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<document id="5E232C29E31099B39782BCCC3619A3F9" ID-DOI="10.11646/zootaxa.5497.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618611" ID-ZooBank="C107D617-5F70-42C5-8FD4-E5C2DE0E48B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725026357643" checkinUser="plazi" docAuthor="Khaustov, Alexander A., Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2024" docId="03EB878FFFBE744F81B438B7FB10FF16" docLanguage="en" docName="zootaxa.5497.3.2.pdf" docOrigin="Zootaxa 5497 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus madagascariensis Khaustov, Frolov &amp; Akhmetova, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="365" masterDocId="FFD2FFF7FFA6745381233D58FFDEFFEF" masterDocTitle="Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)" masterLastPageNumber="368" masterPageNumber="337" pageNumber="361" updateTime="1725028125811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="240BA235D6EB456C6DE29684AF993195">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="AB2B156D908577409FE7A53705AFBD60">Khaustov, Alexander A.</mods:namePart>
<mods:affiliation id="6BA93B3605E2DAF25E2944CA0936778D">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
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<mods:namePart id="DCCECF8792BB100DE84BC7DA029B3045">Frolov, Andrey V.</mods:namePart>
<mods:affiliation id="7F7586DD708BF56982198A886011B76F">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="8C50BBCCCCBC76E9974E143561C414D0">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="428DCBA62F0F36B2B94F61FE860BD308">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:title id="1ADEFC140A32ACB261E527AB3006D264">Zootaxa</mods:title>
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<mods:date id="69325950FA388C5D7BEB3C163090F072">2024</mods:date>
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<mods:number id="9C719DF1D3455ED47AFCF82ED666DE3B">2024-08-26</mods:number>
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<mods:number id="AF3BE221CAF43474061B5B697E9577E1">5497</mods:number>
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<treatment id="03EB878FFFBE744F81B438B7FB10FF16" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16" httpUri="http://treatment.plazi.org/id/03EB878FFFBE744F81B438B7FB10FF16" lastPageId="28" lastPageNumber="365" pageId="24" pageNumber="361">
<subSubSection id="C3586512FFBE744B81B438B7FD52F9E5" box="[151,652,1519,1546]" pageId="24" pageNumber="361" type="nomenclature">
<paragraph id="8BFD3699FFBE744B81B438B7FD52F9E5" blockId="24.[151,652,1519,1583]" box="[151,652,1519,1546]" pageId="24" pageNumber="361">
<heading id="D0B581F5FFBE744B81B438B7FD52F9E5" bold="true" box="[151,652,1519,1546]" fontSize="11" level="1" pageId="24" pageNumber="361" reason="1">
<emphasis id="B936EA8BFFBE744B81B438B7FD52F9E5" bold="true" box="[151,652,1519,1546]" pageId="24" pageNumber="361">
<taxonomicName id="4C424D1AFFBE744B81B438B7FDF9F9E5" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,551,1519,1546]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBE744B81B438B7FDF9F9E5" bold="true" box="[151,551,1519,1546]" italics="true" pageId="24" pageNumber="361">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFBE744B830C38A8FD52F9E5" box="[559,652,1520,1546]" pageId="24" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBE744881B43B4CFE4FF92E" lastPageId="27" lastPageNumber="364" pageId="24" pageNumber="361" type="description">
<paragraph id="8BFD3699FFBE744B81B43B4CFEF9F9C0" blockId="24.[151,652,1519,1583]" box="[151,295,1556,1583]" pageId="24" pageNumber="361">
(
<figureCitation id="13792A1CFFBE744B81BC3B4CFEC1F9C0" box="[159,287,1556,1583]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStart-3="FIGURE 28" captionStartId-0="25.[151,250,1592,1616]" captionStartId-1="26.[151,250,1281,1305]" captionStartId-2="27.[151,250,1597,1621]" captionStartId-3="28.[151,250,1196,1220]" captionTargetBox-0="[151,1435,394,1569]" captionTargetBox-1="[151,1435,187,1252]" captionTargetBox-2="[166,1419,221,1542]" captionTargetBox-3="[151,1436,273,1172]" captionTargetId-0="figure-137@25.[151,1435,381,1569]" captionTargetId-1="figure-24@26.[151,1435,181,1257]" captionTargetId-2="figure-17@27.[151,1436,204,1559]" captionTargetId-3="figure-69@28.[151,1436,273,1172]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi-0="http://doi.org/10.5281/zenodo.13618667" figureDoi-1="http://doi.org/10.5281/zenodo.13618669" figureDoi-2="http://doi.org/10.5281/zenodo.13618671" figureDoi-3="http://doi.org/10.5281/zenodo.13618673" httpUri-0="https://zenodo.org/record/13618667/files/figure.png" httpUri-1="https://zenodo.org/record/13618669/files/figure.png" httpUri-2="https://zenodo.org/record/13618671/files/figure.png" httpUri-3="https://zenodo.org/record/13618673/files/figure.png" pageId="24" pageNumber="361">Figs 2528</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFBE744B81B43B04FA80F998" blockId="24.[151,1437,1628,2014]" box="[151,1374,1628,1655]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81B43B04FEFFF999" bold="true" box="[151,289,1628,1654]" pageId="24" pageNumber="361">Description</emphasis>
. FEMALE. Body weakly sclerotized. Length of idiosoma 185 (170190), width 110 (105115).
</paragraph>
<paragraph id="8BFD3699FFBE744B81E43BD8FEE4F89D" blockId="24.[151,1437,1628,2014]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43BD8FE88F975" box="[199,342,1664,1690]" italics="true" pageId="24" pageNumber="361">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 21 (2021), width 21 (2021). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="B936EA8BFFBE744B83DB3BFCFCFEF951" box="[760,800,1700,1726]" italics="true" pageId="24" pageNumber="361">cha</emphasis>
) 6 (6). Dorsal median apodeme absent. Postpalpal setae (
<emphasis id="B936EA8BFFBE744B81823B91FF63F90D" box="[161,189,1737,1762]" italics="true" pageId="24" pageNumber="361">pp</emphasis>
) 2 (2) spiniform, situated laterad setae
<emphasis id="B936EA8BFFBE744B83583B90FD7DF90D" box="[635,675,1736,1762]" italics="true" pageId="24" pageNumber="361">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFBE744B81FB3BB5FF32F8E9" box="[216,236,1773,1798]" italics="true" pageId="24" pageNumber="361">m</emphasis>
8 (89) and round pits
<emphasis id="B936EA8BFFBE744B80DF3BB5FDD4F8E9" box="[508,522,1773,1798]" italics="true" pageId="24" pageNumber="361">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFBE744B83C23BB5FD2BF8E9" box="[737,757,1773,1798]" italics="true" pageId="24" pageNumber="361">m</emphasis>
. Palps with smooth and pointed setae
<emphasis id="B936EA8BFFBE744B858E3BB4FB07F8E9" box="[1197,1241,1772,1798]" italics="true" pageId="24" pageNumber="361">dFe</emphasis>
6 (67) and
<emphasis id="B936EA8BFFBE744B844D3BB4FA43F8E9" box="[1390,1437,1772,1798]" italics="true" pageId="24" pageNumber="361">dGe</emphasis>
12 (1113) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFBE744B842A3A49FAF3F8C5" box="[1289,1325,1809,1834]" italics="true" pageId="24" pageNumber="361">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFBE744B80373A6DFEF6F8A1" box="[276,296,1845,1870]" italics="true" pageId="24" pageNumber="361">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<paragraph id="8BFD3699FFBE744A81E43A24FEE3FE8A" blockId="24.[151,1437,1628,2014]" lastBlockId="25.[151,1436,151,357]" lastPageId="25" lastPageNumber="362" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43A24FE4AF879" box="[199,404,1916,1942]" italics="true" pageId="24" pageNumber="361">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFBE744B80873A24FDCCF878" box="[420,530,1916,1943]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618667" httpUri="https://zenodo.org/record/13618667/files/figure.png" pageId="24" pageNumber="361">Figs 25A</figureCitation>
,
<figureCitation id="13792A1CFFBE744B833C3A24FD8AF879" box="[543,596,1916,1942]" captionStart="FIGURE 28" captionStartId="28.[151,250,1196,1220]" captionTargetBox="[151,1436,273,1172]" captionTargetId="figure-69@28.[151,1436,273,1172]" captionTargetPageId="28" captionText="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618673" httpUri="https://zenodo.org/record/13618673/files/figure.png" pageId="24" pageNumber="361">28A</figureCitation>
). All dorsal shields with large puncta as illustrated in
<figureCitation id="13792A1CFFBE744B859F3A24FAFAF878" box="[1212,1316,1916,1943]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618667" httpUri="https://zenodo.org/record/13618667/files/figure.png" pageId="24" pageNumber="361">Fig. 25A</figureCitation>
. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFBE744B83233AF8FDC4F855" box="[512,538,1952,1978]" italics="true" pageId="24" pageNumber="361">v1</emphasis>
,
<emphasis id="B936EA8BFFBE744B830B3AF8FD9DF855" box="[552,579,1952,1978]" italics="true" pageId="24" pageNumber="361">v2</emphasis>
,
<emphasis id="B936EA8BFFBE744B83723AF8FDA9F855" box="[593,631,1952,1978]" italics="true" pageId="24" pageNumber="361">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFBE744B859F3AF8FB3FF855" box="[1212,1249,1952,1978]" italics="true" pageId="24" pageNumber="361">sc1</emphasis>
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. Setae
<emphasis id="B936EA8BFFBE744B85053A9CFB9EF831" box="[1062,1088,1988,2014]" italics="true" pageId="24" pageNumber="361">v1</emphasis>
,
<emphasis id="B936EA8BFFBE744B85723A9CFBB5F831" box="[1105,1131,1988,2014]" italics="true" pageId="24" pageNumber="361">v2</emphasis>
,
<emphasis id="B936EA8BFFBE744B85583A9CFB7FF831" box="[1147,1185,1988,2014]" italics="true" pageId="24" pageNumber="361">sc2</emphasis>
and
<emphasis id="B936EA8BFFBE744B85F83A9CFB2BF831" box="[1243,1269,1988,2014]" italics="true" pageId="24" pageNumber="361">c2</emphasis>
pointed; other dorsal setae blunt-tipped; setae
<emphasis id="B936EA8BFFBF744A832B3DCFFDCAFF5F" box="[520,532,151,176]" italics="true" pageId="25" pageNumber="362">e</emphasis>
and
<emphasis id="B936EA8BFFBF744A83703DCFFDB1FF5E" box="[595,623,151,177]" italics="true" pageId="25" pageNumber="362">h2</emphasis>
smooth; other dorsal setae barbed. Alveoli of setae
<emphasis id="B936EA8BFFBF744A85F53DCFFB3CFF5F" box="[1238,1250,151,176]" italics="true" pageId="25" pageNumber="362">e</emphasis>
and
<emphasis id="B936EA8BFFBF744A84023DCFFAF7FF5E" box="[1313,1321,151,177]" italics="true" pageId="25" pageNumber="362">f</emphasis>
distinctly separated. Cupules
<emphasis id="B936EA8BFFBF744A80573DE3FE54FF3B" box="[372,394,187,212]" italics="true" pageId="25" pageNumber="362">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFBF744A83093DE3FD98FF3B" box="[554,582,187,212]" italics="true" pageId="25" pageNumber="362">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFBF744A823F3DE3FCECFF3A" box="[796,818,187,213]" italics="true" pageId="25" pageNumber="362">ih</emphasis>
on tergite H large, round, subequal. Lengths of dorsal setae:
<emphasis id="B936EA8BFFBF744A81FC3D87FF27FF16" box="[223,249,223,249]" italics="true" pageId="25" pageNumber="362">v1</emphasis>
29 (2629),
<emphasis id="B936EA8BFFBF744A80AD3D87FE77FF16" box="[398,425,223,249]" italics="true" pageId="25" pageNumber="362">v2</emphasis>
21 (1822),
<emphasis id="B936EA8BFFBF744A831E3D87FDBDFF16" box="[573,611,223,249]" italics="true" pageId="25" pageNumber="362">sc2</emphasis>
43 (4357),
<emphasis id="B936EA8BFFBF744A83D43D87FCCCFF16" box="[759,786,223,249]" italics="true" pageId="25" pageNumber="362">c1</emphasis>
30 (2734),
<emphasis id="B936EA8BFFBF744A82813D87FC63FF16" box="[930,957,223,249]" italics="true" pageId="25" pageNumber="362">c2</emphasis>
43 (4049),
<emphasis id="B936EA8BFFBF744A85723D87FB81FF16" box="[1105,1119,223,249]" italics="true" pageId="25" pageNumber="362">d</emphasis>
32 (3240),
<emphasis id="B936EA8BFFBF744A85D73D87FADEFF17" box="[1268,1280,223,248]" italics="true" pageId="25" pageNumber="362">e</emphasis>
16 (1516),
<emphasis id="B936EA8BFFBF744A84B73D87FA42FF16" box="[1428,1436,223,249]" italics="true" pageId="25" pageNumber="362">f</emphasis>
36 (3444),
<emphasis id="B936EA8BFFBF744A80023C5BFEE3FEF2" box="[289,317,259,285]" italics="true" pageId="25" pageNumber="362">h1</emphasis>
43 (3944),
<emphasis id="B936EA8BFFBF744A80EC3C5BFE35FEF2" box="[463,491,259,285]" italics="true" pageId="25" pageNumber="362">h2</emphasis>
12 (1214). Distances between setae:
<emphasis id="B936EA8BFFBF744A82B83C5BFC00FEF2" box="[923,990,259,285]" italics="true" pageId="25" pageNumber="362">v1v1</emphasis>
20 (2021),
<emphasis id="B936EA8BFFBF744A854C3C5BFB6CFEF2" box="[1135,1202,259,285]" italics="true" pageId="25" pageNumber="362">v2v2</emphasis>
33 (3336),
<emphasis id="B936EA8BFFBF744A84673C5BFA42FEF2" box="[1348,1436,259,285]" italics="true" pageId="25" pageNumber="362">sc2sc2</emphasis>
38 (3738),
<emphasis id="B936EA8BFFBF744A80073C7FFEB9FEAE" box="[292,359,295,321]" italics="true" pageId="25" pageNumber="362">c1c1</emphasis>
31 (3137),
<emphasis id="B936EA8BFFBF744A80D93C7FFDE3FEAE" box="[506,573,295,321]" italics="true" pageId="25" pageNumber="362">c1c2</emphasis>
24 (2224),
<emphasis id="B936EA8BFFBF744A83F23C7FFD25FEAE" box="[721,763,295,321]" italics="true" pageId="25" pageNumber="362">dd</emphasis>
56 (5357),
<emphasis id="B936EA8BFFBF744A82AC3C7FFC6FFEAE" box="[911,945,295,321]" italics="true" pageId="25" pageNumber="362">ef</emphasis>
5 (56),
<emphasis id="B936EA8BFFBF744A85393C7FFBE6FEAE" box="[1050,1080,295,321]" italics="true" pageId="25" pageNumber="362">ff</emphasis>
51 (4752),
<emphasis id="B936EA8BFFBF744A85E83C7FFACFFEAE" box="[1227,1297,295,321]" italics="true" pageId="25" pageNumber="362">h1h1</emphasis>
41 (3742),
<emphasis id="B936EA8BFFBF744A81B43C13FF03FE8A" box="[151,221,331,357]" italics="true" pageId="25" pageNumber="362">h1h2</emphasis>
6 (68).
</paragraph>
<caption id="DF3D6611FFBF744A81B43B60FA85F9BF" ID-DOI="http://doi.org/10.5281/zenodo.13618667" ID-Zenodo-Dep="13618667" box="[151,1371,1592,1617]" httpUri="https://zenodo.org/record/13618667/files/figure.png" pageId="25" pageNumber="362" startId="25.[151,250,1592,1616]" targetBox="[151,1435,394,1569]" targetPageId="25" targetType="figure">
<paragraph id="8BFD3699FFBF744A81B43B60FA85F9BF" blockId="25.[151,1371,1592,1617]" box="[151,1371,1592,1617]" pageId="25" pageNumber="362">
<emphasis id="B936EA8BFFBF744A81B43B60FEC7F9BF" bold="true" box="[151,281,1592,1616]" pageId="25" pageNumber="362">FIGURE 25</emphasis>
.
<taxonomicName id="4C424D1AFFBF744A80053B60FDA9F9BF" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,631,1592,1616]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="25" pageNumber="362" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBF744A80053B60FDA9F9BF" box="[294,631,1592,1616]" italics="true" pageId="25" pageNumber="362">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBF744A835E3B60FD10F9BF" bold="true" box="[637,718,1592,1616]" pageId="25" pageNumber="362">
<taxonomicNameLabel id="A20557F0FFBF744A835E3B60FD10F9BF" box="[637,718,1592,1616]" pageId="25" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBF744A81E43B26FDBBF833" blockId="25.[151,1437,1662,2013]" pageId="25" pageNumber="362">
<emphasis id="B936EA8BFFBF744A81E43B26FE56F977" box="[199,392,1662,1688]" italics="true" pageId="25" pageNumber="362">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFBF744A80B93B26FDD9F976" box="[410,519,1662,1689]" captionStart="FIGURE 25" captionStartId="25.[151,250,1592,1616]" captionTargetBox="[151,1435,394,1569]" captionTargetId="figure-137@25.[151,1435,381,1569]" captionTargetPageId="25" captionText="FIGURE 25. Spatulaphorus madagascariensis sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618667" httpUri="https://zenodo.org/record/13618667/files/figure.png" pageId="25" pageNumber="362">Figs 25B</figureCitation>
,
<figureCitation id="13792A1CFFBF744A83353B27FD97F976" box="[534,585,1663,1689]" captionStart="FIGURE 28" captionStartId="28.[151,250,1196,1220]" captionTargetBox="[151,1436,273,1172]" captionTargetId="figure-69@28.[151,1436,273,1172]" captionTargetPageId="28" captionText="FIGURE 28. DIC micrographs of Spatulaphorus madagascariensis sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618673" httpUri="https://zenodo.org/record/13618673/files/figure.png" pageId="25" pageNumber="362">28B</figureCitation>
). Anterior and posterior sternal plates with very small, hardly discernible puncta; aggenital and pseudanal plates with slightly larger puncta. Setae
<emphasis id="B936EA8BFFBF744A82E83BFAFC36F953" box="[971,1000,1698,1724]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
,
<emphasis id="B936EA8BFFBF744A82D53BFBFBCEF952" box="[1014,1040,1699,1725]" italics="true" pageId="25" pageNumber="362">2c</emphasis>
,
<emphasis id="B936EA8BFFBF744A853C3BFAFBE5F953" box="[1055,1083,1698,1724]" italics="true" pageId="25" pageNumber="362">4b</emphasis>
, and
<emphasis id="B936EA8BFFBF744A85593BFBFB66F952" box="[1146,1208,1699,1725]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
pointed; other setae weakly blunt-tipped; setae
<emphasis id="B936EA8BFFBF744A80E23B9EFE03F90F" box="[449,477,1734,1760]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
weakly barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFBF744A85433B9FFB59F90E" box="[1120,1159,1735,1761]" italics="true" pageId="25" pageNumber="362">ps3</emphasis>
situated slightly anteriad bases of setae
<emphasis id="B936EA8BFFBF744A80153BB3FEAAF8EA" box="[310,372,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
; setae
<emphasis id="B936EA8BFFBF744A80E23BB3FE39F8EA" box="[449,487,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1</emphasis>
usually shorter than
<emphasis id="B936EA8BFFBF744A83EC3BB3FD28F8EA" box="[719,758,1771,1797]" italics="true" pageId="25" pageNumber="362">ps2</emphasis>
; setae
<emphasis id="B936EA8BFFBF744A82603BB3FC5FF8EA" box="[835,897,1771,1797]" italics="true" pageId="25" pageNumber="362">ps1-2</emphasis>
subequal in some specimens. Ap1 and ap2 well-developed and joined with thick appr; apsej absent; ap3 very thin, hardly discernible; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFBF744A80B23A6AFE73F8A3" box="[401,429,1842,1868]" italics="true" pageId="25" pageNumber="362">3b</emphasis>
; ap5 well developed and exceeding bases of trochanter IV. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate with short tongue-like elongation. Lengths of ventral setae:
<emphasis id="B936EA8BFFBF744A82143A23FC8AF87A" box="[823,852,1915,1941]" italics="true" pageId="25" pageNumber="362">1a</emphasis>
10 (911),
<emphasis id="B936EA8BFFBF744A82F53A22FC2DF87B" box="[982,1011,1914,1940]" italics="true" pageId="25" pageNumber="362">1b</emphasis>
19 (1719),
<emphasis id="B936EA8BFFBF744A85A73A23FB40F87A" box="[1156,1182,1915,1941]" italics="true" pageId="25" pageNumber="362">1с</emphasis>
12 (912),
<emphasis id="B936EA8BFFBF744A84013A23FAE0F87A" box="[1314,1342,1915,1941]" italics="true" pageId="25" pageNumber="362">2a</emphasis>
11 (11),
<emphasis id="B936EA8BFFBF744A81B43AC7FF6FF856" box="[151,177,1951,1977]" italics="true" pageId="25" pageNumber="362">2с</emphasis>
13 (1315),
<emphasis id="B936EA8BFFBF744A80673AC7FEBEF856" box="[324,352,1951,1977]" italics="true" pageId="25" pageNumber="362">3a</emphasis>
13 (1213),
<emphasis id="B936EA8BFFBF744A80D13AC6FDD0F857" box="[498,526,1950,1976]" italics="true" pageId="25" pageNumber="362">3b</emphasis>
11 (1011),
<emphasis id="B936EA8BFFBF744A83BD3AC7FD66F856" box="[670,696,1951,1977]" italics="true" pageId="25" pageNumber="362">3c</emphasis>
12 (1213),
<emphasis id="B936EA8BFFBF744A82693AC7FCB9F856" box="[842,871,1951,1977]" italics="true" pageId="25" pageNumber="362">4a</emphasis>
10 (1011),
<emphasis id="B936EA8BFFBF744A82D43AC6FBCAF857" box="[1015,1044,1950,1976]" italics="true" pageId="25" pageNumber="362">4b</emphasis>
16 (1520),
<emphasis id="B936EA8BFFBF744A85863AC7FB1EF856" box="[1189,1216,1951,1977]" italics="true" pageId="25" pageNumber="362">4c</emphasis>
14 (1215),
<emphasis id="B936EA8BFFBF744A84713AC7FAA7F856" box="[1362,1401,1951,1977]" italics="true" pageId="25" pageNumber="362">ps1</emphasis>
12 (1112),
<emphasis id="B936EA8BFFBF744A81DE3A9BFEFAF832" box="[253,292,1987,2013]" italics="true" pageId="25" pageNumber="362">ps2</emphasis>
15 (1315),
<emphasis id="B936EA8BFFBF744A80973A9BFE05F832" box="[436,475,1987,2013]" italics="true" pageId="25" pageNumber="362">ps3</emphasis>
16 (1617).
</paragraph>
<caption id="DF3D6611FFBC744981B43859FA59FAF6" ID-DOI="http://doi.org/10.5281/zenodo.13618669" ID-Zenodo-Dep="13618669" box="[151,1415,1281,1305]" httpUri="https://zenodo.org/record/13618669/files/figure.png" pageId="26" pageNumber="363" startId="26.[151,250,1281,1305]" targetBox="[151,1435,187,1252]" targetPageId="26" targetType="figure">
<paragraph id="8BFD3699FFBC744981B43859FA59FAF6" blockId="26.[151,1415,1281,1305]" box="[151,1415,1281,1305]" pageId="26" pageNumber="363">
<emphasis id="B936EA8BFFBC744981B43859FEC7FAF6" bold="true" box="[151,281,1281,1305]" pageId="26" pageNumber="363">FIGURE 26</emphasis>
.
<taxonomicName id="4C424D1AFFBC744980053859FDA9FAF6" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,631,1281,1305]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="363" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBC744980053859FDA9FAF6" box="[294,631,1281,1305]" italics="true" pageId="26" pageNumber="363">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBC7449835E3859FD10FAF6" bold="true" box="[637,718,1281,1305]" pageId="26" pageNumber="363">
<taxonomicNameLabel id="A20557F0FFBC7449835E3859FD10FAF6" box="[637,718,1281,1305]" pageId="26" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBC744981E43810FE05F84E" blockId="26.[151,1437,1351,1953]" pageId="26" pageNumber="363">
<emphasis id="B936EA8BFFBC744981E43810FF22FA8E" box="[199,252,1352,1377]" italics="true" pageId="26" pageNumber="363">Legs</emphasis>
(
<figureCitation id="13792A1CFFBC74498028381FFEBCFA8E" box="[267,354,1351,1377]" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618669" httpUri="https://zenodo.org/record/13618669/files/figure.png" pageId="26" pageNumber="363">Figs 26</figureCitation>
,
<figureCitation id="13792A1CFFBC7449804D3810FE50FA8D" box="[366,398,1352,1378]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618671" httpUri="https://zenodo.org/record/13618671/files/figure.png" pageId="26" pageNumber="363">27</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFBC744983B23810FCE8FA8E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[657,822,1351,1378]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="363" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFBC744983B23810FCE8FA8E" box="[657,822,1351,1378]" italics="true" pageId="26" pageNumber="363">S. brevisetosus</emphasis>
</taxonomicName>
except presence of solenidion
<emphasis id="B936EA8BFFBC744985B23810FB7EFA8E" box="[1169,1184,1352,1377]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
on tibia II. Leg I (
<figureCitation id="13792A1CFFBC7449844C381FFF12FA6A" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618669" httpUri="https://zenodo.org/record/13618669/files/figure.png" pageId="26" pageNumber="363">Fig. 26A</figureCitation>
). Tibiotarsus enlarged, with large, thick and strongly curved claw; tibiotarsus internally without oval strongly sclerotized structure. Setae (
<emphasis id="B936EA8BFFBC744980F038C8FE3FFA46" box="[467,481,1424,1449]" italics="true" pageId="26" pageNumber="363">u</emphasis>
) consolidated into a bifid horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFBC744981ED38EBFF02FA22" box="[206,220,1459,1485]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFBC7449825C38EBFC4EFA22" box="[895,912,1459,1485]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of femur and
<emphasis id="B936EA8BFFBC7449850D38EBFBE4FA22" box="[1070,1082,1459,1485]" italics="true" pageId="26" pageNumber="363">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFBC7449808E3880FE1AFA1D" box="[429,452,1496,1522]" italics="true" pageId="26" pageNumber="363">p</emphasis>
,
<emphasis id="B936EA8BFFBC744980F03880FE2FFA1D" box="[467,497,1496,1522]" italics="true" pageId="26" pageNumber="363">p”</emphasis>
,
<emphasis id="B936EA8BFFBC744980DC3880FDC3FA1D" box="[511,541,1496,1522]" italics="true" pageId="26" pageNumber="363">tc</emphasis>
,
<emphasis id="B936EA8BFFBC744983083880FD91FA1D" box="[555,591,1496,1522]" italics="true" pageId="26" pageNumber="363">tc”</emphasis>
,
<emphasis id="B936EA8BFFBC7449837D388FFDA9FA1E" box="[606,631,1495,1521]" italics="true" pageId="26" pageNumber="363">ft</emphasis>
,
<emphasis id="B936EA8BFFBC744983A6388FFD7AFA1E" box="[645,676,1495,1521]" italics="true" pageId="26" pageNumber="363">ft”</emphasis>
) barbed. Setae
<emphasis id="B936EA8BFFBC74498276388FFCBDFA1E" box="[853,867,1495,1521]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of femur and
<emphasis id="B936EA8BFFBC74498527388FFBCEFA1E" box="[1028,1040,1495,1521]" italics="true" pageId="26" pageNumber="363">k</emphasis>
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFBC7449824D38A4FC4EF9F9" box="[878,912,1532,1558]" italics="true" pageId="26" pageNumber="363">ω1</emphasis>
8 (8),
<emphasis id="B936EA8BFFBC744982F838A4FC23F9F9" box="[987,1021,1532,1558]" italics="true" pageId="26" pageNumber="363">ω2</emphasis>
5 (5),
<emphasis id="B936EA8BFFBC7449856A38A4FBB8F9F9" box="[1097,1126,1532,1558]" italics="true" pageId="26" pageNumber="363">φ1</emphasis>
8 (8),
<emphasis id="B936EA8BFFBC7449859238A4FB11F9F9" box="[1201,1231,1532,1558]" italics="true" pageId="26" pageNumber="363">φ2</emphasis>
6 (56); solenidia
<emphasis id="B936EA8BFFBC744981B43B78FF67F9D5" box="[151,185,1568,1594]" italics="true" pageId="26" pageNumber="363">ω1</emphasis>
and
<emphasis id="B936EA8BFFBC744981D33B78FED3F9D5" box="[240,269,1568,1594]" italics="true" pageId="26" pageNumber="363">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFBC744982503B47FC04F9D6" box="[883,986,1567,1593]" captionStart="FIGURE 26" captionStartId="26.[151,250,1281,1305]" captionTargetBox="[151,1435,187,1252]" captionTargetId="figure-24@26.[151,1435,181,1257]" captionTargetPageId="26" captionText="FIGURE 26. Spatulaphorus madagascariensis sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618669" httpUri="https://zenodo.org/record/13618669/files/figure.png" pageId="26" pageNumber="363">Fig. 26B</figureCitation>
). Tarsus with simple claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFBC744980BC3B1CFE6DF9B2" box="[415,435,1604,1629]" italics="true" pageId="26" pageNumber="363">ω</emphasis>
7 (7) weakly clavate, solenidion
<emphasis id="B936EA8BFFBC7449820F3B1CFCE5F9B2" box="[812,827,1604,1629]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
2 (2) very short. Seta
<emphasis id="B936EA8BFFBC744985193B1CFB8FF9B1" box="[1082,1105,1604,1630]" italics="true" pageId="26" pageNumber="363">u</emphasis>
of tarsus smooth; other setae leg setae barbed; seta
<emphasis id="B936EA8BFFBC744980B03B3FFE7FF96E" box="[403,417,1639,1665]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur blunt-tipped; tips of setae (
<emphasis id="B936EA8BFFBC7449826A3B30FCBDF96E" box="[841,867,1640,1665]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly foliate; other leg setae pointed. Leg III (
<figureCitation id="13792A1CFFBC744981DA3BD3FEBAF949" box="[249,356,1675,1702]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618671" httpUri="https://zenodo.org/record/13618671/files/figure.png" pageId="26" pageNumber="363">Fig. 27A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Seta
<emphasis id="B936EA8BFFBC744981EC3BE8FF38F925" box="[207,230,1712,1738]" italics="true" pageId="26" pageNumber="363">u</emphasis>
of tarsus smooth; other leg setae barbed; setae
<emphasis id="B936EA8BFFBC744982223BF7FCD1F926" box="[769,783,1711,1737]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur and
<emphasis id="B936EA8BFFBC744982923BF7FC1CF926" box="[945,962,1711,1737]" italics="true" pageId="26" pageNumber="363">l</emphasis>
of genu blunt-tipped; tips of setae (
<emphasis id="B936EA8BFFBC7449847A3BE8FAADF926" box="[1369,1395,1712,1737]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly flattened; other leg setae pointed; external part of solenidion
<emphasis id="B936EA8BFFBC744982D43B8CFBD8F902" box="[1015,1030,1748,1773]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion. Leg IV (
<figureCitation id="13792A1CFFBC7449852E3BAFFBB0F8FD" box="[1037,1134,1783,1810]" captionStart="FIGURE 27" captionStartId="27.[151,250,1597,1621]" captionTargetBox="[166,1419,221,1542]" captionTargetId="figure-17@27.[151,1436,204,1559]" captionTargetPageId="27" captionText="FIGURE 27. Spatulaphorus madagascariensis sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618671" httpUri="https://zenodo.org/record/13618671/files/figure.png" pageId="26" pageNumber="363">Fig 27B</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium as on tarsi II and III. Seta
<emphasis id="B936EA8BFFBC744982793A43FC5EF8DA" box="[858,896,1819,1845]" italics="true" pageId="26" pageNumber="363">pl”</emphasis>
of tarsus smooth; other leg setae barbed; setae
<emphasis id="B936EA8BFFBC744984AD3A43FA42F8DA" box="[1422,1436,1819,1845]" italics="true" pageId="26" pageNumber="363">d</emphasis>
of femur,
<emphasis id="B936EA8BFFBC744980253A18FEFCF8B5" box="[262,290,1856,1882]" italics="true" pageId="26" pageNumber="363">v”</emphasis>
of tibia,
<emphasis id="B936EA8BFFBC744980AA3A67FE71F8B6" box="[393,431,1855,1881]" italics="true" pageId="26" pageNumber="363">pl”</emphasis>
and
<emphasis id="B936EA8BFFBC744980C53A18FDD4F8B5" box="[486,522,1856,1882]" italics="true" pageId="26" pageNumber="363">tc”</emphasis>
of tarsus blunt-tipped; setae (
<emphasis id="B936EA8BFFBC744982793A18FCAAF8B6" box="[858,884,1856,1881]" italics="true" pageId="26" pageNumber="363">pv</emphasis>
) of tarsus slightly foliate distally; other leg setae pointed; external part of solenidion
<emphasis id="B936EA8BFFBC744983123A3CFD9EF892" box="[561,576,1892,1917]" italics="true" pageId="26" pageNumber="363">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion.
</paragraph>
<caption id="DF3D6611FFBD744881B43B65FF00F996" ID-DOI="http://doi.org/10.5281/zenodo.13618671" ID-Zenodo-Dep="13618671" httpUri="https://zenodo.org/record/13618671/files/figure.png" pageId="27" pageNumber="364" startId="27.[151,250,1597,1621]" targetBox="[166,1419,221,1542]" targetPageId="27" targetType="figure">
<paragraph id="8BFD3699FFBD744881B43B65FF00F996" blockId="27.[151,1436,1597,1657]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881B43B65FEC0F9BA" bold="true" box="[151,286,1597,1621]" pageId="27" pageNumber="364">FIGURE 27</emphasis>
.
<taxonomicName id="4C424D1AFFBD744880133B65FD58F9BA" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[304,646,1597,1621]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBD744880133B65FD58F9BA" box="[304,646,1597,1621]" italics="true" pageId="27" pageNumber="364">Spatulaphorus madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBD744883B23B65FD39F9BA" bold="true" box="[657,743,1597,1621]" pageId="27" pageNumber="364">
<taxonomicNameLabel id="A20557F0FFBD744883B23B65FD39F9BA" box="[657,743,1597,1621]" pageId="27" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBD744881EB3BF0FE4FF92E" blockId="27.[151,1437,1703,2018]" box="[200,401,1703,1730]" pageId="27" pageNumber="364">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBD744881E43B94FD9BF89A" pageId="27" pageNumber="364" type="materials_examined">
<paragraph id="8BFD3699FFBD744881E43B94FB17F8E6" blockId="27.[151,1437,1703,2018]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43B94FEACF90A" bold="true" box="[199,370,1739,1766]" pageId="27" pageNumber="364">Type material</emphasis>
. Female
<materialsCitation id="3B2A3CC4FFBD744880C23B93FC9BF8E6" collectionCode="ZISP, ZIN" country="Madagascar" location="Adringitra Range" municipality="Andohariana" pageId="27" pageNumber="364" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F9883BFFBD744880C23B93FD98F90A" box="[481,582,1739,1765]" pageId="27" pageNumber="364" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFBD744883B73B94FD13F909" box="[660,717,1740,1766]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="27" pageNumber="364">ZISP</collectionCode>
T-Pygm-016,
<collectingCountry id="F3557609FFBD744882563B93FC20F90A" box="[885,1022,1739,1765]" name="Madagascar" pageId="27" pageNumber="364">Madagascar</collectingCountry>
,
<location id="8E9D6042FFBD744885283B93FB0BF909" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744885283B93FB0BF909" box="[1035,1237,1739,1766]" country="Madagascar" municipality="Andohariana" name="Adringitra Range" pageId="27" pageNumber="364">Adringitra Range</location>
,
<collectingMunicipality id="6B99ACE3FFBD744885C03B93FAA9F90A" box="[1251,1399,1739,1765]" pageId="27" pageNumber="364">Andohariana</collectingMunicipality>
, 9- XI/
<date id="FFFC1059FFBD744881E33BA8FE96F8E5" box="[192,328,1776,1802]" pageId="27" pageNumber="364" value="1979-12-10">10-XII.1979</date>
(
<collectionCode id="ED53AE5CFFBD744880743BA8FE55F8E5" box="[343,395,1776,1802]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="364" type="Museum">ZIN</collectionCode>
),
<collectedFrom id="385E7564FFBD744880BD3BA8FC9BF8E6" box="[414,837,1775,1802]" pageId="27" pageNumber="364">
on
<taxonomicName id="4C424D1AFFBD744880E23BB7FC9BF8E6" authority="Orbigny" authorityName="Orbigny" authorityYear="1915" box="[449,837,1775,1801]" class="Insecta" family="Scarabaeidae" genus="Helictopleurus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="corruscus">
<emphasis id="B936EA8BFFBD744880E23BB7FD02F8E6" box="[449,732,1775,1801]" italics="true" pageId="27" pageNumber="364">Helictopleurus corruscus</emphasis>
Orbigny
</taxonomicName>
</collectedFrom>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBD744882733BA8FB1BF8E6" box="[848,1221,1775,1802]" collectionCode="Male" country="Madagascar" location="Adringitra Range" municipality="Andohariana" pageId="27" pageNumber="364" specimenCount="75" specimenCount-female="75" typeStatus="paratype">
<typeStatus id="54F9883BFFBD744882733BA8FC1EF8E5" box="[848,960,1776,1802]" pageId="27" pageNumber="364" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D44FD10FFBD744882E93BB7FB99F8E6" box="[970,1095,1775,1802]" count="75" pageId="27" pageNumber="364" type="female">75 females</specimenCount>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD3699FFBD744881E43A4CFD9BF89A" blockId="27.[151,1437,1703,2018]" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43A4CFE58F8C2" bold="true" box="[199,390,1811,1838]" pageId="27" pageNumber="364">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFBD744880B53A4BFE25F8BE" country="Russia" location="Zoological Institute" pageId="27" pageNumber="364" specimenCount="9" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFBD744880EF3A4BFDF0F8C2" box="[460,558,1811,1837]" pageId="27" pageNumber="364" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFBD7448834A3A4BFCD0F8C1" box="[617,782,1811,1838]" count="9" pageId="27" pageNumber="364" type="generic" typeStatus="paratype">nine paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFBD744885BE3A4BFAA5F8C2" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744885BE3A4BFAA5F8C2" box="[1181,1403,1811,1837]" country="Russia" name="Zoological Institute" pageId="27" pageNumber="364" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFBD744881FC3A6FFE40F8BE" box="[223,414,1847,1873]" country="Russia" name="Saint Petersburg" pageId="27" pageNumber="364">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFBD7448808E3A6FFE25F8BE" box="[429,507,1847,1873]" name="Russia" pageId="27" pageNumber="364">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBD744883293A6FFAC5F8BD" box="[522,1307,1847,1874]" country="Russia" location="Tyumen State" pageId="27" pageNumber="364" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFBD7448836D3A60FD67F8BD" box="[590,697,1848,1874]" pageId="27" pageNumber="364" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFBD7448855E3A60FAC5F8BD" box="[1149,1307,1848,1874]" country="Russia" name="Tyumen'" pageId="27" pageNumber="364">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFBD744884053A6FFD9EF89A" country="Russia" location="University Museum of Zoology" pageId="27" pageNumber="364" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFBD744884053A6FFEA1F89A" LSID="urn:lsid:plazi:treatment:03EB878FFFBE744F81B438B7FB10FF16:8E9D6042FFBD744884053A6FFEA1F89A" country="Russia" name="University Museum of Zoology" pageId="27" pageNumber="364" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFBD744880AA3A04FE39F899" box="[393,487,1884,1910]" country="Russia" name="Tyumen'" pageId="27" pageNumber="364">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFBD744880D13A03FD9EF89A" box="[498,576,1883,1909]" name="Russia" pageId="27" pageNumber="364">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBD744F81E43A27FC77FF3A" lastPageId="28" lastPageNumber="365" pageId="27" pageNumber="364" type="diagnosis">
<paragraph id="8BFD3699FFBD744F81E43A27FC77FF3A" blockId="27.[151,1437,1703,2018]" lastBlockId="28.[151,1436,151,249]" lastPageId="28" lastPageNumber="365" pageId="27" pageNumber="364">
<emphasis id="B936EA8BFFBD744881E43A27FE18F876" bold="true" box="[199,454,1919,1945]" pageId="27" pageNumber="364">Differential diagnosis</emphasis>
. The new species is most similar to
<taxonomicName id="4C424D1AFFBD7448827D3AD8FC16F876" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[862,968,1920,1946]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD7448827D3AD8FC16F876" box="[862,968,1920,1946]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBD744882EC3AD8FBF6F875" bold="true" box="[975,1064,1920,1946]" pageId="27" pageNumber="364">sp, nov.</emphasis>
in having an enlarged tibiotarsus I, simple claws on tarsi II and III and presence of solenidion
<emphasis id="B936EA8BFFBD744882183AFCFC94F852" box="[827,842,1956,1981]" italics="true" pageId="27" pageNumber="364">φ</emphasis>
on tibia II. The new species differs from
<taxonomicName id="4C424D1AFFBD744884353AFCFA5EF852" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1302,1408,1956,1982]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD744884353AFCFA5EF852" box="[1302,1408,1956,1982]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
in having well developed ap5 exceeding trochanters IV (vs. ap5 very short and thin in
<taxonomicName id="4C424D1AFFBD744885613A90FB72F80E" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1090,1196,1992,2018]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="364" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBD744885613A90FB72F80E" box="[1090,1196,1992,2018]" italics="true" pageId="27" pageNumber="364">S. incisus</emphasis>
</taxonomicName>
); posterior margin of posterior sternal plate almost straight medially (vs. posterior margin of posterior sternal plate deeply incised in
<taxonomicName id="4C424D1AFFBA744F84A43DCFFF3BFF3B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBA744F84A43DCFFF3BFF3B" italics="true" pageId="28" pageNumber="365">S. incisus</emphasis>
</taxonomicName>
); and setae
<emphasis id="B936EA8BFFBA744F80483DE3FE77FF3A" box="[363,425,187,213]" italics="true" pageId="28" pageNumber="365">ps1-3</emphasis>
smooth (vs. setae
<emphasis id="B936EA8BFFBA744F835A3DE3FD69FF3A" box="[633,695,187,213]" italics="true" pageId="28" pageNumber="365">ps1-3</emphasis>
barbed in
<taxonomicName id="4C424D1AFFBA744F820D3DE3FC46FF3B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[814,920,187,213]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B936EA8BFFBA744F820D3DE3FC46FF3B" box="[814,920,187,213]" italics="true" pageId="28" pageNumber="365">S. incisus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBA744F81E43D87FB10FF16" box="[199,1230,223,249]" pageId="28" pageNumber="365" type="etymology">
<paragraph id="8BFD3699FFBA744F81E43D87FB10FF16" blockId="28.[151,1436,151,249]" box="[199,1230,223,249]" pageId="28" pageNumber="365">
<emphasis id="B936EA8BFFBA744F81E43D87FE96FF16" bold="true" box="[199,328,223,249]" pageId="28" pageNumber="365">Etymology</emphasis>
. The name of the new species refers to geographical distribution in
<collectingCountry id="F3557609FFBA744F85623D87FB14FF16" box="[1089,1226,223,249]" name="Madagascar" pageId="28" pageNumber="365">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>