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<document id="9022EF7AAB69F15E4B3110C20F6000D9" ID-DOI="10.11646/zootaxa.5497.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618611" ID-ZooBank="C107D617-5F70-42C5-8FD4-E5C2DE0E48B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725026357643" checkinUser="plazi" docAuthor="Khaustov, Alexander A., Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2024" docId="03EB878FFFB2744B81B43BF9FE00FA71" docLanguage="en" docName="zootaxa.5497.3.2.pdf" docOrigin="Zootaxa 5497 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus reductus Khaustov, Frolov &amp; Akhmetova, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="361" masterDocId="FFD2FFF7FFA6745381233D58FFDEFFEF" masterDocTitle="Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)" masterLastPageNumber="368" masterPageNumber="337" pageNumber="357" updateTime="1725028510941" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5948F8DCAC21C4A002AEE587321C9160">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="7809EBE2418489D17B7170505822A516">Khaustov, Alexander A.</mods:namePart>
<mods:affiliation id="B360709791049F6B329B25C2B560FF9A">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
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<mods:namePart id="75AD694EA85248344866E4F6C6AFBA4C">Frolov, Andrey V.</mods:namePart>
<mods:affiliation id="A7173937A54DB90A62B8B9374D291F93">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="94906D770780F9C94FEBC753017B6090">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="033456354AF4594A5D028873483EEA85">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:date id="67C90646C8E1E9A224C7F9CD910ECCCB">2024</mods:date>
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<mods:number id="9A6A3A580C4014A5CD466D043E0D9231">2024-08-26</mods:number>
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<treatment id="03EB878FFFB2744B81B43BF9FE00FA71" ID-DOI="http://doi.org/10.5281/zenodo.13618858" ID-Zenodo-Dep="13618858" LSID="urn:lsid:plazi:treatment:03EB878FFFB2744B81B43BF9FE00FA71" httpUri="http://treatment.plazi.org/id/03EB878FFFB2744B81B43BF9FE00FA71" lastPageId="24" lastPageNumber="361" pageId="20" pageNumber="357">
<subSubSection id="C3586512FFB2744781B43BF9FDC3F953" box="[151,541,1697,1724]" pageId="20" pageNumber="357" type="nomenclature">
<paragraph id="8BFD3699FFB2744781B43BF9FDC3F953" blockId="20.[151,541,1697,1760]" box="[151,541,1697,1724]" pageId="20" pageNumber="357">
<heading id="D0B581F5FFB2744781B43BF9FDC3F953" bold="true" box="[151,541,1697,1724]" fontSize="11" level="1" pageId="20" pageNumber="357" reason="1">
<emphasis id="B936EA8BFFB2744781B43BF9FDC3F953" bold="true" box="[151,541,1697,1724]" pageId="20" pageNumber="357">
<taxonomicName id="4C424D1AFFB2744781B43BF9FE66F953" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[151,440,1697,1724]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB2744781B43BF9FE66F953" bold="true" box="[151,440,1697,1724]" italics="true" pageId="20" pageNumber="357">Spatulaphorus reductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557F0FFB2744780E33BFAFDC3F953" box="[448,541,1698,1724]" pageId="20" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB2744481B43B9EFE4FF8C3" lastPageId="23" lastPageNumber="360" pageId="20" pageNumber="357" type="description">
<paragraph id="8BFD3699FFB2744781B43B9EFEF9F90F" blockId="20.[151,541,1697,1760]" box="[151,295,1734,1760]" pageId="20" pageNumber="357">
(
<figureCitation id="13792A1CFFB2744781BC3B9EFEC1F90F" box="[159,287,1734,1760]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStart-3="FIGURE 24" captionStartId-0="21.[151,250,1357,1381]" captionStartId-1="22.[151,250,1442,1466]" captionStartId-2="23.[151,250,1740,1764]" captionStartId-3="24.[151,247,1089,1113]" captionTargetBox-0="[151,1435,245,1333]" captionTargetBox-1="[151,1417,185,1419]" captionTargetBox-2="[160,1427,190,1716]" captionTargetBox-3="[151,1436,181,1066]" captionTargetId-0="figure-41@21.[151,1435,237,1333]" captionTargetId-1="figure-24@22.[151,1436,181,1419]" captionTargetId-2="figure-17@23.[151,1436,181,1716]" captionTargetId-3="figure-24@24.[151,1436,181,1066]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." captionText-2="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." figureDoi-0="http://doi.org/10.5281/zenodo.13618659" figureDoi-1="http://doi.org/10.5281/zenodo.13618661" figureDoi-2="http://doi.org/10.5281/zenodo.13618663" figureDoi-3="http://doi.org/10.5281/zenodo.13618665" httpUri-0="https://zenodo.org/record/13618659/files/figure.png" httpUri-1="https://zenodo.org/record/13618661/files/figure.png" httpUri-2="https://zenodo.org/record/13618663/files/figure.png" httpUri-3="https://zenodo.org/record/13618665/files/figure.png" pageId="20" pageNumber="357">Figs 2124</figureCitation>
)
</paragraph>
<paragraph id="8BFD3699FFB2744781B43A55FA81F8C7" blockId="20.[151,1437,1805,2012]" box="[151,1375,1805,1832]" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781B43A55FEFFF8C8" bold="true" box="[151,289,1805,1831]" pageId="20" pageNumber="357">Description</emphasis>
. FEMALE. Body weakly sclerotized. Length of idiosoma 155 (150175), width 105 (100110).
</paragraph>
<paragraph id="8BFD3699FFB2744681E43A6AFD94FF3A" blockId="20.[151,1437,1805,2012]" lastBlockId="21.[151,1436,151,213]" lastPageId="21" lastPageNumber="358" pageId="20" pageNumber="357">
<emphasis id="B936EA8BFFB2744781E43A6AFE88F8A3" box="[199,342,1842,1868]" italics="true" pageId="20" pageNumber="357">Gnathosoma</emphasis>
. Gnathosomal capsule, excluding palps, subrectangular, length 19 (1921), width 23 (2023). Dorsum with one pair of pointed cheliceral setae (
<emphasis id="B936EA8BFFB2744783DF3A0EFCFAF89F" box="[764,804,1878,1904]" italics="true" pageId="20" pageNumber="357">cha</emphasis>
) 10 (610). Dorsal median apodeme present, poorly developed. Postpalpal setae (
<emphasis id="B936EA8BFFB2744780FC3A22FE25F87C" box="[479,507,1914,1939]" italics="true" pageId="20" pageNumber="357">pp</emphasis>
) 4 (5) baculiform, situated laterad setae
<emphasis id="B936EA8BFFB2744782EB3A22FC2EF87B" box="[968,1008,1914,1940]" italics="true" pageId="20" pageNumber="357">cha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="B936EA8BFFB2744783013AC6FDE8F858" box="[546,566,1950,1975]" italics="true" pageId="20" pageNumber="357">m</emphasis>
8 (89) and round pits
<emphasis id="B936EA8BFFB27447821D3AC6FC92F858" box="[830,844,1950,1975]" italics="true" pageId="20" pageNumber="357">n</emphasis>
situated posteriad
<emphasis id="B936EA8BFFB2744785033AC6FBEAF858" box="[1056,1076,1950,1975]" italics="true" pageId="20" pageNumber="357">m</emphasis>
. Palps with smooth and pointed setae
<emphasis id="B936EA8BFFB2744781F43A9AFEDDF833" box="[215,259,1986,2012]" italics="true" pageId="20" pageNumber="357">dFe</emphasis>
7 (67) and
<emphasis id="B936EA8BFFB2744780B63A9AFE1AF833" box="[405,452,1986,2012]" italics="true" pageId="20" pageNumber="357">dGe</emphasis>
8 (813) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
<emphasis id="B936EA8BFFB3744680283DCFFEF1FF5F" box="[267,303,151,176]" italics="true" pageId="21" pageNumber="358">ass</emphasis>
) and tiny solenidion
<emphasis id="B936EA8BFFB3744683063DCFFDE7FF5F" box="[549,569,151,176]" italics="true" pageId="21" pageNumber="358">ω</emphasis>
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
</paragraph>
<caption id="DF3D6611FFB3744681B43815FB24FA8A" ID-DOI="http://doi.org/10.5281/zenodo.13618659" ID-Zenodo-Dep="13618659" box="[151,1274,1357,1381]" httpUri="https://zenodo.org/record/13618659/files/figure.png" pageId="21" pageNumber="358" startId="21.[151,250,1357,1381]" targetBox="[151,1435,245,1333]" targetPageId="21" targetType="figure">
<paragraph id="8BFD3699FFB3744681B43815FB24FA8A" blockId="21.[151,1274,1357,1381]" box="[151,1274,1357,1381]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681B43815FEC7FA8A" bold="true" box="[151,281,1357,1381]" pageId="21" pageNumber="358">FIGURE 21</emphasis>
.
<taxonomicName id="4C424D1AFFB3744680053815FDC8FA8A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1357,1381]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="358" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB3744680053815FDC8FA8A" box="[294,534,1357,1381]" italics="true" pageId="21" pageNumber="358">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB37446833E3815FDB3FA8B" bold="true" box="[541,621,1357,1380]" pageId="21" pageNumber="358">
<taxonomicNameLabel id="A20557F0FFB37446833E3815FDB3FA8B" box="[541,621,1357,1380]" pageId="21" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—dorsum of body; B—venter of body. Legs omitted.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB3744681E438CBFBB2F945" blockId="21.[151,1437,1427,1958]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681E438CBFE4AFA42" box="[199,404,1427,1453]" italics="true" pageId="21" pageNumber="358">Idiosomal dorsum</emphasis>
(
<figureCitation id="13792A1CFFB37446808738CBFDCCFA42" box="[420,530,1427,1453]" captionStart="FIGURE 21" captionStartId="21.[151,250,1357,1381]" captionTargetBox="[151,1435,245,1333]" captionTargetId="figure-41@21.[151,1435,237,1333]" captionTargetPageId="21" captionText="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618659" httpUri="https://zenodo.org/record/13618659/files/figure.png" pageId="21" pageNumber="358">Figs 21A</figureCitation>
,
<figureCitation id="13792A1CFFB37446833D38CBFD8DFA42" box="[542,595,1427,1453]" captionStart="FIGURE 24" captionStartId="24.[151,247,1089,1113]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-24@24.[151,1436,181,1066]" captionTargetPageId="24" captionText="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618665" httpUri="https://zenodo.org/record/13618665/files/figure.png" pageId="21" pageNumber="358">24A</figureCitation>
). All dorsal shields smooth. Prodorsal shield with three pairs of setae (
<emphasis id="B936EA8BFFB37446845838CBFA4BFA42" box="[1403,1429,1427,1453]" italics="true" pageId="21" pageNumber="358">v1</emphasis>
,
<emphasis id="B936EA8BFFB3744681B438EFFF6CFA3E" box="[151,178,1463,1489]" italics="true" pageId="21" pageNumber="358">v2</emphasis>
,
<emphasis id="B936EA8BFFB3744681E338EFFF38FA3E" box="[192,230,1463,1489]" italics="true" pageId="21" pageNumber="358">sc2</emphasis>
), one pair of weakly barbed capitate trichobothria (
<emphasis id="B936EA8BFFB37446820D38EFFC8AFA3E" box="[814,852,1463,1489]" italics="true" pageId="21" pageNumber="358">sc1</emphasis>
) and one pair of oval stigmata. Stigmata associated with well-developed tracheal trunks. Setae
<emphasis id="B936EA8BFFB3744683A43884FD4DFA1A" box="[647,659,1500,1525]" italics="true" pageId="21" pageNumber="358">e</emphasis>
,
<emphasis id="B936EA8BFFB3744683803883FD61FA1A" box="[675,703,1499,1525]" italics="true" pageId="21" pageNumber="358">h1</emphasis>
and
<emphasis id="B936EA8BFFB3744683D83883FCC9FA1A" box="[763,791,1499,1525]" italics="true" pageId="21" pageNumber="358">h2</emphasis>
pointed; other dorsal setae blunt-tipped; all dorsal setae barbed. Alveoli of setae
<emphasis id="B936EA8BFFB37446808E3B58FE67F9F6" box="[429,441,1536,1561]" italics="true" pageId="21" pageNumber="358">e</emphasis>
and
<emphasis id="B936EA8BFFB3744680D138A7FE24F9F6" box="[498,506,1535,1561]" italics="true" pageId="21" pageNumber="358">f</emphasis>
distinctly separated. Cupules
<emphasis id="B936EA8BFFB3744682733B58FCB8F9F6" box="[848,870,1536,1561]" italics="true" pageId="21" pageNumber="358">ia</emphasis>
on tergite D,
<emphasis id="B936EA8BFFB3744685263B58FBFFF9F6" box="[1029,1057,1536,1561]" italics="true" pageId="21" pageNumber="358">im</emphasis>
on tergite EF and
<emphasis id="B936EA8BFFB3744685D538A7FAD2F9F6" box="[1270,1292,1535,1561]" italics="true" pageId="21" pageNumber="358">ih</emphasis>
on tergite H large, round;
<emphasis id="B936EA8BFFB3744680133B7CFE92F9D2" box="[304,332,1572,1597]" italics="true" pageId="21" pageNumber="358">im</emphasis>
much larger than subequal
<emphasis id="B936EA8BFFB3744683A93B7CFD7EF9D2" box="[650,672,1572,1597]" italics="true" pageId="21" pageNumber="358">ia</emphasis>
and
<emphasis id="B936EA8BFFB3744683FA3B7BFD31F9D2" box="[729,751,1571,1597]" italics="true" pageId="21" pageNumber="358">ih</emphasis>
. Lengths of dorsal setae:
<emphasis id="B936EA8BFFB3744685343B7BFBEFF9D2" box="[1047,1073,1571,1597]" italics="true" pageId="21" pageNumber="358">v1</emphasis>
21 (2022),
<emphasis id="B936EA8BFFB3744685E43B7BFB3CF9D2" box="[1223,1250,1571,1597]" italics="true" pageId="21" pageNumber="358">v2</emphasis>
16 (1517),
<emphasis id="B936EA8BFFB3744684543B7BFA43F9D2" box="[1399,1437,1571,1597]" italics="true" pageId="21" pageNumber="358">sc2</emphasis>
30 (3033),
<emphasis id="B936EA8BFFB3744680023B1FFEE2F98E" box="[289,316,1607,1633]" italics="true" pageId="21" pageNumber="358">c1</emphasis>
21 (2123),
<emphasis id="B936EA8BFFB3744680EA3B1FFE3AF98E" box="[457,484,1607,1633]" italics="true" pageId="21" pageNumber="358">c2</emphasis>
33 (3033),
<emphasis id="B936EA8BFFB3744683563B1FFD5DF98E" box="[629,643,1607,1633]" italics="true" pageId="21" pageNumber="358">d</emphasis>
24 (2425),
<emphasis id="B936EA8BFFB3744682373B10FCFEF98E" box="[788,800,1608,1633]" italics="true" pageId="21" pageNumber="358">e</emphasis>
20 (1920),
<emphasis id="B936EA8BFFB3744682923B1FFC67F98E" box="[945,953,1607,1633]" italics="true" pageId="21" pageNumber="358">f</emphasis>
25 (2527),
<emphasis id="B936EA8BFFB3744685693B1FFBBBF98E" box="[1098,1125,1607,1633]" italics="true" pageId="21" pageNumber="358">h1</emphasis>
36 (3336),
<emphasis id="B936EA8BFFB3744685D53B1FFACDF98E" box="[1270,1299,1607,1633]" italics="true" pageId="21" pageNumber="358">h2</emphasis>
20 (1320). Distances between setae:
<emphasis id="B936EA8BFFB37446809F3B33FE20F96A" box="[444,510,1643,1670]" italics="true" pageId="21" pageNumber="358">v1v1</emphasis>
19 (1820),
<emphasis id="B936EA8BFFB3744683B63B33FD09F96A" box="[661,727,1643,1669]" italics="true" pageId="21" pageNumber="358">v2v2</emphasis>
34 (3235),
<emphasis id="B936EA8BFFB37446824D3B33FC18F96A" box="[878,966,1643,1669]" italics="true" pageId="21" pageNumber="358">sc2sc2</emphasis>
32 (3236),
<emphasis id="B936EA8BFFB37446857E3B33FB7EF96A" box="[1117,1184,1643,1670]" italics="true" pageId="21" pageNumber="358">c1c1</emphasis>
32 (3235),
<emphasis id="B936EA8BFFB3744684173B33FAA9F96A" box="[1332,1399,1643,1670]" italics="true" pageId="21" pageNumber="358">c1c2</emphasis>
20 (1921),
<emphasis id="B936EA8BFFB3744681DD3BD7FEF6F946" box="[254,296,1679,1706]" italics="true" pageId="21" pageNumber="358">dd</emphasis>
43 (4352),
<emphasis id="B936EA8BFFB37446809B3BC8FE05F946" box="[440,475,1679,1706]" italics="true" pageId="21" pageNumber="358">ef</emphasis>
6 (69),
<emphasis id="B936EA8BFFB3744683623BD7FD81F946" box="[577,607,1679,1706]" italics="true" pageId="21" pageNumber="358">ff</emphasis>
36 (3639),
<emphasis id="B936EA8BFFB3744683D33BD7FCEBF946" box="[752,821,1679,1706]" italics="true" pageId="21" pageNumber="358">h1h1</emphasis>
37 (3643),
<emphasis id="B936EA8BFFB3744682E53BD7FBD2F946" box="[966,1036,1679,1706]" italics="true" pageId="21" pageNumber="358">h1h2</emphasis>
5 (56).
</paragraph>
<paragraph id="8BFD3699FFB3744681E43BEBFAFCF849" blockId="21.[151,1437,1427,1958]" pageId="21" pageNumber="358">
<emphasis id="B936EA8BFFB3744681E43BEBFE5DF922" box="[199,387,1715,1741]" italics="true" pageId="21" pageNumber="358">Idiosomal venter</emphasis>
(
<figureCitation id="13792A1CFFB3744680B23BEBFE24F922" box="[401,506,1715,1741]" captionStart="FIGURE 21" captionStartId="21.[151,250,1357,1381]" captionTargetBox="[151,1435,245,1333]" captionTargetId="figure-41@21.[151,1435,237,1333]" captionTargetPageId="21" captionText="FIGURE 21. Spatulaphorus reductus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618659" httpUri="https://zenodo.org/record/13618659/files/figure.png" pageId="21" pageNumber="358">Figs 21B</figureCitation>
,
<figureCitation id="13792A1CFFB3744683273BEBFDE9F922" box="[516,567,1715,1741]" captionStart="FIGURE 24" captionStartId="24.[151,247,1089,1113]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-24@24.[151,1436,181,1066]" captionTargetPageId="24" captionText="FIGURE 24. DIC micrographs of Spatulaphorus reductus sp. nov., female:A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618665" httpUri="https://zenodo.org/record/13618665/files/figure.png" pageId="21" pageNumber="358">24B</figureCitation>
). All ventral plates smooth.All ventral setae pointed; setae
<emphasis id="B936EA8BFFB3744685E63BEBFADDF922" box="[1221,1283,1715,1741]" italics="true" pageId="21" pageNumber="358">ps1-3</emphasis>
barbed, other ventral setae smooth. Bases of setae
<emphasis id="B936EA8BFFB3744683123B8FFD86F91E" box="[561,600,1751,1777]" italics="true" pageId="21" pageNumber="358">ps3</emphasis>
situated slightly anteriad bases of setae
<emphasis id="B936EA8BFFB3744685393B8FFB86F91E" box="[1050,1112,1751,1777]" italics="true" pageId="21" pageNumber="358">ps1-2</emphasis>
; alveoli of setae
<emphasis id="B936EA8BFFB37446843B3B8FFAE1F91E" box="[1304,1343,1751,1777]" italics="true" pageId="21" pageNumber="358">ps1</emphasis>
and
<emphasis id="B936EA8BFFB3744684563B8FFA42F91E" box="[1397,1436,1751,1777]" italics="true" pageId="21" pageNumber="358">ps2</emphasis>
clearly separated. Ap1 and ap2 well-developed and joined with thick appr; apsej absent; ap3 not evident; ap4 well developed, exceeding beyond bases of setae
<emphasis id="B936EA8BFFB3744683A03A47FD41F8D6" box="[643,671,1823,1849]" italics="true" pageId="21" pageNumber="358">3b</emphasis>
; ap5 absent.Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate straight in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
<emphasis id="B936EA8BFFB3744681B43A3FFF6AF86E" box="[151,180,1895,1921]" italics="true" pageId="21" pageNumber="358">1a</emphasis>
12 (1112),
<emphasis id="B936EA8BFFB3744680653A3FFEBCF86E" box="[326,354,1895,1921]" italics="true" pageId="21" pageNumber="358">1b</emphasis>
14 (1416),
<emphasis id="B936EA8BFFB3744680D53A3FFDCEF86E" box="[502,528,1895,1921]" italics="true" pageId="21" pageNumber="358">1с</emphasis>
13 (1315),
<emphasis id="B936EA8BFFB3744683873A3FFD61F86E" box="[676,703,1895,1921]" italics="true" pageId="21" pageNumber="358">2a</emphasis>
11 (1113),
<emphasis id="B936EA8BFFB3744682723A3FFCB5F86E" box="[849,875,1895,1921]" italics="true" pageId="21" pageNumber="358">2с</emphasis>
12 (1112),
<emphasis id="B936EA8BFFB3744682DD3A3FFBC4F86E" box="[1022,1050,1895,1921]" italics="true" pageId="21" pageNumber="358">3a</emphasis>
14 (1314),
<emphasis id="B936EA8BFFB37446858D3A3FFB14F86E" box="[1198,1226,1895,1921]" italics="true" pageId="21" pageNumber="358">3b</emphasis>
12 (1213),
<emphasis id="B936EA8BFFB37446847D3A3FFAA6F86E" box="[1374,1400,1895,1921]" italics="true" pageId="21" pageNumber="358">3c</emphasis>
14 (1314),
<emphasis id="B936EA8BFFB3744681DD3AD3FEC4F84A" box="[254,282,1931,1957]" italics="true" pageId="21" pageNumber="358">4a</emphasis>
13 (1013),
<emphasis id="B936EA8BFFB3744680893AD3FE19F84A" box="[426,455,1931,1957]" italics="true" pageId="21" pageNumber="358">4b</emphasis>
15 (1516),
<emphasis id="B936EA8BFFB3744683743AD3FDACF84A" box="[599,626,1931,1957]" italics="true" pageId="21" pageNumber="358">4c</emphasis>
16 (1216),
<emphasis id="B936EA8BFFB3744682213AD3FCF7F84A" box="[770,809,1931,1957]" italics="true" pageId="21" pageNumber="358">ps1</emphasis>
13 (1213),
<emphasis id="B936EA8BFFB3744682993AD3FC3FF84A" box="[954,993,1931,1957]" italics="true" pageId="21" pageNumber="358">ps2</emphasis>
13 (1213),
<emphasis id="B936EA8BFFB3744685523AD3FB46F84A" box="[1137,1176,1931,1957]" italics="true" pageId="21" pageNumber="358">ps3</emphasis>
16 (1516).
</paragraph>
<caption id="DF3D6611FFB0744581B438FAFAC7FA55" ID-DOI="http://doi.org/10.5281/zenodo.13618661" ID-Zenodo-Dep="13618661" box="[151,1305,1442,1467]" httpUri="https://zenodo.org/record/13618661/files/figure.png" pageId="22" pageNumber="359" startId="22.[151,250,1442,1466]" targetBox="[151,1417,185,1419]" targetPageId="22" targetType="figure">
<paragraph id="8BFD3699FFB0744581B438FAFAC7FA55" blockId="22.[151,1305,1442,1467]" box="[151,1305,1442,1467]" pageId="22" pageNumber="359">
<emphasis id="B936EA8BFFB0744581B438FAFEC7FA55" bold="true" box="[151,281,1442,1466]" pageId="22" pageNumber="359">FIGURE 22</emphasis>
.
<taxonomicName id="4C424D1AFFB07445800538FBFDC8FA54" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1443,1467]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="359" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB07445800538FBFDC8FA54" box="[294,534,1443,1467]" italics="true" pageId="22" pageNumber="359">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB07445833E38FBFDB3FA54" bold="true" box="[541,621,1443,1467]" pageId="22" pageNumber="359">
<taxonomicNameLabel id="A20557F0FFB07445833E38FBFDB3FA54" box="[541,621,1443,1467]" pageId="22" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB0744581E438B2FE05F85C" blockId="22.[151,1437,1513,1971]" pageId="22" pageNumber="359">
<emphasis id="B936EA8BFFB0744581E438B2FF22F9EC" box="[199,252,1514,1539]" italics="true" pageId="22" pageNumber="359">Legs</emphasis>
(
<figureCitation id="13792A1CFFB07445802A38B1FE80F9EC" box="[265,350,1513,1539]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618661" httpUri="https://zenodo.org/record/13618661/files/figure.png" pageId="22" pageNumber="359">Figs 22</figureCitation>
,
<figureCitation id="13792A1CFFB07445804438B1FE59F9EC" box="[359,391,1513,1539]" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618663" httpUri="https://zenodo.org/record/13618663/files/figure.png" pageId="22" pageNumber="359">23</figureCitation>
). Setation of legs as in
<taxonomicName id="4C424D1AFFB07445835D38B1FCFFF9EC" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[638,801,1513,1539]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="359" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="B936EA8BFFB07445835D38B1FCFFF9EC" box="[638,801,1513,1539]" italics="true" pageId="22" pageNumber="359">S. brevisetosus</emphasis>
</taxonomicName>
except presence of solenidia
<emphasis id="B936EA8BFFB07445854238B2FBAEF9EC" box="[1121,1136,1514,1539]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
on tibiae II-III and absence of seta
<emphasis id="B936EA8BFFB0744581C43B55FED2F9C8" box="[231,268,1549,1575]" italics="true" pageId="22" pageNumber="359">pl”</emphasis>
on tarsus IV. Leg I (
<figureCitation id="13792A1CFFB0744580CE3B55FD88F9C8" box="[493,598,1549,1575]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618661" httpUri="https://zenodo.org/record/13618661/files/figure.png" pageId="22" pageNumber="359">Fig. 22A</figureCitation>
). Tibiotarsus enlarged, with large and thick claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
<emphasis id="B936EA8BFFB0744583F83B69FD27F9A4" box="[731,761,1585,1611]" italics="true" pageId="22" pageNumber="359">φ2</emphasis>
. Setae (
<emphasis id="B936EA8BFFB07445827B3B6AFCB8F9A4" box="[856,870,1586,1611]" italics="true" pageId="22" pageNumber="359">u</emphasis>
) consolidated into a horn-like structure opposing claw. Tarsal claw blunt-tipped. Seta
<emphasis id="B936EA8BFFB0744583113B0DFD9EF980" box="[562,576,1621,1647]" italics="true" pageId="22" pageNumber="359">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="B936EA8BFFB0744585D33B0DFADFF980" box="[1264,1281,1621,1647]" italics="true" pageId="22" pageNumber="359">l</emphasis>
of femur and
<emphasis id="B936EA8BFFB0744581B43B21FF7DF97C" box="[151,163,1657,1683]" italics="true" pageId="22" pageNumber="359">k</emphasis>
of tibiotarsus smooth, other leg setae (except eupathidia
<emphasis id="B936EA8BFFB0744582133B21FC99F97C" box="[816,839,1657,1683]" italics="true" pageId="22" pageNumber="359">p</emphasis>
,
<emphasis id="B936EA8BFFB0744582753B21FCAAF97C" box="[854,884,1657,1683]" italics="true" pageId="22" pageNumber="359">p”</emphasis>
,
<emphasis id="B936EA8BFFB0744582A03B21FC7FF97C" box="[899,929,1657,1683]" italics="true" pageId="22" pageNumber="359">tc</emphasis>
,
<emphasis id="B936EA8BFFB0744582933B21FC0AF97C" box="[944,980,1657,1683]" italics="true" pageId="22" pageNumber="359">tc”</emphasis>
,
<emphasis id="B936EA8BFFB0744582C73B21FC23F97C" box="[996,1021,1657,1683]" italics="true" pageId="22" pageNumber="359">ft</emphasis>
,
<emphasis id="B936EA8BFFB07445852F3B21FBF5F97C" box="[1036,1067,1657,1683]" italics="true" pageId="22" pageNumber="359">ft”</emphasis>
) barbed. Setae
<emphasis id="B936EA8BFFB0744585FE3B21FB35F97C" box="[1245,1259,1657,1683]" italics="true" pageId="22" pageNumber="359">l</emphasis>
of femur and
<emphasis id="B936EA8BFFB0744584B33B21FA42F97C" box="[1424,1436,1657,1683]" italics="true" pageId="22" pageNumber="359">k</emphasis>
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="B936EA8BFFB0744585DB3BC5FAC4F958" box="[1272,1306,1693,1719]" italics="true" pageId="22" pageNumber="359">ω1</emphasis>
6 (6),
<emphasis id="B936EA8BFFB0744584463BC5FA59F958" box="[1381,1415,1693,1719]" italics="true" pageId="22" pageNumber="359">ω2</emphasis>
3 (3),
<emphasis id="B936EA8BFFB0744581E53B99FF3DF934" box="[198,227,1729,1755]" italics="true" pageId="22" pageNumber="359">φ1</emphasis>
5 (5),
<emphasis id="B936EA8BFFB07445800D3B99FE92F934" box="[302,332,1729,1755]" italics="true" pageId="22" pageNumber="359">φ2</emphasis>
4 (4); solenidia
<emphasis id="B936EA8BFFB0744583203B99FDFAF934" box="[515,548,1729,1755]" italics="true" pageId="22" pageNumber="359">ω1</emphasis>
and
<emphasis id="B936EA8BFFB0744583783B99FDA6F934" box="[603,632,1729,1755]" italics="true" pageId="22" pageNumber="359">φ1</emphasis>
thick, clavate; other solenidia weakly clavate. Leg II (
<figureCitation id="13792A1CFFB0744585F93B99FA9FF934" box="[1242,1345,1729,1755]" captionStart="FIGURE 22" captionStartId="22.[151,250,1442,1466]" captionTargetBox="[151,1417,185,1419]" captionTargetId="figure-24@22.[151,1436,181,1419]" captionTargetPageId="22" captionText="FIGURE 22. Spatulaphorus reductus sp. nov., female: A—right leg I, dorsal aspect; B—left leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618661" httpUri="https://zenodo.org/record/13618661/files/figure.png" pageId="22" pageNumber="359">Fig. 22B</figureCitation>
). Tarsus with simple claws and pad-like empodium. Solenidion
<emphasis id="B936EA8BFFB0744582213BBEFCC8F910" box="[770,790,1766,1791]" italics="true" pageId="22" pageNumber="359">ω</emphasis>
6 (56) weakly clavate, solenidion
<emphasis id="B936EA8BFFB0744585843BBEFB68F910" box="[1191,1206,1766,1791]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
2 (2) very short. All leg setae barbed and pointed. Leg III (
<figureCitation id="13792A1CFFB0744583663A51FD70F8CC" box="[581,686,1801,1827]" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618663" httpUri="https://zenodo.org/record/13618663/files/figure.png" pageId="22" pageNumber="359">Fig. 23A</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. All leg setae barbed and pointed; solenidion
<emphasis id="B936EA8BFFB0744582EE3A76FC02F8A8" box="[973,988,1838,1863]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
very small (less than 0.5). Leg IV (
<figureCitation id="13792A1CFFB0744584563A75FF14F884" captionStart="FIGURE 23" captionStartId="23.[151,250,1740,1764]" captionTargetBox="[160,1427,190,1716]" captionTargetId="figure-17@23.[151,1436,181,1716]" captionTargetPageId="23" captionText="FIGURE 23. Spatulaphorus reductus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618663" httpUri="https://zenodo.org/record/13618663/files/figure.png" pageId="22" pageNumber="359">Fig 23B</figureCitation>
). Femur divided into basi- and telofemur. Claws and empodium as on tarsi II and III. All leg setae barbed and pointed; external part of solenidion
<emphasis id="B936EA8BFFB0744583123A2EFD9EF860" box="[561,576,1910,1935]" italics="true" pageId="22" pageNumber="359">φ</emphasis>
absent, but subcuticular very small cup-shaped structure situated at typical insertion point of solenidion.
</paragraph>
<caption id="DF3D6611FFB1744481B43B94FAE0F90C" ID-DOI="http://doi.org/10.5281/zenodo.13618663" ID-Zenodo-Dep="13618663" box="[151,1342,1740,1764]" httpUri="https://zenodo.org/record/13618663/files/figure.png" pageId="23" pageNumber="360" startId="23.[151,250,1740,1764]" targetBox="[160,1427,190,1716]" targetPageId="23" targetType="figure">
<paragraph id="8BFD3699FFB1744481B43B94FAE0F90C" blockId="23.[151,1342,1740,1764]" box="[151,1342,1740,1764]" pageId="23" pageNumber="360">
<emphasis id="B936EA8BFFB1744481B43B94FEC7F90B" bold="true" box="[151,281,1740,1764]" pageId="23" pageNumber="360">FIGURE 23</emphasis>
.
<taxonomicName id="4C424D1AFFB1744480053B94FDC8F90B" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[294,534,1740,1764]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFB1744480053B94FDC8F90B" box="[294,534,1740,1764]" italics="true" pageId="23" pageNumber="360">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFB17444833E3B94FDB3F90B" bold="true" box="[541,621,1740,1764]" pageId="23" pageNumber="360">
<taxonomicNameLabel id="A20557F0FFB17444833E3B94FDB3F90B" box="[541,621,1740,1764]" pageId="23" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
</paragraph>
</caption>
<paragraph id="8BFD3699FFB1744481EB3A4AFE4FF8C3" blockId="23.[151,1436,1810,1981]" box="[200,401,1810,1836]" pageId="23" pageNumber="360">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="C3586512FFB1744B81E43A6EFD9BFB05" lastPageId="24" lastPageNumber="361" pageId="23" pageNumber="360" type="materials_examined">
<paragraph id="8BFD3699FFB1744481E43A6EFC10F852" blockId="23.[151,1436,1810,1981]" pageId="23" pageNumber="360">
<emphasis id="B936EA8BFFB1744481E43A6EFEAFF8BF" bold="true" box="[199,369,1846,1872]" pageId="23" pageNumber="360">Type material</emphasis>
. Female
<typeStatus id="54F9883BFFB1744480FD3A6EFD9DF8BF" box="[478,579,1846,1872]" pageId="23" pageNumber="360" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="ED53AE5CFFB1744483AC3A6EFD16F8BF" box="[655,712,1846,1872]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="23" pageNumber="360">ZISP</collectionCode>
T-Pygm-015
<collectingCountry id="F3557609FFB1744482453A6EFC06F8BF" box="[870,984,1846,1872]" name="Argentina" pageId="23" pageNumber="360">Argentina</collectingCountry>
,
<collectingRegion id="4986F87BFFB1744482C53A6EFB10F8BE" box="[998,1230,1846,1873]" country="Argentina" name="Santiago del Estero" pageId="23" pageNumber="360">Santiago Del Estero</collectingRegion>
, Salado River, on
<taxonomicName id="4C424D1AFFB1744481B43A03FE21F89B" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[151,511,1882,1908]" class="Insecta" family="Scarabaeidae" genus="Parataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="simulator">
<emphasis id="B936EA8BFFB1744481B43A03FE4DF89B" box="[151,403,1882,1908]" italics="true" pageId="23" pageNumber="360">Parataenius simulator</emphasis>
(Harold)
</taxonomicName>
;
<typeStatus id="54F9883BFFB1744483283A03FDA5F89A" box="[523,635,1883,1909]" pageId="23" pageNumber="360" type="paratype">paratypes</typeStatus>
: same locality data,
<specimenCount id="9D44FD10FFB1744482413A02FC03F89B" box="[866,989,1882,1908]" count="47" pageId="23" pageNumber="360" type="female">47 females</specimenCount>
on 9
<taxonomicName id="4C424D1AFFB17444853C3A03FAC4F89B" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1055,1306,1882,1908]" class="Insecta" family="Scarabaeidae" genus="Parataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="simulator">
<emphasis id="B936EA8BFFB17444853C3A03FAC4F89B" box="[1055,1306,1882,1908]" italics="true" pageId="23" pageNumber="360">Parataenius simulator</emphasis>
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744484A33A02FF30F877" count="38" pageId="23" pageNumber="360" type="female">38 females</specimenCount>
on 7
<taxonomicName id="4C424D1AFFB17444800D3A26FDE0F877" authority="Harold" authorityName="Harold" authorityYear="1867" box="[302,574,1918,1944]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="picinus">
<emphasis id="B936EA8BFFB17444800D3A26FE36F877" box="[302,488,1918,1944]" italics="true" pageId="23" pageNumber="360">Ataenius picinus</emphasis>
Harold
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744483823A26FCD0F877" box="[673,782,1918,1944]" count="8" pageId="23" pageNumber="360" type="female">8 females</specimenCount>
on 2
<taxonomicName id="4C424D1AFFB17444826D3A26FBA6F877" authority="Schmidt" authorityName="Schmidt" authorityYear="1916" box="[846,1144,1918,1944]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis id="B936EA8BFFB17444826D3A26FBCAF877" box="[846,1044,1918,1944]" italics="true" pageId="23" pageNumber="360">Ataenius clavatus</emphasis>
Schmidt
</taxonomicName>
beetles;
<specimenCount id="9D44FD10FFB1744485FE3A26FA97F877" box="[1245,1353,1918,1945]" count="5" pageId="23" pageNumber="360" type="female">5 females</specimenCount>
on one
<taxonomicName id="4C424D1AFFB1744481B43AFAFE2FF853" authority="(Blanchard)" baseAuthorityName="Blanchard" baseAuthorityYear="1846" box="[151,497,1954,1980]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="360" phylum="Arthropoda" rank="species" species="platensis">
<emphasis id="B936EA8BFFB1744481B43AFAFEBAF853" box="[151,356,1954,1980]" italics="true" pageId="23" pageNumber="360">Ataenius platensis</emphasis>
(Blanchard)
</taxonomicName>
beetle. All host beetles are housed in
<collectionCode id="ED53AE5CFFB1744482BB3AFBFC14F852" box="[920,970,1955,1981]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="23" pageNumber="360" type="Museum">ZIN</collectionCode>
.
</paragraph>
<caption id="DF3D6611FFBE744B81B43919FA42FBB5" ID-DOI="http://doi.org/10.5281/zenodo.13618665" ID-Zenodo-Dep="13618665" box="[151,1436,1089,1114]" httpUri="https://zenodo.org/record/13618665/files/figure.png" pageId="24" pageNumber="361" startId="24.[151,247,1089,1113]" targetBox="[151,1436,181,1066]" targetPageId="24" targetType="figure">
<paragraph id="8BFD3699FFBE744B81B43919FA42FBB5" blockId="24.[151,1436,1089,1114]" box="[151,1436,1089,1114]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81B43919FEC8FBB6" bold="true" box="[151,278,1089,1113]" pageId="24" pageNumber="361">FIGURE 24</emphasis>
. DIC micrographs of
<taxonomicName id="4C424D1AFFBE744B80CD391AFD09FBB5" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[494,727,1090,1114]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFBE744B80CD391AFD09FBB5" box="[494,727,1090,1114]" italics="true" pageId="24" pageNumber="361">Spatulaphorus reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBE744B83FE391AFCF5FBB5" bold="true" box="[733,811,1090,1114]" pageId="24" pageNumber="361">
<taxonomicNameLabel id="A20557F0FFBE744B83FE391AFCF5FBB5" box="[733,811,1090,1114]" pageId="24" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A—general view dorsally; B—general view ventrally.
</paragraph>
</caption>
<paragraph id="8BFD3699FFBE744B81E439D0FD9BFB05" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E439D0FE58FB4D" bold="true" box="[199,390,1160,1186]" pageId="24" pageNumber="361">Type deposition</emphasis>
.
<materialsCitation id="3B2A3CC4FFBE744B80B439D0FE25FB29" country="Russia" location="Zoological Institute" pageId="24" pageNumber="361" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
The
<typeStatus id="54F9883BFFBE744B80EF39D0FDF0FB4D" box="[460,558,1160,1186]" pageId="24" pageNumber="361" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD10FFBE744B834939D0FCD3FB4D" box="[618,781,1160,1186]" count="4" pageId="24" pageNumber="361" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are deposited in the collection of
<location id="8E9D6042FFBE744B85BF39D0FAA5FB4D" LSID="urn:lsid:plazi:treatment:03EB878FFFB2744B81B43BF9FE00FA71:8E9D6042FFBE744B85BF39D0FAA5FB4D" box="[1180,1403,1160,1186]" country="Russia" name="Zoological Institute" pageId="24" pageNumber="361" stateProvince="Saint Petersburg">Zoological Institute</location>
of RAS,
<collectingRegion id="4986F87BFFBE744B81FC39F4FE40FB29" box="[223,414,1196,1222]" country="Russia" name="Saint Petersburg" pageId="24" pageNumber="361">Saint Petersburg</collectingRegion>
,
<collectingCountry id="F3557609FFBE744B808E39F4FE25FB29" box="[429,507,1196,1222]" name="Russia" pageId="24" pageNumber="361">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B2A3CC4FFBE744B832939F4FAC5FB29" box="[522,1307,1196,1222]" country="Russia" location="Tyumen State" pageId="24" pageNumber="361" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
other
<typeStatus id="54F9883BFFBE744B836D39F4FD67FB29" box="[590,697,1196,1222]" pageId="24" pageNumber="361" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="4986F87BFFBE744B855E39F4FAC5FB29" box="[1149,1307,1196,1222]" country="Russia" name="Tyumen'" pageId="24" pageNumber="361">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2A3CC4FFBE744B840539F4FD9EFB05" country="Russia" location="University Museum of Zoology" pageId="24" pageNumber="361" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location id="8E9D6042FFBE744B840539F4FEA1FB05" LSID="urn:lsid:plazi:treatment:03EB878FFFB2744B81B43BF9FE00FA71:8E9D6042FFBE744B840539F4FEA1FB05" country="Russia" name="University Museum of Zoology" pageId="24" pageNumber="361" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4986F87BFFBE744B80AA3988FE39FB05" box="[393,487,1232,1258]" country="Russia" name="Tyumen'" pageId="24" pageNumber="361">Tyumen</collectingRegion>
,
<collectingCountry id="F3557609FFBE744B80D13988FD9EFB05" box="[498,576,1232,1258]" name="Russia" pageId="24" pageNumber="361">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBE744B81E439ACFB2EFAB9" pageId="24" pageNumber="361" type="diagnosis">
<paragraph id="8BFD3699FFBE744B81E439ACFB2EFAB9" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E439ACFE18FAE1" bold="true" box="[199,454,1268,1294]" pageId="24" pageNumber="361">Differential diagnosis</emphasis>
. The new species significantly differs from other species of the genus
<taxonomicName id="4C424D1AFFBE744B85FF39ACFAA1FAE1" authorityName="Rack" authorityYear="1993" box="[1244,1407,1268,1294]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFBE744B85FF39ACFAA1FAE1" box="[1244,1407,1268,1294]" italics="true" pageId="24" pageNumber="361">Spatulaphorus</emphasis>
</taxonomicName>
in having tarsus IV with five setae (
<emphasis id="B936EA8BFFBE744B83293840FDEEFADD" box="[522,560,1304,1330]" italics="true" pageId="24" pageNumber="361">pl”</emphasis>
absent) and the alveoli of setae
<emphasis id="B936EA8BFFBE744B82B43840FC60FADD" box="[919,958,1304,1330]" italics="true" pageId="24" pageNumber="361">ps1</emphasis>
and
<emphasis id="B936EA8BFFBE744B82D73840FBC5FADD" box="[1012,1051,1304,1330]" italics="true" pageId="24" pageNumber="361">ps2</emphasis>
clearly separated. In other species of
<taxonomicName id="4C424D1AFFBE744B81953864FE87FAB9" authorityName="Rack" authorityYear="1993" box="[182,345,1340,1366]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFBE744B81953864FE87FAB9" box="[182,345,1340,1366]" italics="true" pageId="24" pageNumber="361">Spatulaphorus</emphasis>
</taxonomicName>
tarsus IV has six setae (
<emphasis id="B936EA8BFFBE744B834A3864FD50FAB9" box="[617,654,1340,1366]" italics="true" pageId="24" pageNumber="361">pl”</emphasis>
present) and the alveoli of setae
<emphasis id="B936EA8BFFBE744B82DC3864FBE3FAB9" box="[1023,1085,1340,1366]" italics="true" pageId="24" pageNumber="361">ps1-2</emphasis>
are contiguous.
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBE744B81E43838FE00FA71" pageId="24" pageNumber="361" type="etymology">
<paragraph id="8BFD3699FFBE744B81E43838FE00FA71" blockId="24.[151,1437,1160,1438]" pageId="24" pageNumber="361">
<emphasis id="B936EA8BFFBE744B81E43838FE96FA95" bold="true" box="[199,328,1376,1402]" pageId="24" pageNumber="361">Etymology</emphasis>
. The name of the new species is derived from Latin
<taxonomicName id="4C424D1AFFBE744B82A63838FC3AFA95" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[901,996,1376,1402]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="361" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B936EA8BFFBE744B82A63838FC3AFA95" box="[901,996,1376,1402]" italics="true" pageId="24" pageNumber="361">reductus</emphasis>
</taxonomicName>
meaning
<emphasis id="B936EA8BFFBE744B85733838FB77FA95" box="[1104,1193,1376,1402]" italics="true" pageId="24" pageNumber="361">reduced</emphasis>
, and refers to reduced number of setae on tarsus IV.
</paragraph>
</subSubSection>
</treatment>
</document>