="57" pageNumber="58">Remarks.</emphasis>
This specimen shares features with
<taxonomicName id="4C424D37FFE9FFBD95C8FBA7FCFCFBFB" authorityName="Lamarck" authorityYear="1801" box="[719,843,1060,1083]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="varicosa">
<emphasis id="B936EAA6FFE9FFBD95C8FBA7FCFCFBFB" box="[719,843,1060,1083]" italics="true" pageId="57" pageNumber="58">P. varicosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C424D37FFE9FFBD945EFBA7FC63FBFB" authorityName="in Yonow" authorityYear="2012" box="[857,980,1058,1083]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="coelestis">
<emphasis id="B936EAA6FFE9FFBD945EFBA7FC63FBFB" box="[857,980,1058,1083]" italics="true" pageId="57" pageNumber="58">P. coelestis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C424D37FFE9FFBD94E5FBA7FBC7FBFB" baseAuthorityName="Yonow &amp; Hayward" baseAuthorityYear="1991" box="[994,1136,1058,1083]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="marindica">
<emphasis id="B936EAA6FFE9FFBD94E5FBA7FBC7FBFB" box="[994,1136,1058,1083]" italics="true" pageId="57" pageNumber="58">P. marindica</emphasis>
</taxonomicName>
and particularly
<taxonomicName id="4C424D37FFE9FFBD9229FBA7FE15FB9E" authority="(Fahrner &amp; Beck, 2000)" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD9229FBA7FA2BFBFB" box="[1326,1436,1058,1083]" italics="true" pageId="57" pageNumber="58">P. haegeli</emphasis>
(
<bibRefCitation id="EFD34B45FFE9FFBD9799FBC6FE2CFB9E" author="Fahrner" box="[158,411,1093,1118]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Beck, L. A. (2000) Identification key to the Indo-Pacific species of the nudibranch family Phyllidiidae Rafinesque, 1814 including the description of two new species. Archiv fur Molluskenkunde, 128, 189 - 211." type="journal article" year="2000">Fahrner &amp; Beck, 2000</bibRefCitation>
)
</taxonomicName>
. All have rhinotubercles, a greyish background, ridges formed by yellow-capped tubercles, black bands and golden-yellow rhinophores (
<bibRefCitation id="EFD34B45FFE9FFBD9594FBE4FCD5FB40" author="Brunckhorst" box="[659,866,1127,1152]" pageId="57" pageNumber="58" refString="Brunckhorst, D. J. (1993) The Systematics and Phylogeny of Phyllidiid Nudibranchs (Doridoidea). Records of the Australian Museum, 16 (Supplement), 1 - 107. https: // doi. org / 10.3853 / j. 0812 - 7387.16.1993.79" type="journal article" year="1993">Brunckhorst 1993</bibRefCitation>
;
<bibRefCitation id="EFD34B45FFE9FFBD9468FBE4FBDDFB40" author="Fahrner" box="[879,1130,1127,1152]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Beck, L. A. (2000) Identification key to the Indo-Pacific species of the nudibranch family Phyllidiidae Rafinesque, 1814 including the description of two new species. Archiv fur Molluskenkunde, 128, 189 - 211." type="journal article" year="2000">Fahrner &amp; Beck 2000</bibRefCitation>
).
<taxonomicName id="4C424D37FFE9FFBD937BFBE4FAFCFB40" authorityName="in Yonow" authorityYear="2012" box="[1148,1355,1127,1152]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="coelestis">
<emphasis id="B936EAA6FFE9FFBD937BFBE4FAFCFB40" box="[1148,1355,1127,1152]" italics="true" pageId="57" pageNumber="58">Phyllidia coelestis</emphasis>
</taxonomicName>
differs by having a middle longitudinal black band and
<taxonomicName id="4C424D37FFE9FFBD95BBFB09FC29FB63" baseAuthorityName="Yonow &amp; Hayward" baseAuthorityYear="1991" box="[700,926,1162,1187]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="marindica">
<emphasis id="B936EAA6FFE9FFBD95BBFB09FC29FB63" box="[700,926,1162,1187]" italics="true" pageId="57" pageNumber="58">Phyllidia marindica</emphasis>
</taxonomicName>
by a single median ridge.
<taxonomicName id="4C424D37FFE9FFBD93CAFB09FA2BFB63" authorityName="Lamarck" authorityYear="1801" box="[1229,1436,1162,1187]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="varicosa">
<emphasis id="B936EAA6FFE9FFBD93CAFB09FA2BFB63" box="[1229,1436,1162,1187]" italics="true" pageId="57" pageNumber="58">Phyllidia varicosa</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C424D37FFE9FFBD9608FB2EFE67FB06" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" box="[271,464,1197,1222]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD9608FB2EFE67FB06" box="[271,464,1197,1222]" italics="true" pageId="57" pageNumber="58">Phyllidia haegeli</emphasis>
</taxonomicName>
dorsally, but living specimens have a distinctive line on the sole of the foot and they are usually larger, lighter in colour and less flattened (
<bibRefCitation id="EFD34B45FFE9FFBD941CFB4CFB95FB28" author="Fahrner" box="[795,1058,1231,1256]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Beck, L. A. (2000) Identification key to the Indo-Pacific species of the nudibranch family Phyllidiidae Rafinesque, 1814 including the description of two new species. Archiv fur Molluskenkunde, 128, 189 - 211." type="journal article" year="2000">Fahrner &amp; Beck 2000</bibRefCitation>
,
<bibRefCitation id="EFD34B45FFE9FFBD9336FB4CFAE2FB28" author="Fahrner" box="[1073,1365,1231,1256]" pageId="57" pageNumber="58" refString="Fahrner, A. &amp; Schrodl, M. (2000) Redescription of Phyllidiopsis sinaiensis (Yonow, 1988) (Nudibranchia: Doridoidea: Phyllidiidae), with a review of the Red Sea Phyllidiidae. Journal of Molluscan Studies, 66, 467 - 476. https: // doi. org / 10.1093 / mollus / 66.4.467" type="journal article" year="2000">Fahrner &amp; Schrödl 2000</bibRefCitation>
). Few differences exist between our specimen and
<taxonomicName id="4C424D37FFE9FFBD9588FB71FCE7FACB" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" box="[655,848,1266,1291]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD9588FB71FCE7FACB" box="[655,848,1266,1291]" italics="true" pageId="57" pageNumber="58">Phyllidia haegeli</emphasis>
</taxonomicName>
. The collected specimen has outer black bands that unite anterior to the rhinophores, while in
<taxonomicName id="4C424D37FFE9FFBD957DFA95FD5DFAEE" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" box="[634,746,1301,1326]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD957DFA95FD3FFAED" box="[634,648,1302,1325]" italics="true" pageId="57" pageNumber="58">P</emphasis>
.
<emphasis id="B936EAA6FFE9FFBD959FFA96FD5DFAEE" box="[664,746,1301,1326]" italics="true" pageId="57" pageNumber="58">haegeli</emphasis>
</taxonomicName>
they converge but do not merge. Furthermore, the specimen exhibits inner longitudinal bands that unite posteriorly but in
<taxonomicName id="4C424D37FFE9FFBD945AFABAFC78FA90" baseAuthorityName="Fahrner &amp; Beck" baseAuthorityYear="2000" box="[861,975,1335,1360]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="haegeli">
<emphasis id="B936EAA6FFE9FFBD945AFABAFCDCFA90" box="[861,875,1337,1360]" italics="true" pageId="57" pageNumber="58">P</emphasis>
.
<emphasis id="B936EAA6FFE9FFBD947BFAB4FC78FA90" box="[892,975,1335,1360]" italics="true" pageId="57" pageNumber="58">haegeli</emphasis>
</taxonomicName>
they are shorter and do not merge. The main distinction is associated with the outer longitudinal black bands: in the collected specimen, these bands are shorter and connect with the radial lines to create an inverted “V”. As
<taxonomicName id="4C424D37FFE9FFBD94B6FAFEFBAFFA56" authorityName="Cuvier" authorityYear="1797" box="[945,1048,1405,1430]" class="Gastropoda" family="Phyllidiidae" genus="Phyllidia" kingdom="Animalia" order="Nudibranchia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B936EAA6FFE9FFBD94B6FAFEFBAFFA56" box="[945,1048,1405,1430]" italics="true" pageId="57" pageNumber="58">Phyllidia</emphasis>
</taxonomicName>
can be highly variable (
<bibRefCitation id="EFD34B45FFE9FFBD922FFAFEFF40FA78" author="Stoffels" pageId="57" pageNumber="58" refString="Stoffels, B., van der Meij, S E T Hoeksema, B. W., van Alphen, J., van Alen, T., Meyers-Munoz, M. A., de Voogd, N. J., Tuti, Y. &amp; van der Velde, G. (2016) Phylogenetic relationships within the Phyllidiidae (Opisthobranchia, Nudibranchia). ZooKeys, 605, 1 - 35. https: // doi. org / 10.3897 / zookeys. 605.7136" type="journal article" year="2016">
Stoffels
<emphasis id="B936EAA6FFE9FFBD928FFAFDFF03FA78" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
2016
</bibRefCitation>
), it is not possible to confirm the taxonomic status without further molecular or anatomical review.
</paragraph>
</subSubSection>
</treatment>
</document>